Sciasmomyia tubata, Shi, Li, Gaimari, Stephen D. & Yang, Ding, 2013

Shi, Li, Gaimari, Stephen D. & Yang, Ding, 2013, Revision of Sciasmomyia Hendel (Diptera: Lauxaniidae), with eight new species, Zootaxa 3691 (4), pp. 401-435 : 431-434

Sciasmomyia tubata sp. nov.

( Figs 101–111 )

Diagnosis. Face with blackish brown median stripe, a pair of blackish brown oblique lateral stripes confuluent with a blackish brown ventral margin. Mesonotum dark grayish pruinose, with two brown median spots anterirly; anepisternum with a brown oblique I-shaped spot and katepisternum blackish brown (mostly blackish brown in female), with grayish pruinosity. Wing with two hyaline triangular marginal spots between R2+3 and M1; large brown spot on r-m and brown rhombic spot on dm-cu. Abdomen dark brown to black, with grayish pruinosity. Male with syntergosternite 7+8 forming large blackish brown ring, truncate on ventral margin with setulae; epandrium blackish brown, small, broad; surstylus small, semicircular; hypandrium V-shaped, thick, with short hypandrial apodeme; phallus tubular, broad apically, with a pair of slender median subuliform sclerites and a pair of hooked dorsal processes; phallapodeme narrowly separated from and shorter than phallus, consisting of two broad sclerites. Female with hypogynial valve absent.

Description. MALE. Body length 4.5–5.6 mm, wing length 5.3–5.9 mm. FEMALE. Body length 4.3–5.1 mm, wing length 5.3–5.6 mm.

Head ( Fig. 102 ) yellow. Frons with two narrow brown stripes extending to vertex; ocellar triangle black; oc longer than posterior or. Face with a blackish brown median stripe, a pair of blackish brown oblique lateral stripes confluent with a wide blackish brown stripe along ventral margin, and a large brown spot at ventral corner; parafacial yellow except for inner margin black, with a brown elliptical spot at ventral corner. Gena about 5/8 height of eye, with a brown spot. Antenna yellow, 1st flagellomere 1.6 times longer than high; arista with longest setula nearly as long as 1/4 height of 1st flagellomere. Proboscis pale brown. Palpus blackish brown.

Thorax ( Figs 101, 103 ) blackish brown with grayish pruinosity. Mesonotum with two brown median spots anteriorly; anepisternum with a brown oblique I-shaped spot below anepisternal seta; katepisternum blackish brown (darker in female), with grayish pruinosity. Legs yellow except all femora black except at apex, all tibiae with subapical ring, all basitarsi with a brown basal and apical ring, tarsomeres 2–5 entirely brown. Fore femur with 4–5 pv and 8 pd, mid femur with 3 weak a, hind femur with 1 weak preapical ad. Wing ( Fig. 105 ) brown with hyaline spots, all of which display white on that part of the vein; subcostal cell with brown spot; R2+3 with series of irregular longitudinal spots, including preapically; M1 with series of irregular hyaline spots, include large circular spot near middle of wing; triangular hyaline marginal spot between tips of R2+3 and M1; large brown spot on r-m, surrounded by hyaline ring; large brown rhombic spot on dm-cu, with C-shaped hyaline spot proximally and basally, with hyaline spots large on M1 just basal and proximal to dorsal tip of dm-cu; M1 with large preapical hyaline spot; costa with 2nd (between R1 and R2+3), 3rd (between R2+3 and R4+5) and 4th (between R4+5 and M1) sections in proportion of 4.8:1.3:1; ultimate and penultimate sections of M 1 in proportion of 1:1.2; ultimate section of CuA1 about 1/8 of penultimate. Halter pale yellow.

Abdomen ( Fig. 104 ) unicolorous brown to black, with grayish pruinosity. Male genitalia ( Figs 106–109 ): syntergosternite 7+8 blackish brown, forming a large complete ring, truncate on ventral margin with setulae, spiracle very small, with three setae in front of spiracle in lateral view; epandrium blackish brown, small, broad; surstylus small, semicircular with setulae, curled upwards apically in posterior view; hypandrium V-shaped, thick, with hypandrial apodeme short; phallus tubular, broad apically, with a pair of slender median subuliform sclerites and a pair of hooked dorsal processes; phallapodeme narrowly separated from and shorter than phallus, consisting of two broad sclerites, curving backwards and converging, narrowed on dorsoapical margin. Female terminalia ( Figs 110, 111 ): epiproct triangular and hypoproct rectangular; hypogynial valve absent.

