Diplazium urticifolium Christ

MONRO, A. K., SANTAMARÍA-AGUILAR, D., GONZÁLEZ, F., CHACÓN, O., SOLANO, D., RODRÍGUEZ, A., ZAMORA, N., FEDELE, E. & CORREA, M., 2017, A first checklist to the vascular plants of La Amistad International Park (PILA), Costa Rica-Panama, Phytotaxa 322 (1), pp. 448-450 : 448-450

Diplazium urticifolium Christ Thelypteris atrovirens (C.Chr.) C.F.Reed

Terrestrial herbs. Distribution in PILA 900–1700 m, mixed Terrestrial herbs. Distribution in PILA 1200–2500 m, forest in transition to low elevation forest, mixed forest in mixed forest in transition to low elevation forest, mixed transition to cloud forest. Global distribution: Mexico to forest in transition to cloud forest, cloud forest, low Venezuela. Records: A. Rodríguez 10818 (BM, INB, MO, elevation oak forest. Global distribution: Mexico to Colombia. Records: A. Rodríguez 10817 (BM, INB, MO,

PMA), A. Rodríguez 10897 (BM, INB, MO, PMA), G. forest. Global distribution: Costa Rica to Peru. Records: Davidse 28945 (CR, MO), G. Herrera 3272 (INB, MO), G. Herrera 3120 (MO). G. Herrera 5277 (CR, INB, MO), G. Herrera 6066 (CR, INB), J. Bittner 1912 (INB, MO), M. Grayum 10365 Thelypteris funckii (Mett.) Alston (INB, MO) . Terrestrial herbs. Distribution in PILA 2200–3000 m, low elevation oak forest, high elevation oak forest. Global

Thelypteris brachypus (Sodiro) R.M.Tryon & A.F.Tryon distribution: Costa Rica to the Guyanas. Records: G. Terrestrial herbs. Distribution in PILA 2500–2800 m, low Davidse 25096 (CR, MO), G. Davidse 25787 (MO), G. elevation oak forest, high elevation oak forest. Global Davidse 29266 (CR, MO), M. Moraga 320 (CR, INB). distribution: Panama to Peru. Records: L.D. Gómez 22283 (MO). Thelypteris germaniana (Fée) Proctor Terrestrial herbs. Distribution in PILA 900–2000 m, mixed ε-PILA Thelypteris chiriquiana (A.R.Sm.) Salino & forest in transition to low elevation forest, mixed forest in T.E.Almeida transition to cloud forest. Global distribution: Costa Rica Terrestrial herbs. Distribution in PILA 2200– 2200 m, to Venezuela, French Guyana, Greater Antilles, Lesser low elevation oak forest. Global distribution: endemic to Antilles. Records: A. Chacón 271 (CR, MO). Panama. Records: H. Van der Werff 6319 (MO). Thelypteris gigantea (Mett.) R.M.Tryon

Thelypteris cinerea A.R.Sm. Terrestrial herbs. Distribution in PILA 100–1400 m, low Terrestrial herbs. Distribution in PILA 1300–2300 m, elevation mixed forest, mixed forest in transition to low mixed forest in transition to cloud forest, cloud forest, elevation forest, mixed forest in transition to cloud forest. low elevation oak forest. Global distribution: Mexico Global distribution: Costa Rica to Peru. Records: G. to Ecuador, Venezuela. Records: G. Herrera 6031 (CR, Herrera 3094 (MO). INB, MO). Thelypteris gomeziana A.R.Sm. & Lellinger

