Hycleus amoenus

Bologna, Marco A., Amore, Valentina & Pitzalis, Monica, 2018, Meloidae of Namibia (Coleoptera): taxonomy and faunistics with biogeographic and ecological notes, Zootaxa 4373 (1), pp. 1-141: 67-70

publication ID

https://doi.org/10.11646/zootaxa.4373.1.1

publication LSID

lsid:zoobank.org:pub:202AFD20-7B37-405A-9CBA-051EF24E9FB2

persistent identifier

http://treatment.plazi.org/id/F72687E4-FFD5-FF88-FF5F-F9E5FC124829

treatment provided by

Plazi

scientific name

Hycleus amoenus
status

 

Cb) Hycleus amoenus  species group

This southern African group of species is well distinct by the reduced number of antennomeres (eigth) and molecular analyses ( Pitzalis 2007) pointed out its division in some sub-groups. Because of the antennal feature, the species of this group were erroneously referred to the genus Actenodia  (see Bologna et al. 2008a), but actually they belong to Hycleus  , as pointed out by Bologna & Pinto (2002), and by Pitzalis (2007). This group includes (Pitzalis & Bologna in preparation): H. amoenus  , H. bushmanicus  , H. deserticolus  , H. devylderi  , H. kochi  , H. politus  , H. vansoni  , and the new species H. planitiei  .

Two other eastern African and Arabic species, namely H. afrotropicus ( Kaszab, 1983)  and H. yemenicus ( Kaszab, 1983)  have a similar reduced number of antennomeres (Bologna 1990; Bologna & Turco 2007) and for this reason they were erroneously described as Actenodia ( Kaszab 1983)  . Relationships between these two assemblages of species are not defined, but probably the reduction of antennomeres is only a parallelism more than a synapomorphic condition.

Hycleus amoenus ( Marseul, 1872)  comb. n. ( Fig. 4JView FIGURE4)

Actenodia amoena Marseul, 1872  ; Kaszab 1952

Actenodia amoena ssp. anthicoides Kaszab, 1955b  syn. n.

Types. Types of this species were examined at MNHN, and those of the ssp. anthicoides  at NHP.

Distribution. Botswana, southwestern Namibia  (new species record for this country), and western South Africa.

Material examined. [Karas] Lüderitz: Kaukausib Riverbed , Diamond Area 1, 26.8833°S 15.4167°E (CB; SMWNAbout SMWN)GoogleMaps  ; C13, 4.5 km N Rosh Pinah , 27.82909°S 16.72726°E (CB).GoogleMaps 

Other records: Namibia ( Pitzalis et al. 2014)  .

Remarks. Kaszab (1955b) described the ssp. anthicoides  from the South African Richtersveld, just close to the southwestern Namibian border. After the examination of several specimens of different populations, we consider this subspecies as a synonym of the nominate form.

Hycleus bushmanicus ( Kaszab, 1952)  ( Fig. 4KView FIGURE4)

Actenodia bushmanica Kaszab, 1952 

Distribution. South Namibia  and northwestern South Africa.

Material examined and literature records. [Hardap] Mariental Urban: 11 km W Mariental (road C19), 24.6167°S 17.8500°E (CB). [Karas] Keetmanshoop Rural: D608, 35 km S Keetmanshoop, 26.90790°S 18.26411°E (CB); D608, 87 km S Keetmanshoop, 27.32571°S 18.27616°E (CB); D608, 95.5 km S Keetmanshoop, 27.37944°S 18.22418°E (CB); C12, 33 km W Grunau, 27.57847°S, 18.10557°E (CB). Karasburg: M21, 21.5 km S Karasburg, 28.20340°S 18.72978°E (CB); M21, 4 km N Warmbad, 28.39832°S 18.75766°E (CB); C13, 44.5 km N Noordoewer, Grape Valley, 28.41803°S 17.44408°E (CB); C10, 23 km N Velloorsdrif, 28.52194°S 19.18402°E (CB).

Other records: Namibia  ( Bologna 2000a; Pitzalis et al. 2014).

Remarks. Types of this species were examined at NHP.

Hycleus deserticolus Wellman, 1908  ( Fig. 4LView FIGURE4)

Actenodia deserticola Wellman, 1908  Zonabris (Actenodia) annulipes Pic, 1910  Actenodia annulipes, Kaszab 1955 

Distribution. Southern Angola, Namibia  .