Type material. Holotype 3 (CAUC): CHINA, Guizhou Province: Fanjingshan, Mianxuling, 1900 m, 29. VII. 2001, Zhiqi Liu. Paratypes: CHINA, Guizhou Province: data same as holotype (13, CAUC; 1Ƥ, CSCA); CHINA, Yunnan Province: Jinping, Yakou, 18. V. 2006, Junhua Zhang (1Ƥ, CAUC); CHINA, Sichuan Province: E’mei Mountain, Jiulaodong, 1800–1900 m, 11. VIII. 1957, Youcai Lu (1Ƥ, CAUC); E’mei Mountain, Jiulaodong, 1800– 1900 m, 11. VIII. 1957, Keren Huang (1Ƥ (IOZCAS1218968), IZAS); E’mei Mountain, Jiulaodong, 1800–1900 m, 16. VIII. 1957, Keren Huang (1Ƥ (IOZCAS1218973), IZAS); E’mei Mountain, Jiulaodong, 1800–1900 m, 8. V. 1957, Keren Huang (13 (IOZCAS1218982), IZAS); E’mei Mountain, Jiulaodong, 1800–1900 m, 17–19. VIII. 1957, Youcai Lu (13, 2Ƥ (IOZCAS1218976–1218978), IZAS); E’mei Mountain, Jiulaodong, 1800–1900 m, 7. VII. 1957, Zongyuan Wang (13 (IOZCAS1218975), IZAS); E’mei Mountain, Jiulaodong, 1800–1900 m, 25. IX. 1957, Youcai Lu (13 (IOZCAS1218970), IZAS); E’mei Mountain, Xixiangchi, 1800–2000 m, 19. VIII. 1957, Fuxing Zhu (13 (IOZCAS1218964), IZAS); E’mei Mountain, Xixiangchi, 1800–2000 m, 14. VIII. 1957, Keren Huang (13 (IOZCAS1218974), IZAS); E’mei Mountain, Jiulaodong, 1800–1900 m, 3. VIII. 1957, Keren Huang (13 (IOZCAS1218979), IZAS); E’mei Mountain, Jiulaodong, 1800–2000 m, 27. VIII. 1957, Keren Huang (1Ƥ (IOZCAS1218971), IZAS); E’mei Mountain, Jiulaodong, 1800–1900 m, 29. VIII. 1957, Fuxing Zhu (1Ƥ (IOZCAS1218966), IZAS).

Distribution. China (Sichuan, Guizhou, Yunnan).

Remarks. The new species is similar to Sciasmomyia longicurvata sp. nov. in having the face with a pair of brown oblique lateral stripes (not Y- or V-shaped), and the wing having a distinctly apical hyaline apical margin between R2+3 and M1. But, the new species can be differentiated by color patterns on the katepisternum and abdomen, along with characteristics of the male genitalia. Females of S. longicurvata are unknown.

Etymology. Latin, tubata , meaning tubular; refers to the tubular phallus; a feminine adjective.

Figures

FIGURES 101 – 105. Sciasmomyia tubata sp. nov. Holotype male and paratype female. 101. male habitus, lateral view; 102. male head, anterior view; 103. female thorax, dorsal view. 104. male abdomen, lateral view; 105. male wing.

FIGURES 106 – 111. Sciasmomyia tubata sp. nov. Paratype male. 106. syntergosternite 7 + 8 and epandrial complex, lateral view; 107. syntergosternite 7 + 8 and epandrial complex, posterior view; 108. phallus complex, ventral view; 109. phallus complex, lateral view. Paratype female. 110. abdomen tergites and stergites 7 – 9, lateral view; 111. spermathecae. Scale 0.1 mm. Abbreviation: st = syntergosternite 7 + 8.