Mecyclothorax picdupinsensis Liebherr, 2018

Liebherr, James K., 2018, Cladistic classification of Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) and taxonomic revision of the New Caledonian subgenus Phacothorax Jeannel, Deutsche Entomologische Zeitschrift 65 (1), pp. 1-63 : 43-44

publication ID	https://dx.doi.org/10.3897/dez.65.21000
publication LSID	lsid:zoobank.org:pub:73DEE0F3-2BB0-4A21-B445-5E168FE50F54
persistent identifier	https://treatment.plazi.org/id/469A110B-37CD-48A9-B118-E78BED4D6367
taxon LSID	lsid:zoobank.org:act:469A110B-37CD-48A9-B118-E78BED4D6367
treatment provided by	Deutsche Entomologische Zeitschrift by Pensoft
scientific name	Mecyclothorax picdupinsensis Liebherr
status	sp. n.

Treatment

15. Mecyclothorax picdupinsensis Liebherr sp. n. Figures 20H View Figure 20 , 21L-N View Figure 21 , 22B View Figure 22 , 23F View Figure 23 , 24E View Figure 24 , 25 View Figure 25

Diagnosis.

This final member of the species triplet also including M. mouensis and M. kanak can be diagnosed from the former by characters listed in the M. kanak species diagnosis. As stated above, this species cannot be diagnosed from M. kanak based on external characters. The basis for species recognition lies both in the form of the male aedeagal median lobe apex-i.e., broadly rounded with a minute hitch present (Fig. 21L View Figure 21 )-and female gonocoxae with a broad mediobasal patch covered with quadrate cuticular scales (Fig. 24E View Figure 24 ). Also, based on specimens at hand, the female bursa copulatrix is shorter and broader in this species, and the helminthoid sclerite is more narrowly extended as a laminar structure that is heavily melanized (Fig. 23F View Figure 23 ). Standardized body length 3.03.4 mm. Chaetotaxy +/+//+/-//+/2/+/+.

Description

(n = 5). The description of M. kanak can serve to describe this species, with substitution of the following ranges of ratios of the measured specimens: ocular ratio 1.44-1.52; ocular lobe ratio 0.71-0.80; EyL/EyD = 2.08-2.38; MPW/BPW = 2.30-2.44; MPW/PL = 1.33-1.38; APW/BPW = 1.32-1.36; MEW/EL = 0.85-0.89.

Male genitalia (n = 2). Antecostal margin of abdominal mediotergite IX angulate, not extended (Fig. 21M View Figure 21 ); right paramere narrow, evenly narrowed from narrow base to whiplike apex, with one seta near midlength on ventral margin complementing two long apical setae (Fig. 22B View Figure 22 ); left paramere narrow basally, evenly narrowed to whiplike apex, two longer apical setae present; aedeagal median lobe robust, broad dorsoventrally, evenly curved dorsally and ventrally to a broadly downturned apex which bears a variously developed apical invagination, present as a small tubercle or change of angle of the apical margin (Fig. 21L View Figure 21 ); aedeagal median lobe internal sac with sclerotized region in position of dorsal plate, internal sac folded longitudinally toward apex (Fig. 21L, N View Figure 21 ).

Female reproductive tract (n = 1). Bursa copulatrix length subequal to circumference, its surface thin, translucent (Fig. 23F View Figure 23 ); spermathecal duct entering at apex of bursa separate from bursa-common oviduct juncture; rounded laminar helminthoid sclerite extended from bursal wall between, and separate from both, bursal-common oviduct juncture and spermatheca; spermatheca of only slightly greater circumference than spermathecal duct, reservoir about 1/2 length of spermathecal duct, spermathecal gland duct entering at base of spermathecal reservoir (Fig. 23F View Figure 23 ); ligular apophysis present on common oviduct; basal gonocoxite 1 with apical fringe of two setae laterally, a very small setae may be present immediately mesad, and several smaller setae occur near apex of medial margin (Fig. 24E View Figure 24 ); medial margin bearing a dense field of mi crotrichia basally, this field about 8 microtrichia maximal breadth, with the macrotrichia appearing like flattened scales; gonocoxite 2 moderately narrow basally, basal width 0.4 × medial length; two gracile lateral ensiform setae of moderate length present, the apical seta broader.

Types - Holotype male (MNHN): NEW CALEDONIA 11787 / 22°15 ’Sx166°49’, 280m / Pic du Pin, site 1,rainfor. / 26Nov2004, berlesate / G.Monteith,P.Grimbacher // QUEENSLAND / MUSEUM LOAN / DATE: July 2005 / No. LE 05.24 // New Caledonia Mecyclothorax revision / measured specimen 1 // HOLOTYPE / Mecyclothorax / Mecyclothorax picdupinsensis / J.K.Liebherr 2017 (black-bordered red label) /

Paratypes (11 specimens). NEW CALEDONIA: Pic du Pin[code PA 48], 22°15'S 166°49'E, 28-vi-1970, Franz (NHMW, 3), base, 280 m el., 22°14.9'S 166°49.7'E, sifted rainforest, litter, 26-xii-2006, Wanat & Dobosz (MNHW, 1), 22°15'S 166°49'E, sifted litter, 22-x-2008, Wanat (MNHW, 1), site 1, rainforest, 280 m el., 22°15'S 166°49'E, berlesate, 26-xi-2004, lot 11787, Monteith & Grimbacher (QMB, 5), 21-xii-2004, lot 12045, Monteith (QMB, 1).

Etymology.

This species name represents the genitive adjectival form of the type locality, Pic du Pin; Mecyclothorax picdupinsensis . This final species name includes reference to the New Caledonian Pine, Araucaria columnaris J. R. Forster and W. J. Hooker, and partially recapitulates the final species named in Liebherr (2016); Cyphocoleus iledespinsensis Liebherr.

Distribution and habitat.

This species is restricted to the vicinity of Pic du Pin, southern Grande Terre (Fig. 25 View Figure 25 ), with all collecting localities recorded at 280 m elevation. Nine of the 12 specimens were collected from sieved litter, whereas three specimens (NHMW) are associated with the collecting note: "Total stehendes Stammen von Aruacaria biramulata (Herbert Franz field notebook: Jäch, pers. comm.)": Thus this species may climb the lower reaches and boles of larger trees in addition to occupying the ground-level litter layer.