Glyptasida Casey 1912

Lockwood, Stephanie A. & Pollock, Darren A., 2009, A review of the genus Glyptasida Casey (Coleoptera: Tenebrionidae: Asidini), Zootaxa 2023, pp. 1-27 : 5-7

publication ID

https://doi.org/ 10.5281/zenodo.186116

DOI

https://doi.org/10.5281/zenodo.5657492

persistent identifier

https://treatment.plazi.org/id/F9076430-FFA5-CC26-4285-FC0CC843FEE5

treatment provided by

Plazi

scientific name

Glyptasida Casey 1912
status

 

Glyptasida Casey 1912

Glyptasida Casey 1912: 95 ; Leng 1920: 224; Wilke 1922: 269; Bradley 1930: 185, 322; Gebien 1937: 704 –705; Blackwelder 1945: 516; Pallister 1954: 18; Papp 1961: 85; Arnett 1968: 651, 673; Brown 1971:19, 27, Fig. 20 View FIGURES 18 – 27 ; Broadus 1974:16, 21, Fig. 12 View FIGURES 10 – 13 ; Blackwelder and Arnett 1975:R 73.22; Lavigne 1980:39,41; Bousquet 1991: 254; Poole and Gentili 1996: 435; Stapp 1997 a: 353; Stapp 1997 b: 304; Arnett 2000: 467; Fagerlund 2000: 36; Aalbu et al. 2002: 488; NMGF 2008; Quinn 2005; NAU 2006.

Type species: Pelecyphorus sordidus LeConte. Fixation : Casey (1912: 75) by original designation.

Derivation of generic name: The name Glyptasida , derived from the Greek glyptos - meaning carved ( Brown 1991: 371) and asida derived from the Old-world generic name Asida , alludes to the “carved-like” texture of the elytra and that the members resemble the genus Asida . Individuals of Glyptasida have been informally called “carved darkling beetles” ( Quinn 2005).

Diagnosis. With general features of Asidini (sensu Aalbu et al. 2002) and the following unique combination of characteristics: Ligula small, hidden under apical margin of mentum; maxillary palpi sexually dimorphic; width of pronotum at basal angles at least 1 / 5 greater than width of elytra at humeri; base of pronotum broadly bisinuate.

Description. Body form. Ovate to ovate-elongate ( Figs. 1, 2, 3 View FIGURES 1 – 9 ); GHW 2.5–4.4 mm; GPW 4.7–9.6 mm; GEW 7.2–13.2 mm; HL 1.9–4.1 mm; PL 3.3–5.5 mm; EL 7.1–18.7 mm; TL 15.3 –27.0 mm; Ratio of GPW/ GEW 0.50–0.95; Ratio TL/GEW 1.3–2.5.

Color. Black, shining to dull; individuals can appear brown due to dirt and debris between the elytral costae.

Head. Trapezoidal, slightly wider than long, widest in front of eyes, slightly concave where clypeus and frons are fused medially ( Fig. 14 View FIGURES 14 – 17 ); frons extended laterally in front of eyes; head covered with setae; antennae inserted ventrally under frons; eyes longitudinal, long and narrow, oriented vertically ( Fig. 16 View FIGURES 14 – 17 ).

Antenna. Clavate, moderately thick, of 11 antennomeres, setose from scape; antennomere 3 longest; antennomere 10 distinctly wider than long; antennomere eleven conical and distinctly smaller than antennomere 10, antennomere 10 with fine sensory setal patches extending laterally, forming two separate patches (on some individual antennae, one patch may be separated to form three total patches) ( Fig. 15 View FIGURES 14 – 17 ); antennomere 11 with circular patch of sensory setae with several long setae extending distally ( Fig. 15 View FIGURES 14 – 17 ).

Mouthparts. Labrum dorsally punctate with short discal setae; apical edge with two patches of setae directed medially. Mandibles large with only medial edges covered by labrum, evenly arcuate to apex ( Fig. 17 View FIGURES 14 – 17 ); mandibles punctate and covered with setae on dorsolateral surface; mola large, subquadrate, anterior edge evenly arcuate, convex from posterior to anterior edge. Two-segmented cardo about one-half length of the stipes, cardo inverted triangular; stipes elongate, two segmented, inner segment not sclerotized; lacinia and galea fringed with dense setae; galea with proximally curved convexly shaped setae, galea narrowed distally; lacinia with setae along medial margin; palpifer present, hinged to allow movement of maxillary palp; palpus of three articles with long setae; palpomere 1 large, concave, allowing for reception of apical palpomere, about one-half length of apical palpomere; palpomere 2 small, about one-third length of apical palpomere, concave, allowing for reception of apical palpomere; palpomere 1 width about twice as wide as apical palpomere; apical palpomere flattened, with extra fine sensory setae distally. Mentum twice as wide as long, widest medially, lateral edges sinuate basally, covered with setae dorsally, apical edge bisinuate, concave medially; ligula not visible behind mentum, ligula about twice as wide as long with anterior edge deeply emarginated, apicolateral corner rounded with one long seta; labial palpi three-segmented; palpomere three widest medially; palpomere two with several long setae; palpomere three with several long setae medially and very fine sensilla apically.

