Scipopus (Phaeopterina) brunneus, Lindsay & Marshall, 2023

Lindsay, Kate & Marshall, Stephen A., 2023, A revision of Scipopus Enderlein including the subgenera Scipopus s. str., Phaeopterina Frey and Parascipopus subgen. nov. (Diptera, Micropezidae, Taeniapterinae), European Journal of Taxonomy 904, pp. 1-189 : 120-122

Scipopus (Phaeopterina) brunneus sp. nov.

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Fig. 50

Differential diagnosis

Scipopus (Phaeopterina) brunneus sp. nov. resembles S. (Ph.) sexguttatus and S. (Ph.) musculosus sp. nov. in having three subapical hyaline spots on the wing and a mostly white first fore tarsomere. Scipopus (Ph.) brunneus can be differentiated from these by its dark brown clypeus, antenna and parafacial.

Etymology

The species name, from the Latin noun for ‘brown’, refers to the brown parafacial and clypeus of this species.

Type material examined

Holotype

COLOMBIA • 1 ♀; Caqueta San José de Fragua R Yuruyaco; 1°20′55″ N, 76°06′11″ W; 1300 m a.s.l.; 9–13 Sep. 2000; E. Gonzalez leg.; Malaise 5; debu 01089018/MYCRO600-19 sequenced for CO1–5′, CO1–3′, 12S, 28S; IAVH (dissected and photographed, Fig. 50A–D ).

Description

LENGTH. 12 mm.

HEAD. Palpus light brown, darker apically, entirely pale microtrichose and setulose on ventral margin, narrow (length 5.3× height). Antenna and parafacial dark brown in ground colour, white microtrichose. Clypeus dark brown, width ~2.0× height, silvery microtrichose on entire surface, denser in posterolateral corners. Frontal vitta dull, orange, microtrichose, strongly tapered posteriorly. Orbital plate orange, microtrichose, bare and slightly shiny on lateral edges. Epicephalon light brown, shiny, white microtrichose except bare anteriorly, wide (width ⅔ or more of upper frontal vitta width at inner verticals), clearly delineated from upper frontal vitta. Postocellar seta absent; all other head chaetotaxy well-developed.

THORAX. Scutum black-brown with wide, blue median sheen. Postpronotal lobe black-brown, densely setulose on anterior half. Notopleuron black-brown, pale brown microtrichose ventromedially and posteriorly. Female cervical sclerite very slightly swollen posteriorly. Pleuron dark brown with blue sheen, completely silvery microtrichose but denser posteriorly, forming silvery transverse band. Legs dark brown; fore tarsomere 1 almost entirely white; hind tarsomere 1 pale basally, with ventral white fringe. Wing brown-black infuscate on r 1, distal ½ and apex, three hyaline window-like spots in m 1, r 2+3, r 3+4.

FEMALE ABDOMEN. T1 with fine, long, dark brown setae. Pleuron off-white, dorsal quarter of P3–5 dark brown. T1+2 ~2.0× length of T3; posterior margin of T2 ~1.6× as wide as T1. Oviscape dark brown, darker apically, anterior ⅔ white microtrichose, ~3.0× length of T6. Common duct very short, smooth, ventral receptacle apparently originating from duct. Paired spermathecal duct (duct leading to large paired spermathecae) arising from bursa, long (>10.0 × length of paired spermathecae), mostly parallel-sided but very swollen distally. Paired spermathecal stems with very short, narrow, basal section and long, bulbous, swollen distal section ( Fig 50C ). Paired spermathecae nearly spherical, sparsely tuberculate especially near base. “Single” spermathecal duct (smaller spermathecal duct, in this case leading to two small spermathecae) short and narrow, ½ diameter and>½ length of paired duct, arising from base of paired duct, terminating in pair of elongate, bulbous and distally rounded spermathecae.

MALE ABDOMEN. Unknown.

Remarks

See S. (Ph.) musculosus sp. nov. for discussion.

Distribution

Colombia.

Figures

Fig. 50. Scipopus (Phaeopterina) brunneus sp. nov. ♀, Colombia, holotype (IAVH). A. Head and thorax, dorsal view. B. Head, anterolateral view. C. Female spermathecae and associated structures. D. Habitus. Abbreviations: pd = paired spermathecal duct; ps = paired spermatheca; sd = single spermathecal duct; ss = single spermatheca; vr = ventral receptacle.