RAPHITOMIDAE Bellardi, 1875

Coan, Eugene V., Pacaud, Jean-Michel & Kabat, Alan R., 2024, Gérard-Paul Deshayes (1796 - 1875) and his taxa, Geodiversitas 46 (16), pp. 553-930 : 657-658

Family RAPHITOMIDAE Bellardi, 1875

alba , Pleurotoma – Deshayes, 1863: 110-111, pl. 12, figs 17-18. — Type locality: La Réunion. — Type age: Recent. — Current status: Raphitoma alba (Deshayes, 1863) , but taxon inquirendum (MolluscaBase).

attenuata , Pleurotoma – Deshayes, 1834-DescrCoqFoss: 483-484, 36, pl. 68, figs 6-8, non J. Sowerby, 1816; 1865-DescrCoqFoss: 386. — Type locality: Retheuil. — Type age: Eocene (Ypresian). Pleurotoma subattenuata d’Orbigny, 1850b (d’Orbigny 1850b: 316, no. 340), replacement name. Raphitoma (Raphitoma) subattenuata (d’Orbigny, 1850) (Pacaud 2007: 48) . — Current status: Amblyacrum subattenuatum (d’Orbigny, 1850) (Pacaud herein).

baudoni , Pleurotoma – Deshayes, 1865-DescrCoqFoss: 393, 96, pl. 96, figs 25-26. — Type localities: Thiverval-Grignon, Mouchy-le-Châtel & Coincourt. — Type age: Eocene (Lutetian). Raphitoma (Raphitoma) costellata [Lamarck, 1804] baudoni (Deshayes, 1865) ( Le Renard & Pacaud 1995: 126) . Raphitoma (Raphitoma) baudoni (Deshayes, 1865) ( Pacaud 2008: 68) . — Current status: Amblyacrum costellatum baudoni (Deshayes, 1865) (Pacaud herein).

capellinii, Pleurotoma – Deshayes, 1865-DescrCoqFoss: 391, 96, pl. 96, figs 27-28. — Type locality: La Ferté-sous-Jouarre. — Type age: Eocene (Bartonian). Raphitoma (Raphitoma) costellata [Lamarck, 1804] capellinii (Deshayes, 1865) ( Le Renard & Pacaud 1995: 126). — Current status: Amblyacrum costellatum capellinii (Deshayes, 1865) (Pacaud herein).

carinata , Pleurotoma – Deshayes, 1834-DescrCoqFoss: 489-490, 36, pl. 66, figs 26-29, ex Defrance ms, non “ Pleurotome ” [incorrect subsequent spelling] carinata Link, 1808 . — Type localities: Thiverval-Grignon, Parnes, Mouchy-le-Châtel & Chaumont-en-Vexin. — Type age: Eocene (Lutetian). Raphitoma (Raphitoma) defrancei Tucker & Le Renard, 1993 ( Tucker & Le Renard 1993: 2), replacement name ( Le Renard & Pacaud 1995: 126; Pacaud 2008: 68, as “Defrance in Deshayes”). Both preoccupy Pleurotoma carinata J. E. Gray, 1834 , and Pleurotoma carinata Bivona, 1838 . — Current status: synonym of Amblyacrum dameriaciense (Deshayes, 1865) , making the replacement name unnecessary (Pacaud 2021: 43).

clandestina , Pleurotoma –Deshayes, 1863: 110, pl. 12, figs 15-16. — Type locality: La Réunion. — Type age: Recent. — Type material: MNHN-IM-2000-2841, syntype. — Current status: synonym of Kermia pumila (Mighels, 1845) [ Pleurotoma ] ( Kilburn 2009: 230- 231, figs 34-38).

columnella , Pleurotoma – Deshayes, 1865-DescrCoqFoss: 396, 96, pl. 96, figs 19-20. — Type locality: Le Fayel. — Type age: Eocene (Bartonian). Raphitoma (Amblyacrum) perplexa [ Tucker & Le Renard, 1993] columnella (Deshayes, 1865) ( Le Renard & Pacaud 1995: 126) , an error in rank. — Current status: synonym of Amblyacrum perplexum (Deshayes, 1865) (Pacaud herein).

dubia , Pleurotoma – Deshayes, 1834-DescrCoqFoss: 481, 36, pl. 67, figs 12-14, ex Defrance ms, non De Cristofori & Jan, 1832. — Type locality: Thiverval-Grignon. — Type age: Eocene (Lutetian). — Current status: Crassispira (Crassispira) tuckeri Le Renard, 1994 ( Le Renard 1994: 38), replacement name ( Pacaud 2008: 71).

elongata , Pleurotoma – Deshayes, 1834-DescrCoqFoss: 484, 36, pl. 68, figs 4-5, 9, non Deshayes, 1834 [ Conorbidae ]; 1837-DescrCoqFoss: 812, 814, species renamed P. striolaris Deshayes, 1837 (see below); 1865-DescrCoqFoss: 387– Type locality: Retheuil. — Type age: Eocene (Ypresian).