Thelypteris decussata (L.) Proctor Terrestrial herbs. Distribution in PILA 2900–3500 m, Terrestrial herbs. Distribution in PILA 0–2300 m, low high elevation oak forest, páramo. Global distribution: elevation mixed forest, mixed forest in transition to low Mexico, Costa Rica, Panama. Records: G. Davidse 26047 elevation forest, mixed forest in transition to cloud forest, (CR, MO), G. Davidse 29323 (CR, MO). cloud forest, low elevation oak forest. Global distribution: Guatemala to Peru, the Guyanas, Brazil, Bolivia, Greater Thelypteris inaequans (C.Chr.) Lellinger Antilles, Lesser Antilles. Records : A.K. Monro 5579 (BM, Terrestrial herbs. Distribution in PILA 1300–2200 m, INB, MO, PMA), A. Rodríguez 10768 (BM, INB, MO, mixed forest in transition to cloud forest, cloud forest, PMA), A. Rodríguez 10871 (BM, INB, MO, PMA), A. low elevation oak forest. Global distribution: Mexico to Rodríguez 11507 (BM, INB, MO, PMA), D. Santamaría Ecuador. Records: M. Grayum 10290 (INB, MO). 6616 (BM, INB, MO, PMA), M. Moraga 507 (INB, MO). Thelypteris jimenezii (Maxon & C.Chr.) C.F.Reed Terrestrial herbs. Distribution in PILA 1200–2200 m,

Thelypteris deflexa (C.Presl) R.M.Tryon mixed forest in transition to low elevation forest, mixed Terrestrial herbs. Distribution in PILA 2400–3000 m, low forest in transition to cloud forest, cloud forest, low elevation oak forest, high elevation oak forest. Global elevation oak forest. Global distribution: Costa Rica, distribution: Mexico to Venezuela. Records: G. Davidse Panama. Records: G. Herrera 2670 (INB, MO), G. 28689 (CR, MO), G. Herrera 3729 (INB, MO). Herrera 5479 (INB, MO), G. Herrera 5497 (CR, INB, MO).

Thelypteris delasotae A.R.Sm. & Lellinger Terrestrial herbs. Distribution in PILA 2200–3200 m, Thelypteris leprieurii (Hook.) R.M.Tryon low elevation oak forest, páramo. Global distribution: Terrestrial herbs. Distribution in PILA 100–1800 m, low Mexico, Honduras, Costa Rica, Panama. Records: G. elevation mixed forest, mixed forest in transition to low Davidse 24000 (CR, MO), G. Davidse 24001 (CR, MO), elevation forest, mixed forest in transition to cloud forest. G. Davidse 28642 (CR, MO). Global distribution: Mexico to Colombia. Records: A. Rodríguez 10999 (BM, INB, MO, PMA), D. Santamaría

Thelypteris eggersii (Hieron.) C.F.Reed 6669 (BM, INB, MO, PMA), G. Herrera 2820 (INB, Terrestrial herbs. Distribution in PILA 100–3000 m, low MO). elevation mixed forest, mixed forest in transition to low elevation forest, mixed forest in transition to cloud forest, Thelypteris linkiana (C.Presl) R.M.Tryon cloud forest, low elevation oak forest, high elevation oak

Terrestrial herbs. Distribution in PILA 900–1200 m, Antilles, Lesser Antilles. Records: G. Davidse 28885 mixed forest in transition to low elevation forest. Global (CR, MO), G. Davidse 28944 (CR, MO), G. Davidse distribution: Mexico to Bolivia, Greater Antilles, Lesser 29222 (CR, MO), G. Herrera 5871 (CR, INB, MO), F. Antilles. Records: A. Rodríguez 11047 (BM, INB, MO, Quesada 1435 (INB).

PMA).

Thelypteris rupestris (Klotzsch) C.F.Reed

Thelypteris longipilosa (Sodiro) C.F.Reed Terrestrial herbs. Distribution in PILA 1800–2000 m, Terrestrial herbs. Distribution in PILA 2100– 2100 m, cloud forest. Global distribution: Costa Rica to Ecuador, cloud forest, low elevation oak forest. Global distribution: Venezuela, Greater Antilles. Records: M. Grayum 10366 Costa Rica to Peru. Records: M. Moraga 479 (CR, INB). (INB, MO).