Material examined and literature records. [Kunene] Epupa: Hartmann's Valley, 17.3833°S 12.2500°E (SMWN); Dunes, 17.3833°S 12.2500°E (SMWN); Okakatuwo, 17.4000°S 12.7333°E (SMWN); Okakatuwo, 17.4500°S 12.7000°E (SMWN); Kandao, 17.6167°S 12.5000°E (SMWN); Ondundujengo River, 17.8000°S 12.3167°E (SMWN). Opuwo: 30 km WNW Orupembe, 18.1167°S 12.3500°E (SMWN); Oropembe, 18.1833°S 12.5167°E (SMWN); Kaokoveld, Anabib (Orupembe), 100 miles W Ohopoho, 18.1833°S 12.5167°E ( Kaszab 1956, as ab. anticedisrupta: Kaszab, 1956); Kaokoveld, Orupembe, 18.1833°S 12.5167°E ( Kaszab 1956, as ab. anticedisrupta); Kaokoveld, Sanitatas, 85 miles WSW Ohopoho, 18.2833°S 12.6667°E ( Kaszab, 1956); Kunene distr., C43, 18.9562°S 13.7573°E (CB). Sesfontein: Khowarib River, 19.2667°S 13.8667°E (SMWN); D2650, near Kamanjab, 19.8087°S 14.5128°E (CB). Kamanjab: C35, 35 km NW Kamanjab, 19.5000°S 14.8000°E (CB); C40, 26 km E Kamanjab, 19.6957°S 15.1046°E (CB); C39, 25 km E Khorixas, 20.3061°S 15.1867°E (CB). Khorixas: Bethanis 514. 20.4000°S 14.4000°E (SMWN); Twyfelfontein, 20.5667°S 14.3667°E (SMWN); C35, 54 km N Uis, 20.7788°S 14.8692°E (CB); Duineveld 529, 20.7833°S 14.6333°E (SMWN); Onverwag 412, 20.8370°S 14.9610°E (SMWN). [Kavango] Mashari: Kaukau-Kungveld, Samengaigai, 19.0833°S 20.2000°E ( Kaszab 1956 as ab. anticedisrupta:). [Otjozondjupa] Otjiwarongo: C63, Outjo-Kalkfeld 15 km N of Kalkfeld, 20.7537°S 16.2210°E (CB). [Erongo] Dâures: Brandberg, Hungorob Valley, 21.1900°S 14.5282°E ( Bologna 2000a; SMWN); Brandberg, Messum valley, 21.2215°S 14.5163°E ( Bologna 2000a; SMWN); C35, 2 km E Uis, 21.2325°S 14.8973°E (CB); Gross Spitzkoppe, 21.8167°S 15.1667°E (CB); Spitskoppe, 21.8518°S 15.1484°E (CP; SMWN). Omaruru: near Omaruru, 21.4333°S 15.9333°E (CP). Karibib: D1935, 30 km N jct. B2, 21.7732°S 15.4699°E (CB); D1935, 30 km NNW Usakos, 21.7833°S 15.5000°E (CB); Karibib-Okahandja, 21.9333°S 15.8333°E (CP); B2, 16 km W Karibib, 21.9358°S 15.7056°E (CB); Karibib, 21.9417°S 15.8509°E (CB); Usakos, D1935 6 km N jct. B2, 21.9475°S 15.5851°E (CB); Karibib, D 1953, 100 m jct. C32, 21.9552°S 15.8493°E (CB); B2, 16 km W Karibib, 21.9702°S 15.6868°E (CB); B2, 2, 6 km W Usakos, 21.9989°S 15.5647°E (CB); B2, 2 km W Usakos, 21.9992°S 15.5696°E (CB); Navachab 67, 22.0167°S 15.7333°E (SMWN); D1980, 27 km S Karibib, 22.1500°S 15.8333°E (CB); C32, 42 km S Karibib, 22.29447°S 15.8351°E (CB); D1953, near Karibib, 45 km jct. C32, 22.2963°S 16.0658°E (CB); C32, 54 km S Karibib, Swakop River, 22.3954°S 15.8343°E (CB). Arandis: Rossing Mine, 22.4667°S 15.0333°E (SMWN); N Namib, 22.5667°S 14.7167°E (SMWN); C14, Kuiseb Pass, 23.3045°S 15.7526°E (CB); C14, Kuiseb River, 23.30457°S 15.77184°E (CB). Walvis Bay Rural: Rostock, Windhoek, 23.3833°S 15.7500°E (SMWN); C14, 36 km N Solitaire, 23.5823°S 15.8182°E (CB). [Omaheke] Aminius: C20, Gobabis-Leonardville 5 km S of Aais, 23.2593°S 18.7237°E (CB). [Khomas] Windhoek Rural: Kos 28, 23.2667°S 16.1333°E (SMWN); D1228, 7 km ENE Rehoboth, 23.2833°S 17.2333°E (CB). [Hardap] Rehoboth Rural: B1, 28.5 km S Rehoboth, 23.5833°S 17.1500°E (CB). Gibeon: C19, 8 km S Solitaire, 23.9774°S 16.0063°E (CB); Waltevrede Game Farm, 24.1779°S 15.9802°E (CB); C14, 10 km S Waltevrede, 24.2377°S 15.9040°E (CB); Sesriem 137, River, 24.4833°S 15.9500°E (SMWN); C14, 1 km W Maltahöhe, 24.8589°S 16.9751°E (CB); Wereldend 115, 25.1500°S 16.2333°E (SMWN); Gorrasis 99, 25.3184°S 15.9089°E (SMWN). Mariental Urban: C19, 11 km W Mariental, 24.6167°S 17.8500°E (CB). Mariental Rural: M29, 25 km E Mariental, 24.7825°S 18.1612°E (CB); M 29, 118 km S Mariental, 25.4479°S 18.6138°E (CB). [Karas] Keetmanshoop Rural: Khabus 146, 26.2833°S 18.2333°E (SMWN); Khabus 146, 26.3000°S 18.2167°E (SMWN); Karas distr., 27.1352°S 19.4941°E (CB). Keetmanshoop Urban: B1, near Keetmanshoop, 26.5833°S 18.1333°E (CB). Berseba: B4, 73 km W Keetmanshoop, 26.7809°S 17.4657°E (CB). Karasburg: 5 km S Warmbad, 28.4833°S 18.7667°E (SMWN).

