Sphictostethus papua Loktionov & Lelej

Loktionov, Valery M. & Lelej, Arkady S., 2018, Discovery of the spider wasp genus Sphictostethus Kohl, 1884 (Hymenoptera: Pompilidae: Pepsinae) in New Guinea, with description of two new species, Zootaxa 4429 (2), pp. 295-302: 296-298

publication ID

https://doi.org/10.11646/zootaxa.4429.2.5

publication LSID

lsid:zoobank.org:pub:1893E602-6ED1-433A-B1EC-A1732E0D1347

persistent identifier

http://treatment.plazi.org/id/EF4BC719-FFDB-002E-FF63-FD48022199D8

treatment provided by

Plazi

scientific name

Sphictostethus papua Loktionov & Lelej
status

sp. nov.

Sphictostethus papua Loktionov & Lelej  , sp. nov.

( Figs 1–15 View Figure )

Material examined. Holotype, ♀, Indonesia, West Papua, Arfak Mts, Duebei env[irons], 20 km S Wamere, 1190 m, Manokwari distr. , 21.I–8.II.2009 (S. Jákl) [OLL]. 

Diagnosis. Female. Head in frontal view with vertex straight between eye tops ( Fig. 1 View Figure ). Clypeus in anterior half conspicuously convex, vaulted forwards, its apical rim swollen, forming flattened surface below, curved in arc and inclined towards mentum ( Figs 2, 3 View Figure ); anterior margin of clypeus almost straight ( Fig. 1 View Figure ). Protibia with distinct elongate and stout spine dorsoapically ( Fig. 11 View Figure ). Mesopleuron with distinct conical tubercle posterolaterally ( Fig. 7 View Figure ). Propodeum with small tubercle dorsolaterally ( Figs 7, 9 View Figure ). T1 with long and narrow petiole anteriorly ( Figs 8, 9 View Figure ). Metasoma black with T1 and S1 completely, T2 anterolaterally and anterior half of S2 yellow ( Figs 8, 12 View Figure ). Male. Unknown.

Description. FEMALE. Length: body 10 mm; fore wing 9 mm. Head 1.1 times as wide as height; UID: MIDAbout MID: LID = 32: 45: 50; MIDAbout MID 0.45 times as long as head width in frontal view; half of MIDAbout MID 0.8 times as long as eye width ( Fig. 1 View Figure ). Ocelli large, noticeably raised; ocellar triangle obtuse-angled; POD: OOD = 0.6 ( Fig. 4 View Figure ). Head in frontal view with vertex straight between eye tops ( Fig. 1 View Figure ). Posterior margin of vertex in dorsal view concave medially ( Fig. 4 View Figure ). Frons on both sides of midline barely convex ( Fig. 2 View Figure ). Gena in dorsal view barely developed ( Fig. 4 View Figure ), in profile conspicuously receding toward vertex ( Fig. 2 View Figure ). Malar space short ( Fig. 3 View Figure ). Clypeus 2.1 times as wide as long; anterior half conspicuously convex, vaulted forwards; apical rim very swollen forming flattened surface below curved in arc and inclined towards mentum ( Figs 2, 3 View Figure ); anterolateral corner barely rounded; anterior margin almost straight ( Fig. 1 View Figure ). Mandible with stout subapical tooth. Flagellum filiform; ratio of scape, pedicel and F1–F4 29:11:34:35:42:42; F1 4.3 times as long as maximum width and 1.06 times as long as UID; apical flagellomere pointed apically.

Mesosoma. In dorsal view subparallel, 2.3 times as long as maximum width. Pronotum convex posterolaterally; posterior border deeply angulate ( Fig. 5 View Figure ). Discs of mesoscutum, mesoscutellum and metanotum convex. Metanotum medially 1.7 times as long as metapostnotum medially ( Fig. 7 View Figure ). Metapostnotum with longitudinal depression medially; posterior margin arcuately emarginate. Mesopleuron with distinct conical tubercle posterolaterally ( Fig. 7 View Figure ). Propodeum evenly convex with small tubercle dorsolaterally ( Figs 7, 9 View Figure ).

Legs. Protibia with distinct elongate and stout spine dorsoapically ( Fig. 11 View Figure ). Metatibia barely serrate dorsally ( Fig. 10 View Figure ). Longest spur of metatibia 0.6 times as long as metatarsomere 1. Ratio of protarsomeres 1–5 52:21:16:8:18. Protarsomere 1–3 ventrally with median row of short spines and short scattered spines on each side of median row; protarsomere 4 with few spines ventroapically; protarsomere 5 symmetrical without spines ventrally. Meso- and metatarsomeres 1–4 with spines as on corresponding protarsomeres but spines longer and denser; meso- and metatarsomere 5 without spines ventrally. Orbicula with 8–10 radiating long bristles; longest bristle 0.75 times as long as claw. Tarsal claws symmetrical with large inner tooth ( Fig. 13 View Figure ).