Thelypteris oaxacana A.R.Sm. Thelypteris serrata (Cav.) Alston Terrestrial herbs. Distribution in PILA 1300–2200 m, Terrestrial herbs. Distribution in PILA 0–2200 m, low mixed forest in transition to cloud forest, cloud forest, elevation mixed forest, mixed forest in transition to low low elevation oak forest. Global distribution: Mexico to elevation forest, mixed forest in transition to cloud forest, Panama. Records: A. Rodríguez 10960 (BM, INB, MO, cloud forest, low elevation oak forest. Global distribution: PMA), M. Grayum 10380 (INB, MO). USA to Mexico to Argentina, Greater Antilles, Lesser Antilles. Records: A. Rodríguez 11193 (BM, INB, MO,

Thelypteris oligocarpa (Humb. & Bonpl. ex Willd.) Ching PMA ). Terrestrial herbs. Distribution in PILA 1200–1600 m, mixed forest in transition to low elevation forest. Global Thelypteris subscandens A.R.Sm. distribution: Mexico to Argentina, Greater Antilles, Lesser Terrestrial herbs. Distribution in PILA 1900–2700 m, Antilles. Records: J. Bittner 1915 (INB, MO), M. Moraga cloud forest, low elevation oak forest, high elevation 478 (INB). oak forest. Global distribution: Endemic to Costa Rica. Records: F. Quesada 1931 (INB), M. Grayum 10372

Thelypteris pachyrhachis (Kunze ex Mett.) Ching (INB, MO) . Terrestrial herbs. Distribution in PILA 1900–2600 m, cloud forest, low elevation oak forest, high elevation oak Thelypteris thomsonii (Jenman) Proctor forest. Global distribution: Costa Rica to Brazil, Greater Terrestrial herbs. Distribution in PILA 1300–2600 m, Antilles, Lesser Antilles. Records: G. Davidse 25820 mixed forest in transition to cloud forest, cloud forest, (CR, MO), M. Grayum 10320 (INB, MO). low elevation oak forest, high elevation oak forest. Global distribution: Mexico to Peru, Greater Antilles. Records:

Thelypteris pilosula (Klotzsch & H.Karst. ex Mett.) G. Davidse 25821 (CR, MO). R.M.Tryon Terrestrial herbs. Distribution in PILA 1600–2600 m, Thelypteris valdepilosa (Baker) C.F.Reed mixed forest in transition to cloud forest, cloud forest, Terrestrial herbs. Distribution in PILA 700–1100 m, low low elevation oak forest, high elevation oak forest. Global elevation mixed forest, mixed forest in transition to low distribution: Mexico, Guatemala, Costa Rica to Bolivia, elevation forest. Global distribution: Costa Rica to Peru. Greater Antilles. Records: G. Davidse 25778 (CR, MO), Records: A. Chacón 289 (CR, MO), G. Herrera 3249 M. Chinchilla 5 (CR, INB, MO), M. Moraga 453 (INB), (INB, MO). M. Moraga 480 (CR, INB). CYATHEACEAE

Thelypteris pusilla (Mett.) Ching Terrestrial herbs. Distribution in PILA 1900–3000 m, Alsophila erinacea (H.Karst.) D.S.Conant cloud forest, low elevation oak forest, high elevation Terrestrial trees. Distribution in PILA 0–2200 m, low oak forest. Global distribution: Costa Rica to Bolivia. elevation mixed forest, mixed forest in transition to low Records: G. Davidse 25245 (CR, MO), G. Davidse 29233 elevation forest, mixed forest in transition to cloud forest, (CR, MO), G. Herrera 3624 (MO), M. Grayum 10304 cloud forest, low elevation oak forest. Global distribution: (CR, INB, MO). Costa Rica to Bolivia. Records: A. Fernández 898 (INB), A. Rodríguez 11592 (BM, INB, MO, PMA), J. Bittner