Other records: Namib Sand Sea desert ( Seely 2012); Namibia  ( Bologna 2000a; Pitzalis et al. 2014).

Remarks. Types of annulipes  were examined at MNHNAbout MNHN.

Hycleus devylderi ( Borchmann, 1928)  ( Fig. 4MView FIGURE4)

Actenodia devylderi Borchmann, 1928 

Distribution. Namibia  (endemic).

Material examined and literature records. [Kunene] Khorixas: N of Doros Crater, Skeleton Coast Park, 20.3667°S 14.1333°E ( SMWNAbout SMWN). [Otjozondjupa] Otjiwarongo: C63, Outjo-KalkfeldGoogleMaps  , 15 km N of Kalkfeld, 20.7537°S 16.2210°E (CB). Okahandja GoogleMaps  : 30 km W Okahandja, 21.9260°S 16.5768°E (CB; CP). [Erongo] Dâures: Brandberg, Hungorob Mouth, 21.2267°S 14.5167°E ( Bologna 2000a, as sp. aff. devylderi  ; SMWNAbout SMWN); Messum River GoogleMaps  , 21.2500°S 14.4667°E (SMWN); D2342, 17 km SE Messum Valley, 21.3048°S 14.6574°E (CB). Omaruru: C33, 3– 9 km S of Omaruru, 21.4948°S 15.9705°E (CB). Karibib: D2306, 33 km N Usakos, 21.7764°S 15.4578°E (CB); D1935, 30 km NNW Usakos, 21.7833°S 15.5000°E (CB); C33, 17 km jct. B2, 21.8390°S 15.9233°E (CB); B2, 16 km W Karibib, 21.9358°S 15.7056°E (CB); D1953, jct. C32, 21.9552°S 15.8493°E (CB); B2, 8 km W Usakos , 21.9876°S 15.5057°E (CB); B2, 2GoogleMaps  , 6 km W Usakos, 21.9989°S 15.5647°E (CB); D1953, 6 km jct. C32, 22.0078°S 15.8861°E (CB); B2, Swakopmund-Usakos GoogleMaps  , 210 km W Usakos, 22.0199°S 15.5296°E (CB); B2, 52 km W Usakos, 22.1499°S 15.1930°E (CB); D1953, 45 km jct. C32, 22.2963°S 16.0658°E (CB). Arandis GoogleMaps  : 6 km N Arandis, 22.3667°S 14.9833°E ( SMWNAbout SMWN); Upper Ostrich Gorge, 22.4833°S 14.9833°E ( SMWNAbout SMWN); Upper Panner Gorge, 22.4833°S 15.0167°E ( SMWNAbout SMWN). Walvis Bay Rural: C14, 45 km N Solitaire, 23.5364°S 15.7827°E (CB). [Khomas] Windhoek Rural: Windhoek, Heroes Acre, 22.62°S 17.07°E, ( OSUAbout OSU); D1261, Nauchas-Rehoboth GoogleMaps  , 1–40 km NW Nauchas, 23.5763°S 16.4588°E (CB). [Hardap] Gibeon: C14, 10 km S Waltevrede, 24.2377°S 15.9040°E (CB); C14, Maltahohe-Bullsport, after Fish River GoogleMaps  , 24.4022°S 16.8217°E (CB). [Karas] Lüderitz: near Aus, 26.6667°S 16.2667°E (CP).GoogleMaps 

Other records: Damaraland ( Borchmann 1928); Namibia  ( Bologna 2000a, also as “sp. aff. devylderi  ”; Pitzalis et al. 2014).

