Pinguisoperla yangzhouensis

Chen, Zhi-Teng, 2018, Pinguisoperla, a new fossil genus of Perlidae (Insecta: Plecoptera) from mid-Cretaceous Burmese amber, Zootaxa 4425 (3), pp. 596-600: 597-598

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Pinguisoperla yangzhouensis


Pinguisoperla yangzhouensis  Chen, sp. nov.

Figs. 1–4View FIGURE 1View FIGURE 2View FIGURE 3.

Diagnosis. As for the genus.

Description. Macropterous; body length (excluding antennae and cerci) ca. 10 mm, generally dark ( Fig. 1 View Figure ). Dorsal head and thorax of the specimen are covered by cracks of the amber; apices of antennae and wings are not preserved; dorsal abdomen is covered by wings; ventral abdominal tip slightly decayed, whose detailed structure is unclear.

Head ( Figs. 1 View Figure , 2 View Figure ): Head inserted in prothorax, dark brown and sub-quadrate except for the dark, bulbous compound eyes; ocelli invisible. Antenna filiform, at least 24-segmented, each segment almost dark except for the paler apices. Maxillary palp four-segmented, two basal segments longest, third segment slightly thinner and shorter, apical segment shortest and thinnest, about half length of the third segment. Labial palp three-segmented, median segment longest, apical segment shortest and thinnest, about 2/3 length of the median segment.

Thorax ( Figs. 1 View Figure , 2 View Figure ): Prothorax much narrower than head, but subequal in length; pronotum dark, trapezoidal with acute corners, anterior margin wider than posterior margin. Mesothorax and metathorax unclear, but strongly sclerotized and wider than prothorax. Legs mostly dark; forelegs shortest, hindlegs longest; femur subequal in length with tibia in forelegs and midlegs, but shorter than tibia in hindlegs; tibia ventrally with two subapical spurs; tarsus three-segmented, two basal segments shortest, subequal in size and with conspicuous euplantulae, apical segment slender and with long apical bristles; claws simple and pointed.

Wings ( Figs. 1 View Figure , 3 View Figure ): Apex of left forewing lost, estimated length ca. 11 mm. ScP not clear; at least five crossveins present between ScP and the anterior margin. RP originated from basal 1/3 of RA and with three branches. M simple, not forked. Cu forked to CuA and CuP, CuA with four branches; CuP simple; area between M and CuA with six crossveins; area between CuA and CuP with six crossveins. AA1 simple, AA2 forked. Hind wings invisible.

Abdomen ( Figs. 1 View Figure , 4): Abdomen dark, near half the length of the body. Sternum 9 darkly sclerotized, posterior margin distinctly extended backwards, median area paler and decayed, unclear, perhaps with specific structures. Paraprocts invisible, not well developed; abdominal tip triangular in shape. Cercus hairy, with about 7 segments; first basal segment strongly sclerotized, enlarged and elongated, curved outward and backward; several subsequent segments still thick but gradually tapering to the hairy apex; each cercal segment subapically fringed with long setae.

Type material. Holotype male, an almost complete specimen in a piece of mid-Cretaceous Burmese amber from Kachin, Hukawng Valley (26°20´N, 96°36´E) of northern Myanmar (98.79 ± 0.62 Ma). The holotype is deposited in the Chen Amber Collection (No. CZT-PLE-MA1), Yangzhou, ChinaGoogleMaps  .

Etymology. The species is named after Yangzhou City, where the author has studied and worked for nearly ten years.

Syninclusions. Plant remains, including leaves and twigs.