Harpagonella arizonica (I.M. Johnston) Guilliams & B.G. Baldwin

Guilliams, C. Matt, Jang, Timothy & Baldwin, Bruce G., 2016, Molecular and morphological evidence for recognition of two species within Harpagonella (Amsinckiinae, Boraginaceae), PhytoKeys 70, pp. 17-30: 23

publication ID

http://dx.doi.org/10.3897/phytokeys.70.9053

persistent identifier

http://treatment.plazi.org/id/FBA8FC46-7F33-1957-A9D5-676BDE544294

treatment provided by

PhytoKeys by Pensoft

scientific name

Harpagonella arizonica (I.M. Johnston) Guilliams & B.G. Baldwin
status

comb. nov.

Harpagonella arizonica (I.M. Johnston) Guilliams & B.G. Baldwin  comb. nov.

BASIONYM.

Harpagonella palmeri A. Gray var. arizonica  I.M. Johnston. Contr. Gray Herb. 73: 75. 1924. TYPE: U.S.A. Arizona: "plains, Lowell," W.F. Parish 162, May 3, 1884, (holotype: GH! digital image).

SPECIMENS EXAMINED.

Specimens listed alphanumerically by collector within a region. (*=specimen measured; è =specimen also used in molecular study; bold =type specimen) Harpagonella arizonica  : MÉXICO. Baja California. Moran 12682 ( DS). Sonora. Keck 3963 ( DS, POM), Reina & Van Devender 2003-194 è ( ARIZ, ASU), Van Devender 2005-842 è ( ARIZ). UNITED STATES. Arizona. Abrams 12944 ( DS), Baker 8203 ( ASU), Baker 15963 ( ASU), Barr 67-78 ( ASU), Barr 67-82 * ( ARIZ, ASU), Benson 9302 ( POM), Bingham 527 * ( ARIZ), Bingham 1402 ( ASU), Bowers 2250 * ( ARIZ), Bowers 2280 * ( ARIZ), Bowers 2395 * è ( ARIZ), Bowers 2461 * ( ARIZ), Boyle 8026 ( ARIZ), Brandegee, T.S. s.n. 19 April 1889 ( UC), Butterwick 4349 ( ASU), Butterwick 4550 ( ASU), Butterwick & Hillyard 5793 ( ARIZ, ASU), Butterwick 7419 ( ASU), Carter s.n. 17 March 1936 ( ARIZ), Cave 16 ( ARIZ), Damrel 1618-B8 ( ASU), Daniel 2581 (ASU), Daniel & Butterwick 3853 ( CAS), Daniel 3907 ( ASU), Doan 441 ( ASU), Ducote 683 ( ASU), Eastwood 8130 ( CAS), Farruggia 1832 ( ASU), Felger 05-218 ( ASU), Fosberg 10605 ( CAS, RSA), Fosberg 10664 ( CAS, POM), Freeman ( ASU), Gillespie 5429 ( DS), Griffiths s.n. date unknown * ( ARIZ), Halse 1701 ( CAS), Halverson 379 ( ASU), Harrison & Fulton 6608 ( POM), Harrison & Kearney 6654 ( POM), Higgins 6480 ( ASU), Hitchcock 25598 ( DS, RSA), Imdorf & Rice 427 ( ASU, ARIZ), Imdorf 587 ( ASU), Kearney 6654 * ( ARIZ), Keck 2998 ( DS), Keil 1051 ( ASU), Keil 1484 ( ASU), Keil 2864 ( ASU), Keil 4082 ( ASU), Keil 4168 ( ASU), Keil K-11216 ( ASU), Landrum 6656 ( ASU), Landrum 11176 ( ASU), Lane 1035 ( ASU), Lane 1067 ( ASU), Lehto 181 ( ASU), Lehto 307 ( ASU), Lehto 1648 ( ASU), Lehto 1652 ( ASU), Lehto 4594 ( ASU), Lehto 7766 ( ASU), Lehto 10374 ( ASU), Lehto 10389 ( ASU), Lehto 10408 ( ASU), Lehto 10687 ( ASU), Lehto 11733 ( ASU), Lehto 17494 ( ASU), Lehto 17504 ( ASU), Lehto 17541 ( ASU), Lehto 12874-b ( ASU), Lehto L-19732 ( ASU), Lehto L-19740 ( ASU), Makings 2018 ( ASU), Makings, L. Fertig, & W. Fertig 4346 ( ASU, RSA), Manton 236 ( ASU), Mason 1663 * ( ARIZ, CAS), Mauz, Rosen, & Rautenkranz 2005-19 ( ARIZ), McGill LAM1280 ( ASU, RSA), McLaughlin 4476 * è ( ARIZ), Orcutt 173 ( CAS), Parfitt 2498 ( ASU), Parish 162 ( GH; holotype), Parish s.n. 1909 ( DS), Pase 1599 ( ASU), Peebles 1426 * ( ARIZ), Peebles 3693 * ( ARIZ), Pierce 296 ( ASU), Pinkava 4672 ( ASU), Pinkava 10122 ( ASU), Pinkava 10261 ( ASU), Pinkava 10893 ( ASU), Pinkava 11655 ( ASU), Price 829 ( ASU), Rand 15 ( ASU), Rand 152 ( ASU), Reeves 6447-a ( ASU), Reina & Van Devender 97-269 ( ARIZ), Rice 328 ( ASU), Rice 1121 ( ASU), Rice 1586-a ( ASU), Rice 1598 ( ASU), Jones, S. 1433 ( ASU), Schramm, Bond, & Bond 9 ( ASU, RSA), Shreve 7497 ( ARIZ), Shreve 10113 * ( ARIZ, DS), Smith 1577 ( ASU), Swingle s.n. 1914 ( ARIZ), Tedford 582 * ( ARIZ), Tedford 599 * è ( ARIZ), Tedford 614 ( ARIZ), Tedford & Rose 1034 * ( ARIZ), Thornber 2562 * ( ASU, ARIZ, CAS, RSA), Thornber 2581 * ( ARIZ, CAS, RSA), Thornber 4683 ( ARIZ), Thornber 5488 * ( ARIZ), Thornber s.n. 1905 * ( ARIZ), Thornber s.n. 1913 * ( ARIZ), Toumey 5014 * ( ARIZ), Turner 78-41 * ( ARIZ), VanDevender 88-54 * è ( ARIZ), Van Devender 2003-23 * ( ASU, ARIZ), W. Fertig, Makings, & Alcock 29265 ( ASU), Warren 68-25 * ( ARIZ), Warren 68-51 * ( ARIZ), Wiggins 8420 * ( ARIZ), Wiggins 8690 ( DS), Wood ( ASU). Harpagonella palmeri  : MÉXICO. Baja California. Bacigalupi 3067 ( DS, RSA, UC), Boyd 5319 * ( RSA, UC), Boyd & Ross 5464 ( RSA), Boyd & Ross 5761 ( RSA), Boyd, Gross, O’Brien, & Hamilton 10352 ( RSA), Breedlove 62271 ( CAS, RSA), Carter, Chisaki, & Moran 1056 ( UC), Dressler 668 * ( ARIZ), Epling & Stewart s.n. 9 April 1936 ( DS), Haines & Stewart s.n. 7 February 1935 ( DS), Howell 8306 ( CAS), Jones, M.E. s.n. 11 April 1882 ( POM), Moran 6562 ( POM), Moran 6677 ( DS), Moran 6750 ( DS, RSA), Moran 12770 ( UC), Moran 19378 ( CAS), Moran 19992 ( POM), Porter 10551 ( RSA), Rebman & Delgadillo 1638 ( ASU), Rebman & Roberts 4856 ( ASU), Sanders, Rodriguez, West, et al. 5466 ( ASU), Thomas 15730 ( DS), Thorne, Liston, Mistretta 62122 ( RSA), Van Devender 91-348 ( ARIZ), Van Devender, T.R. & R.K. Van Devender 91-239 ( ARIZ), Wiggins & Ernst 12 ( UC), Wiggins & Thomas 67 ( CAS), Wiggins & Ernst 120 ( DS), Wiggins 4265 ( DS, POM), Wiggins 4415 ( POM), Wiggins 4463 ( DS, POM), Wiggins 7600 ( DS, UC). UNITED STATES. California. Atwood 17833 * ( UC), Bacigalupi 8261 * ( JEPS), Banks & Boyd 57 ( RSA), Banks & Boyd 316 ( RSA), Banks & Boyd 398 ( RSA), Banks 1652 ( RSA), Banks 1680 ( RSA), Bell, Clark, Goss, Green, & Rusiniak 3546 ( RSA), Boyd 1384 ( ARIZ, CAS, RSA), Boyd 1396 ( CAS, RSA), Boyd 1399 ( CAS, RSA), Boyd 1589 * ( ARIZ, CAS, RSA), Boyd 1644 * ( ARIZ, CAS, RSA), Boyd 1767 * ( ARIZ, CAS, RSA), Boyd 1790 * ( ARIZ, CAS, RSA), Boyd 1816 * ( CAS, RSA, UC), Boyd 3045 * ( UC), Boyd, Ross, & Arnseth 3029 ( RSA), Boyd, Ross, & Arnseth 3036 ( RSA), Boyd, Ross, & Arnseth 3045 ( RSA), Boyd, Ross, & Arnseth 3116 ( RSA), Boyd, Ross, & Arnseth 3133 ( RSA), Boyd, Ross, & Arnseth 3196 ( RSA), Boyd, Ross, & Arnseth 3206 * ( RSA, UC), Boyd, Ross, & Arnseth 