Remarks. Types of this species were not examined, but specimens compared with types were studied at HMNHAbout HMNH.

Hycleus kochi ( Kaszab, 1952)  ( Fig. 4NView FIGURE4)

Actenodia kochi Kaszab, 1952 

Distribution. Namibia  (endemic).

Material examined and literature records. [Kunene] Epupa: C43, near Opuwo , 17.8554°S 13.7981°E (CB). Opuwo: D3710, near Opuwo, 18.1589°S 13.9173°E (CB); D3710, near Opuwo, 18.2011°S 13.8994°E (CB)GoogleMaps  ; Kunene distr., C43, 18.6788°S 13.7135°E (CB)GoogleMaps  ; Kunene distr., C43, 18.7599°S 13.7489°E (CB); C 35, 110 km NW Kamanjab GoogleMaps  , 18.9167°S 14.3833°E (CB). Outjo: Abachaus , 19.7167°S 16.5800°E ( Kaszab, 1952; SMWNAbout SMWN)GoogleMaps  . Kamanjab: C38, 40 km N Outjo, 19.80883°S 15.91281°E (CB). [Oshikoto] Guinas: C38, 1 km jct. B1, 18.7725°S 17.2491°E (CB). Omuthiyagwiipundi: Site 1/ Row D, Term. / Tamboti forest GoogleMaps  , 18.8725°S 17.0550°E (SMWN). [Otjozondjupa] Grootfontein: D2859, Hoba Meteorite , 19.5833°S 17.9333°E (CB). Otjiwarongo: B1, 13.6 km N OtjiwarongoGoogleMaps  , 20.3728°S 16.7393°E (CB); Hedwigstal 77, 20.8750°S 16.1250°E ( SMWNAbout SMWN)GoogleMaps  . Omatako: 140 km N Okahandja, 20.4925°S 16.9872°E (CP); Waterberg , jct. C22-2512, 20.6333°S 17.1500°E (CB); B1, 28 km S Otjiwarongo, 20.6806°S 16.7787°E (CB); Erindi Osave 170, Otjiwarongo , 20.9800°S 16.7600°E ( SMWNAbout SMWN)GoogleMaps  . Okahandja: Ovita , 21.5228°S 16.2539°E ( SMWNAbout SMWN)GoogleMaps  . [Omaheke] Steinhausen: Alkm ar 512, 21.8700°S 19.8800°E ( SMWNAbout SMWN)GoogleMaps  . [Khomas] Windhoek Rural: Windhoek, D 1535, 5 km jct. B6, 22.3681°S 17.6675°E (CB). [Karas] Keetmanshoop Rural: Noachabeb 97/ Rotegab 95, 27.3833°S 18.4667°E ( SMWNAbout SMWN)GoogleMaps  . Karasburg: Eendoorn 106, 28.7333°S 18.9667°E ( SMWNAbout SMWN)GoogleMaps  .

Other records: Namib Sand Sea desert ( Seely 2012); Namibia  ( Bologna 2000a; Pitzalis et al. 2014).

Remarks. Types of this species were examined at NHP and at HMNHAbout HMNH.

SMWN

State Museum

MNHN

Museum National d'Histoire Naturelle

OSU

Ohio State University

HMNH

Hayashibara Museum of Natural History

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Meloidae

Genus

Hycleus

Loc

Hycleus amoenus

Bologna, Marco A., Amore, Valentina & Pitzalis, Monica 2018

2018
Loc

Hycleus amoenus ( Marseul, 1872 )

Bologna & Amore & Pitzalis 2018

2018
Loc

Actenodia amoena ssp. anthicoides

Kaszab 1955

1955
Loc

ssp. anthicoides

Kaszab 1955

1955
Loc

Actenodia annulipes

, Kaszab 1955

1955
Loc

Hycleus bushmanicus (

Kaszab 1952

1952
Loc

Actenodia bushmanica

Kaszab 1952

1952
Loc

Hycleus kochi (

Kaszab 1952

1952
Loc

Actenodia kochi

Kaszab 1952

1952
Loc

Hycleus devylderi (

Borchmann 1928

1928
Loc

Actenodia devylderi

Borchmann 1928

1928
Loc

Zonabris (Actenodia) annulipes

Pic 1910

1910
Loc

Hycleus deserticolus

Wellman 1908

1908
Loc

Actenodia deserticola

Wellman 1908

1908
Loc

Actenodia amoena

Marseul 1872

1872