Thelypteris rudis (Kunze) Proctor 1919 (INB, MO). Terrestrial herbs. Distribution in PILA 0–3000 m, low elevation mixed forest, mixed forest in transition to low Culcita coniifolia (Hook.) Maxon elevation forest, mixed forest in transition to cloud forest, Terrestrial trees. Distribution in PILA 1500–3200 m, cloud forest, low elevation oak forest, high elevation oak mixed forest in transition to cloud forest, cloud forest, forest. Global distribution: Mexico to Bolivia, Greater low elevation oak forest, páramo. Global distribution: Mexico to Brazil, Greater Antilles. Records: A.K. Monro Cyathea gracilis Griseb. 5286 (BM, INB, MO), A.K. Monro 5484 (BM, INB, MO, Terrestrial shrubs. Distribution in PILA 1200–1800 m, PMA), A. Rodríguez 10659,1 (BM, INB, MO, PMA), A. mixed forest in transition to low elevation forest, mixed Soto 335 (INB), D. Santamaría 5691 (BM, INB, MO, forest in transition to cloud forest. Global distribution: PMA), G. Davidse 25828 (CR, MO), G. Davidse 25829 Costa Rica to Colombia, Greater Antilles. Records: A. (MO), G. Davidse 28641 (MO), G. Davidse 29238 (MO), Rodríguez 10683 (BM, INB, MO, PMA), J. Bittner 1779 G. Herrera 3712 (INB, MO), M.A. Thomas 1582 (MO), (INB, MO). M.A. Thomas 1617 (MO), M. Moraga 338 (INB, MO), R. Aguilar 1160 (INB, MO). Cyathea mutica (Christ) Domin Terrestrial shrubs. Distribution in PILA 200–2100 m, low

Cyathea acutidens (Christ) Domin elevation mixed forest, mixed forest in transition to low Terrestrial trees. Distribution in PILA 100–2000 m, low elevation forest, mixed forest in transition to cloud forest, elevation mixed forest, mixed forest in transition to low cloud forest, low elevation oak forest. Global distribution: elevation forest, mixed forest in transition to cloud forest. Mexico to Panama, Venezuela. Records: G. Herrera 5476 Global distribution: Costa Rica, Ecuador, French Guyana, (INB), G. Herrera 6045 (INB, MO). Bolivia. Records: E. Alfaro 2606 (INB), E. Navarro 6 (INB), F. González 442 (INB), G. Davidse 28452 (INB, Cyathea schiedeana (C.Presl) Domin MO ). Terrestrial trees. Distribution in PILA 400–1500 m, low elevation mixed forest, mixed forest in transition to low

Cyathea albomarginata R.C.Moran elevation forest, mixed forest in transition to cloud forest. Terrestrial shrubs. Distribution in PILA 2200– 2200 m, Global distribution: Mexico to Colombia. Records: D. low elevation oak forest. Global distribution: Endemic to Santamaría 6648 (BM, INB, MO, PMA), G. Herrera Costa Rica. Records: J. Bittner 1765 (INB, MO). 5329 (INB, MO), J. Bittner 1431 (MO).

Cyathea caracasana (Klotzsch) Domin Cyathea suprastrigosa (Christ) Maxon Terrestrial trees. Distribution in PILA 1600–2200 m, Terrestrial trees. Distribution in PILA 1300–3300 m, mixed forest in transition to cloud forest, cloud forest, mixed forest in transition to cloud forest, cloud forest, low low elevation oak forest. Global distribution: Costa elevation oak forest, páramo. Global distribution: Costa Rica, Colombia to the Guyanas. Records: A. Fernández Rica to Colombia. Records: A.K. Monro 5461 (BM, INB, 715 (INB), A. Rojas 3216 (CR, INB), G. Davidse 23998 MO, PMA), E. Mora 289 (CR, INB), G. Davidse 28679 (CR). (CR, MO), G. Davidse 29247 (MO), F. Quesada 1560 (INB), M. Moraga 337 (INB).