Prothorax. Pronotum large, about 1.75 wider than long (GPW/PL 1.0– 2.1) ( Figs. 1, 2, 3 View FIGURES 1 – 9 ); variable in shape from evenly arcuate to basally sinuate, lateral margins equally arcuate or basally sinuate; prominent hind angles present or absent; lateral edges may have serrations, from just a few to entire edge covered; discal punctures small ( Fig. 11 View FIGURES 10 – 13 ) or large ( Fig. 10 View FIGURES 10 – 13 ), bearing a hair protruding over the puncture, hairs directed from anterior to posterior. Prosternum trapezoidal, convex, widest posteriorly; anterior margin with long setae, projected anteriorly around head, covered with setiferous punctures, setae projected posteriorly; prosternal process large, widest medially, projected ventrally; coxal cavities closed, circular with medial concavity. Proepisternum impunctate to sparsely punctate, sparsely covered with setae.

Pterothorax. Apterous. Mesosternum hexagonal in shape with well developed mesosternal process, concave laterally with prosternal process projecting ventrally, anterolateral margins straight, posterior lateral margins sinuate, covered with long setae, dense near margins; mesosternal process concave medially between closed coxal cavities. Mesoepisternum subquadrate, lateral margins straight, anterior margin concave, posterior margin convex, impunctate, sparsely covered with short setae. Mesoepimeron rectangular, narrow, anterior lateral angle acute, flared anteriorly; surface impunctate, sparsely covered with short setae. Metasternum elongate laterally, narrowest between meso- and meta-coxal openings; anterior margin convex, posterior margin concave, lateral anterior angles acute. Metepisternum impunctate, sparsely covered with setae, separated from metasternum by a fold. Mesosternum and metasternum meet to form tear drop shape middle coxal cavities, metasternum and abdomen meet to form teardrop-shaped hind coxal cavities.

Elytra. Elytra fused, oblong-oval, convex, widest near midlength, Ratio of EL/GEW 0.6–1.7 ( Figs. 1, 2, 3 View FIGURES 1 – 9 ); posteriorly subarcuate, with colliculate microsculpturing ( Figs. 12, 13 View FIGURES 10 – 13 ); two elevated longitudinal costae present ( Figs. 4, 6 View FIGURES 1 – 9 ), reduced in one species ( Fig. 5 View FIGURES 1 – 9 ); elevated transverse sinuous rugose lines between the costae and margins, reduced in one species; setae present, with ( Fig. 12 View FIGURES 10 – 13 ) or without nodes ( Fig. 13 View FIGURES 10 – 13 ) (raised circular areas), setae may be dense (in some specimens); epipleural fold narrow from base to apex.

Legs. Front, middle, and hind legs similar, slender and long; all surfaces with setae; tarsomere arrangement 5 - 5 - 4; tarsal claw arcuate; tibial spurs 2 - 2 - 2, proximal, slightly arcuate; femur enlarged distally; tibia acutely pointed at apex. Coxa globose and projecting, covered with setiferous punctures; trochanter rectitriangular, covered with setiferous punctures. Mesotrochantin not visible.

Abdomen. Five ventrites, covered with setae; defensive glands absent; visible membrane along hind margin of ventrites 3 and 4.

Male genitalia. Aedeagus with tegmen inverted (ventral) ( Fig. 22 View FIGURES 18 – 27 ); setae extended laterally from the apex of the apicale; tegmen greatly narrowed toward the apex; apicale distinctly narrower than the basale. Median lobe elongate, narrow; rounded ( Fig. 20 View FIGURES 18 – 27 ) or flared ( Fig. 24 View FIGURES 18 – 27 ) basally.

Female genitalia. Ovipositor elongate ( Fig. 27 View FIGURES 18 – 27 ); styli small, stylus with small depression with sensilla; internal tract ovate with oviduct extending laterally before the tract narrows; spermatheca extending from the apex, with multiple branches; accessory gland branching off just before the spermatheca ( Fig. 26 View FIGURES 18 – 27 ).

Sexual Dimorphism. Male apical maxillary palpomere enlarged, scalene with proximal angle acute ( Fig. 18 View FIGURES 18 – 27 ); female apical maxillary palpomere smaller and right-triangular ( Fig. 19 View FIGURES 18 – 27 ).

NAU

Nanjing Agricultural University, Department of Plant Science

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Tenebrionidae

Loc

Glyptasida Casey 1912

Lockwood, Stephanie A. & Pollock, Darren A. 2009
2009
Loc

Glyptasida

Aalbu 2002: 488
Arnett 2000: 467
Fagerlund 2000: 36
Stapp 1997: 353
Stapp 1997: 304
Poole 1996: 435
Bousquet 1991: 254
Broadus 1974: 16
Arnett 1968: 651
Papp 1961: 85
Pallister 1954: 18
Blackwelder 1945: 516
Gebien 1937: 704
Bradley 1930: 185
Wilke 1922: 269
Leng 1920: 224
Casey 1912: 95
1912