Pamphilius benesi Shinohara, 1985

Shinohara, Akihiko, Kramp, Katja & Taeger, Andreas, 2022, The Pamphiliinae of the Russian Far East and Korea (Hymenoptera, Pamphiliidae), Zootaxa 5167 (1), pp. 1-251 : 79

Pamphilius benesi Shinohara, 1985

( Figs 64 , 65 ) (https://doi.org/10.6084/m9.figshare.11405151)

Pamphilius benesi Shinohara, 1985a: 327 ; Shinohara, 1988a: 319; Shinohara & Hara, 1997b: 851; Shinohara, 2001: 108, 113, 116; Shinohara, 2002a: 189; Shinohara, 2002b: 429; Shinohara, 2004: 265; Shinohara & Hara, 2005: 274; Taeger et al., 2010: 86; Shinohara, 2013: 94; Lee et al., 2019: 9; Shinohara, 2019: 9; Shinohara, 2020: 14, 245.

Material examined. About 340 specimens, including the type series. Seventy-two specimens are from South Korea ( Shinohara 2001) .

Distribution. South Korea, Japan (Hokkaido, Honshu, Shikoku, Kyushu) ( Shinohara 2001).

Host plant. Betulaceae : Corylus sieboldiana Blume var. sieboldiana ( Shinohara & Hara 1997b, 2005).

Remarks. This species belongs to the P. sylvaticus group ( Shinohara 1985 a, 1988 a, 2001, 2002b). From Korea, only a series of male specimens collected in 1992 and 1993 have been recorded ( Shinohara 2001) and no material for molecular studies was available. The larvae are gregarious leaf rollers on Corylus .

The maximum intraspecific p -distance within P. benesi from Japan was 2.2% in COI (n=7) and 0.6% in NaK (n=7) and the nearest neighbour was P. pallidus , diverging by a minimum of 2.9% in the COI analysis, and by a minimum of 0.5% in the NaK analysis. In both COI and NaK trees ( Figs 145 , 159 ), P. benesi was the sister of P. pallidus with 100% UFBoot support.

Figures

FIGURE 64. Pamphilius benesi, female, Japan – (a) dorsally, (b) lateroventrally, (c) head & thorax dorsally, (d) head & thorax ventrolaterally, (e) face, (f) sawsheath laterally, (g) antenna.

FIGURE 65. Pamphilius benesi, male, Korea – (a) dorsally, (b) lateroventrally, (c) head & thorax dorsally, (d) head & thorax ventrolaterally, (e) face, (f) antenna, (g) genitalia dorsally, (h) genitalia ventrally, (i) claw.

FIGURE 145. Condensed ML tree based on CO1 gene sequences, with details of Pamphilius sylvaticus group, part 1.

FIGURE 159. Condensed ML tree based on NaK gene sequences, with details of Pamphilius sylvaticus group.