Wings. Fore wing barely infuscated with two wide dark bands and dark apical portion; vein M completely ending shortly beyond 3rs-m vein and not continuous as a spectral vein; venation as in Fig. 14 View Figure . Hind wing slightly infuscated with dark apical portion; venation as in Fig. 15 View Figure .

Metasoma. T1 with long and narrow petiole anteriorly ( Figs 8, 9 View Figure ). S2 with small arcuate depression anteromedially and deep transverse arcuate groove medially ( Fig. 12 View Figure ).

Sculpture. Head and mesosoma matt, metasoma polished. Frons, discs of scutum and scutellum with denser fine punctures. Pronotum and disc of metanotum with hardly visible punctures. Metapostnotum smooth with one transverse stria anteriorly. Propodeum with micropunctures anteriorly and posteriorly and fine transverse furrows medially. T1–T5, S1 and S2 with scattered micropunctures. T6 and S3–S6 with scattered pits. Antenna matt. Legs mostly matt with femur polished ventrally.

Colour. Head black; narrow yellowish strip along inner orbit ( Fig. 1 View Figure ); labrum and clypeus yellow brown anteriorly; antenna yellow brown ventrally. Mesosoma completely black. Metasoma black with T1 and S1 completely, T2 anterolaterally and anterior half of S2 yellow ( Figs 8, 12 View Figure ). Legs from black to dark brown. Frons, vertex, mesoscutum, mesoscutellum and disc of propodeum with bluish green metallic sheen.

Pubescence. Body without setae except following: upper frons with 1–2 pale long erect setae along orbit; anterior portion of clypeus with few light brown setae; mandible and labrum with few long and hard brown setae; prementum with few long erect barely curved bristles ( Fig. 6 View Figure ); propleuron with few long pale setae; disc of scutum with few scattered and short brown setae; propodeum laterally with short silver setae; procoxa with few pale setae; T6 and S6 with long brown suberect setae. Lower face, clypeus, gena, pronotum and propleuron laterally, metanotum and mesopleuron laterally with denser silver pubescence. Disc of propodeum with spot of silver pubescence on each side posterolaterally. Legs with gray micropubescence. Metasoma without pubescence.

MALE. Unknown.

Comparison. The female of the species is similar to that of Sphictostethus gadali Krogmann & Austin, 2011  , S. haoae Krogmann & Austin, 2011  and S. picadillycircus Krogmann & Austin, 2011  from Australia and S. fugax ( Fabricius, 1775)  and S. nitidus ( Fabricius, 1775)  from New Zealand in having distinct elongate and stout dorsoapical spine on the protibia ( Fig. 11 View Figure ), but it can be easily distinguished by the following characters: head in frontal view with vertex straight between top of eyes ( Fig. 1 View Figure ) vs distinctly convex in S. gadali  , S. haoae  and S. picadillycircus  ; specialized shape of clypeus which in anterior half conspicuously convex, vaulted forwards, with its apical rim very swollen, forming a flattened surface below that is curved in an arc and inclined towards the mentum ( Figs 2, 3 View Figure ), its anterior margin almost straight ( Fig. 1 View Figure ) vs clypeus barely convex and not forming a flattened surface below that is curved in an arc at the apical rim, its anterior margin barely or conspicuously emarginate in S. gadali  , S. haoae  and S. picadillycircus  and S. nitidus  ; petiole of T1 well-developed ( Fig. 9 View Figure ) vs barely developed in S. gadali  , S. haoae  , S. picadillycircus  and S. nitidus  ; mesopleuron with distinct conical tubercle posterolaterally ( Fig. 7 View Figure ) vs without any tubercle in S. fugax  and S. nitidus  or with indistinct tubercle in S. gadali  , S. haoae  and S. picadillycircus  ; propodeum with small tubercle dorsolaterally ( Figs 7, 9 View Figure ) vs without any tubercle in S. gadali  , S. haoae  , S. picadillycircus  and S. fugax  or with large somewhat flattened tubercle giving the propodeum a hexagonal outline in dorsal view in S. nitidus  .

Distribution. New Guinea: Indonesia (West Papua).

Etymology. The new species is named after the indigenous Papuan people living in New Guinea. The specific epithet is to be treated as a noun in apposition.

Remark. The true data on the label of the holotype should be 21.I–8.II.2008 (pers. comm. of S. Jákl).

MID

Middlebury College