fragilis , Pleurotoma – Deshayes, 1834-DescrCoqFoss: 480-481, pl. 67, 36, figs 25-27; 1843-HistNatAnim2: 489, possibly synonym of Pleurotoma striatulatus (Lamarck, 1803) [ Fusus ]; 1843-HistNatAnim3: 631; 1865-DescrCoqFoss: 388. — Type locality: Thiverval-Grignon. — Type age: Eocene (Lutetian). Preoccupies Pleurotoma fragilis Reeve, 1845 , which became a synonym of Daphnella interrupta Pease, 1860 .Sherborn (1926: 2505) misunderstood Deshayes’ (1843-HistNatAnim: 489) footnote and listed Pleurotoma striatulata as a new species by Deshayes. — Current status: Pleurotomella (Anomalotomella) fragilis (Deshayes, 1834) ( Le Renard & Pacaud 1995: 126; Pacaud 2008: 69).

margaritula , Pleurotoma – Deshayes, 1834-DescrCoqFoss: 479- 480, 36, pl. 67, figs 8-11; 1865-DescrCoqFoss: 386. — Type localities: Thiverval-Grignon, Parnes & Mouchy-le-Châtel. — Type age: Eocene (Lutetian). — Current status: Crassispira (Tripia) margaritula (Deshayes, 1834) ( Le Renard & Pacaud 1995: 125; Pacaud 2008: 71). nana , Pleurotoma – Deshayes, 1834-DescrCoqFoss: 482-483, 37, pl. 68, figs 19-22; 1865-DescrCoqFoss: 387. — Type locality: Parnes. — Type age: Eocene (Lutetian). Preoccupies P. nana Deshayes, 1835 [ Turridae ; see below], P. nana Scacchi, 1836 , and P. nana Thiele, 1925 . Scacchi’s species was renamed Bella menkhorsti Aartsen, 1988 . Thiele’s species is a synonym of Micropleurotoma melvilli (Sykes, 1906) . — Current status: Crassispira (Tripia) nana (Deshayes, 1834) ( Le Renard & Pacaud 1995: 125; Pacaud 2008: 71, as “1865”).

perplaxa, Pleurotoma – Deshayes, 1865-DescrCoqFoss: 395, 99, as Pleurotoma perplexa , pl. 99, figs 10-12; 666 [text should have been spelled perplexa ]. — Type locality: Thiverval-Grignon. — Type age: Eocene (Lutetian). Raphitoma (Amblyacrum) perplexa (Deshayes, 1865) ( Tucker & Le Renard 1993: 38; Le Renard & Pacaud 1995: 126; Pacaud 2008: 68). — Current status: Amblyacrum perplexum (Deshayes, 1865) ( Tréguier & Pacaud 2018: 211, 212, pl. 4, fig. 4).

perplexa , Pleurotoma – see entry above.

quantula , Pleurotoma – Deshayes, 1865-DescrCoqFoss: 394, 99, pl. 99, figs 4-6. — Type locality: Chaussy, Parnes & Saint-Félix. — Type age: Eocene (Lutetian). Raphitoma (Raphitoma) costellata [Lamarck, 1804] quantula (Deshayes, 1865) ( Le Renard & Pacaud 1995: 126) . — Current status: Raphitoma (Raphitoma) quantula (Deshayes, 1865) ( Pacaud 2008: 68) .

reeveana , Pleurotoma – Deshayes, 1863: 106-107, pl. 12, figs 5-7. — Type locality: La Réunion. — Type age: Recent. — Type material: MNHN-IM-2000-3218, syntype. — Current status: Daphnella reeveana (Deshayes, 1863) (B. Li & X. Li 2014: 12, fig. 3E; Hasegawa & Okutani, in Okutani 2017: 1028, pl. 321, fig. 5; Poppe 2017: 566, pl. 1583, figs 1-2; Boutet et al. 2020: 449).

rugosa , Pleurotoma – Deshayes, 1834-DescrCoqFoss: 486-487, 35, pl. 66, figs 20-21, non I. Lea , 1833. — Type locality: Parnes. — Type age: Eocene (Lutetian). — Type species (OD) of Amblyacrum Cossmann, 1889 . Paphitoma gougeroti Tucker & Le Renard, 1993 ( Tucker & Le Renard 1993: 2), replacement name. Raphitoma (Amblyacrum) gougeroti ( Tucker & Le Renard 1993) ( Le Renard & Pacaud 1995: 126; Pacaud 2008: 68). — Current status: Amblyacrum gougeroti ( Tucker & Le Renard 1993; Pacaud 2021: 43).

striolaris , Pleurotoma – Deshayes, 1834-DescrCoqFoss: 812, replacement name for Pleurotoma elongata Deshayes, 1834 , non Deshayes, 1834 [ Conorbidae ]. — Type locality: Retheuil. — Type age: Eocene (Ypresian). — Current status: Raphitoma (Raphitoma) striolaris (Deshayes, 1834) ( Le Renard & Pacaud 1995: 126) .