3920 ( RSA), Boyd, Ross, Arnseth, & Bonilla 4008 ( RSA), Boyd, Ross, Arnseth, & Bonilla 4060 ( CAS, RSA), Boyd, Ross, Arnseth, & Bonilla 4110 ( RSA), Boyd, Arnseth, Rasmussen, & Cota 4605 ( RSA), Boyd 6165 ( RSA), Boyd & Mistretta 6311 ( RSA), Boyd 6901 ( RSA), Boyd 6962 ( RSA), Boyd & Ross 7302 ( RSA), Boyd & Ross 7906 è ( RSA, UC), Boyd & Ross 8212 è ( RSA, SBBG, UC), Boyd & Ross 8220 ( RSA), Boyd & Ross 8244 ( RSA), Boyd & Ross 8249 * ( ARIZ, RSA), Boyd & Banks 8279 ( RSA), Boyd 10414 ( RSA, UC), Boyd s.n. 28 March 1982 ( RSA), Boyd s.n. 27 April 1982 ( RSA), Bramlet 2301 * ( ARIZ), Bramlet 2370 ( CAS), Bramlet 2394 ( RSA), Bramlet 2399 ( RSA), Bramlet & Coleman 2418 ( RSA), Bramlet 2982 ( RSA), Bramlet 2988 ( RSA), Bramlet 3352B ( RSA), Brandegee T.S. 824 * ( CAS, POM, UC), Brandegee s.n. 12 April 1894 ( DS), Brandegee s.n. 15 April 1894 * ( RSA, UC), Brandegee T.S. s.n. 8 April 1895 * ( UC), Gander 1128 * ( DS, POM, UC), Gander 3112 * ( JEPS), Gander 5072 * ( JEPS, RSA, UC), Grant 5218 ( DS), Grant & Wheeler 540 ( UC), Gross, Fraga, Virgen, Thibault 1781 ( RSA), Gross, Fraga, Virgen, Thibault 1845 ( RSA), Hamilton s.n. 17 May 2001 ( RSA), Hirshberg 290 ( RSA), Jones, C. 10 ( RSA), Jones, M.E. 3066 ( ARIZ, CAS, DS, POM, UC), Jones, M.E. s.n. 5 April 1882 ( RSA), Junak, Hoefs, & Crockett SCa-351 ( SBBG), Junak, Hoefs, & Crockett SCa-355 ( SBBG), Junak SCa-361 ( SBBG), Junak, Hoefs, & Crockett SCa-379 ( SBBG), Junak, Hoefs, Takara SCa-399 ( SBBG), Junak, Hoefs, & Stratton SCa-497 ( SBBG), Junak, Hoefs, Takara SCa-514 ( SBBG), Junak & Kirkland SCa-573 ( SBBG), Junak & Kirkland SCa-577 ( SBBG), Junak, Hoefs, Kirkland, & Stratton SCa-631 ( SBBG), Junak, Hoefs, & Kirkland SCa-1439 ( SBBG), Junak SCa-1465 ( SBBG), Junak & Philbrick SCa-1529 ( SBBG), Leatherman 65 ( RSA), Marsh & Marsh s.n. 10 June 1991 ( RSA), Moran & Barber s.n. 8 June 2001 ( RSA), Munz & Johnston 5335a * ( CAS, POM, UC), Palmer 70 ( MO; isotype) Parikh 156 ( SBBG), Parikh & Gale 1739 ( SBBG), Parish 12060 ( CAS), Parry s.n. 17 March 1882 ( DS), Peirson 3029 ( RSA), Philbrick & Thorne B67-175 ( SBBG), Pringle 269 ( CAS), Purer 6927 * ( UC), Rebman 8031 * è ( UC), Rebman 8348 * è ( UC), Rebman, Gregory, Mulligan, & Ricks 11673 ( RSA), Rebman, Gregory, Rich, & Principe 12817 * ( RSA, UC), Riefner 20-391 ( RSA), Riefner 20-393 ( RSA), Riefner 95-62 ( RSA), Roberts 3870 ( RSA), Roberts & Bontrager 4565 ( RSA), Roberts, Roberts, & Bontrager 4587 ( RSA), Roberts 4855 ( RSA), Roberts & Bomkamp 4981 ( RSA), Roberts & Bramlet 5563 ( RSA), Roberts & Bramlet 5691 ( RSA), Ross 6853 * ( UC), Ross 6869 ( CAS), Ross & Takara 6939 ( CAS), Ross, Takara, & Otte 6947 ( CAS), Sanders 26178 ( SBBG), Sanders 32379 ( RSA, SBBG), Sanders, Salvato, Volansky, & Balk 32568 ( RSA), Sanders, Wotipka, Elvin, et al. 26153 ( CAS, SBBG), Thorne 35873 ( SBBG), Thorne 35949 * ( UC), True 152 ( POM), Vanderwerff 4235 ( RSA), White 8381 ( ASU, RSA), White & Duchardt 8862 ( RSA).