Cyathea delgadii Sternb. Terrestrial trees. Distribution in PILA 0–1800 m, low Lophosoria quadripinnata (J.F.Gmel.) C.Chr. elevation mixed forest, mixed forest in transition to low Terrestrial herbs. Distribution in PILA 1100–2900 m, elevation forest, mixed forest in transition to cloud forest, mixed forest in transition to low elevation forest, mixed cloud forest. Global distribution: Costa Rica to Argentina. forest in transition to cloud forest, cloud forest, low Records: A. Chacón 277 (CR, INB, MO). elevation oak forest, high elevation oak forest. Global distribution: Mexico, Honduras to El Salvador, Costa

Cyathea divergens Kunze Rica to Brazil, Chile, Lesser Antilles, Greater Antilles. Terrestrial trees. Distribution in PILA 1200–2400 m, Records: A.K. Monro 5536 (BM, INB, MO, PMA), G. mixed forest in transition to low elevation forest, mixed Davidse 28706 (MO), G. Davidse 28834 (CR), G. Herrera forest in transition to cloud forest, cloud forest, low 3733 (INB), J. Bittner 1760 (CR), J. Bittner 1474 (MO), elevation oak forest. Global distribution: Mexico to the F. Quesada 1395 (CR), F. Quesada 1957 (INB). Guyanas. Records: J. Bittner 1440 (MO), M. Moraga 650 (INB). Lophosoria quesadae A.Rojas Terrestrial herbs. Distribution in PILA 2300–2900 m, low

Cyathea fulva (M.Martens & Galeotti) Fée elevation oak forest, high elevation oak forest. Global Terrestrial trees. Distribution in PILA 1100–2900 m, distribution: Endemic to Costa Rica. Records: F. Quesada mixed forest in transition to low elevation forest, mixed 1396 (CR), M. Moraga 305 (CR). forest in transition to cloud forest, cloud forest, low elevation oak forest, high elevation oak forest. Global Loxsomopsis pearcei (Baker) Maxon distribution: Mexico to Venezuela. Records: A.K. Monro Terrestrial herbs. Distribution in PILA 2300–2600 m, low 5535 (BM, INB, MO, PMA), A. Rodríguez 10684 (BM, elevation oak forest, high elevation oak forest. Global INB, MO, PMA), J. Bittner 1918 (INB, MO). distribution: Costa Rica, Colombia to Bolivia. Records: M. Grayum 10329 (INB), M. Moraga 339 (CR).

Global distribution: Mexico to Brazil, Greater Antilles,

Sphaeropteris brunei (Christ) R.M.Tryon Lesser Antilles , Africa, Asia, New Zealand, Tasmania.

Terrestrial shrubs or trees. Distribution in PILA 500–2000 Records: A.K. Monro 4140 (BM, INB, MO), G. Davidse

m, low elevation mixed forest, mixed forest in transition 23882 (CR, MO), G. Davidse 26209 (MO), G. Davidse to low elevation forest, mixed forest in transition to cloud 28589 (CR, MO), G. Davidse 28671 (MO), F. Quesada forest. Global distribution: Costa Rica to Colombia. 1393 (CR, INB, MO), L. Angulo 244 (INB, MO), M.A.

Records: A. Fernández 842 (INB), A. Rodríguez 10893 Thomas 1581 (MO).

(BM, INB, MO, PMA). Hypolepis blepharochlaena Mickel & Beitel

Terrestrial herbs. Distribution in PILA 2600–2700 m, high

DENNSTAEDTIACEAE elevation oak forest, sabana natural grassland. Global distribution: Mexico to Panama. Records: M. Moraga 469

Blotiella lindeniana (Hook.) R.M.Tryon (CR, INB) .

Facultatively epiphytic herbs. Distribution in PILA

1000–2000 m, mixed forest in transition to low elevation Hypolepis bogotensis H.Karst.

forest, mixed forest in transition to cloud forest. Global Terrestrial herbs. Distribution in PILA 2400–3300 m, low

distribution: Costa Rica to Brazil, Greater Antilles, Lesser elevation oak forest, páramo. Global distribution: Mexico

Antilles. Records: A.K. Monro 5469 (BM, INB, MO, to Venezuela, Greater Antilles. Records: A. Rojas 3229

PMA), A. Rodríguez 11034 (BM, INB, MO, PMA), G. (INB, MO), A. Rojas 3230 (INB).

Herrera 2844 (MO), J. Bittner 1465 (MO), J. Bittner

1758 (INB). Hypolepis ditrichomatis R.C.Moran Terrestrial herbs. Distribution in PILA 1400–2200 m,

Dennstaedtia arborescens (Willd.) Ekman ex Maxon mixed forest in transition to cloud forest, cloud forest,

Terrestrial herbs. Distribution in PILA 100–2600 m, low low elevation oak forest. Global distribution: Costa Rica,

elevation mixed forest, mixed forest in transition to low Panama. Records: J. Bittner 1888 (INB, MO), M. Grayum

elevation forest, mixed forest in transition to cloud forest, 10319 (INB, MO).

cloud forest, low elevation oak forest, high elevation oak forest. Global distribution: Mexico to Honduras, Costa Hypolepis grandis Lellinger

Rica to Bolivia. Records: G. Herrera 3028 ( MO), J. Terrestrial herbs. Distribution in PILA 0–3000 m, low

Bittner 1490 (MO). elevation mixed forest, mixed forest in transition to low elevation forest, mixed forest in transition to cloud forest,

Dennstaedtia auriculata H.Navarrete & B.Ollg. cloud forest, low elevation oak forest, high elevation oak

Terrestrial herbs. Distribution in PILA 0–700 m, low forest, sabana natural grassland. Global distribution: Costa

elevation mixed forest, sabana natural grassland. Global Rica, Panama. Records: G. Davidse 28677 (CR, MO).

distribution: Costa Rica to Bolivia, Greater Antilles.

Records: M. Moraga 484 (INB). Hypolepis pulcherrima Underw. & Maxon

Terrestrial herbs. Distribution in PILA 1900–3300 m,

Dennstaedtia bipinnata (Cav.) Maxon cloud forest, low elevation oak forest, páramo. Global

Terrestrial herbs. Distribution in PILA 0–2200 m, low distribution: Mexico to Panama, Greater Antilles, Lesser

elevation mixed forest, mixed forest in transition to low Antilles. Records: G. Davidse 26210 (MO).

elevation forest, mixed forest in transition to cloud forest,

cloud forest, low elevation oak forest. Global distribution: Hypolepis trichobacilliformis R.C.Moran

USA to Bolivia, Lesser Antilles, Greater Antilles. Records: Terrestrial herbs. Distribution in PILA 1600–2200 m,

G. Herrera 5511 ( INB). mixed forest in transition to cloud forest, cloud forest,

low elevation oak forest. Global distribution: Costa Rica,

Dennstaedtia distenta (Kunze) T.Moore Panama. Records: F. Quesada 1438 ( INB).

Terrestrial herbs. Distribution in PILA 1800–2200 m,

cloud forest, low elevation oak forest. Global distribution: Hypolepis viscosa Mett.

Mexico to Panama, Greater Antilles, Lesser Antilles. Facultatively epiphytic herbs. Distribution in PILA

Records: C. Hamilton 3614 ( MO). 400–2500 m, low elevation mixed forest, mixed forest in transition to low elevation forest, mixed forest in

Histiopteris incisa (Thunb.) J.Sm. transition to cloud forest, cloud forest, low elevation oak

Terrestrial herbs. Distribution in PILA 1200–3300 m, forest. Global distribution: Mexico to Venezuela, Bolivia.

mixed forest in transition to low elevation forest, mixed Records: G. Herrera 5246 ( MO).

forest in transition to cloud forest, cloud forest, low elevation oak forest, páramo, sabana natural grassland.