Moraria hudsoni, Reid, Janet W. & Lesko, Lynn T., 2003

Reid, Janet W. & Lesko, Lynn T., 2003, A new species of Moraria (Crustacea: Copepoda: Harpacticoida) from the Laurentian Great Lakes, Zootaxa 205, pp. 1-19 : 2-8

publication ID	https://doi.org/ 10.5281/zenodo.156303
DOI	https://doi.org/10.5281/zenodo.5657192
persistent identifier	https://treatment.plazi.org/id/FC587B6C-E320-FFB5-B35C-0F1FB692FE47
treatment provided by	Plazi
scientific name	Moraria hudsoni
status	sp. nov.

Treatment

Moraria hudsoni n. sp.

Figures 1 6 View FIGURE 1 View FIGURE 2 View FIGURE 3 View FIGURE 4 View FIGURE 5 View FIGURE 6

Synonymy: Moraria laurentica . Hudson et al. 1998: 34 [partim]. Moraria mrazeki . Hudson et al. 1998: 34 [partim].

Material examined: Holotype female, dissected on slide (USNM 1007895); allotype male, dissected on slide (USNM 1007896); and 7 female, 5 male, and 6 copepodid paratypes, ethanolpreserved (USNM 264250); all from Michigan: Mackinac County: Lake Huron: Prentiss Bay north of Highway M134, 45 °59'26"N, 084°13'37"W, collected with hand grab, 18 August 1994, coll. P.L. Hudson.

Nonparatype specimens: 1 male, Michigan: Lake Michigan: Trails End Bay, 45°44'50"N, 084°48'06"W, among aquatic vegetation, water depth 3060 cm, 6 June 1994, coll. P.L. Hudson (USNM 259905). 1 female and 1 copepodid, Michigan: Lake Michigan: Trails End Bay, location as above, 15 August 1994, coll. P.L. Hudson (USNM 278064). 2 females, Michigan: Mackinac County: Lake Huron: Prentiss Bay: Whitefish Point, Cedar Camp, 45°58'49"N, 084°13'52"W, 7 June 1996, coll. P.L. Hudson (USNM 284307). 7 females, Michigan: Mackinac County: Lake Huron: Prentiss Bay, north of Highway M134, among Carex , water depth 4050 cm, 45°59'26"N, 084°13'37"W, 9 May 1996, coll. P.L. Hudson (USNM 284313). 4 females, Michigan: Mackinac Co.: Lake Huron: Prentiss Bay, location as above, sedge meadow, surface grab, 15 May 2000, coll. P.L. Hudson (USNM 305420).

Female: Length of holotype, from tip of rostrum to tips of caudal rami (specimen extended in lactic acid), 448 m; range of lengths of 7 female paratypes (in glycerin), 392 424 m.

Body ( Figs. 1 View FIGURE 1 , 2 View FIGURE 2 , 3 View FIGURE 3 A) compact, tapering anteroposteriorly. Except for cephalosome, body with numerous rows of minute hairs; integument covered with refractile points, appearing as tiny pores (arrowed circle in Fig. 2 View FIGURE 2 B). Rostrum distinct from cephalosome, bluntly triangular with 2 subapical sensilla. Cephalosome with ovoid dorsal hyaline window; pedigers 2 and 3 each with paired lateral hyaline windows ( Fig. 2 View FIGURE 2 A). Hyaline frills of pedigers 24 smooth; frills of pediger 5, genital doublesomite, and abdominal somites 2 and 3 crenate. Genital doublesomite with row of spines extending dorsally and laterally along posterior margin; abdominal somites 2 and 3 each with row of spines along entire posterior margin. Urosomite with spines along lateral and ventral margins.

Anal operculum ( Figs. 2 View FIGURE 2 B, 3C) rounded, reaching approximately to posterior end of urosomite, its free margin bare.

Caudal ramus ( Figs. 1 View FIGURE 1 B, 2, 3AD) about 2 times longer than wide, tapering posteriorly and with prominent longitudinal dorsal crest extending to 4/5 length of ramus and ending in acute point. Ramus bearing 2 lateral (outer) setae (anterior lateral seta also with tiny hair at its base); 3 terminal setae of which the medial (inner) terminal seta is short and slender, and the middle and lateral terminal setae are stout and spinulose, without interior breaking planes; and 1 dorsal seta. Medial margin of ramus with diagonal row of about 5 spines.

Antennule ( Fig. 3 View FIGURE 3 E) 7segmented, set on prominent base; segments 17 bearing respectively, 1, 8, 6, 2, 1, 2, and 8 setae, segments 4 and 8 also each bearing aesthetasc.

Antenna ( Fig. 3 View FIGURE 3 F, G) with allobasis; exopodite a single segment bearing 1 large stout spinulose seta on margin and 3 slender smooth setae terminally.

Mandible ( Fig. 3 View FIGURE 3 H), endopodite with 2 apical setae.

Maxillule (not illustrated), precoxal arthrite with 7 spines and 2 setae; coxa with apical spine; basis with apical spine and 2 subapical setae.

Maxilla ( Fig. 3 View FIGURE 3 I), syncoxa with 2 endites, each with 2 plumose terminal setae; basis with spatulate spinulose spine and 2 pairs of setae.

Maxilliped ( Fig. 3 View FIGURE 3 J), basis with large plumose distal seta.

Legs 14 ( Fig. 4 View FIGURE 4 AD) with 3segmented exopodites and 2segmented endopodites. Couplers (intercoxal sclerites) without ornament. Leg 1, endopodite slightly shorter than exopodite, segment 1 with slender seta on mediodistal corner, segment 2 with 3 terminal and subterminal setae of which middle seta is about 3 times longer than endopodite. Legs 24, exopodite segment 3 respectively with 4, 4, and 5 socketed spines and setae; endopodite segment 1 with 1, and segment 2 respectively with 2, 3, and 3 setae.

Leg 5 ( Fig. 4 View FIGURE 4 E), baseoendopodites separated, medial lobe of each leg bearing 6 setae. Exopodite distinct from baseoendopodite, ovoid, reaching about to end of baseoendopodite, usually bearing 5 setae, but holotype female with 6th seta on medial margin (arrowed in figure).

Preserved specimens colorless.

Male: Length of allotype, from tip of rostrum to tips of caudal rami (specimen extended in lactic acid), 420 m; range of lengths of 5 male paratypes (in glycerin), 360 400 m.

Body ( Fig. 5 View FIGURE 5 ) like that of female in general shape, number and position of hyaline windows, and ornamentation; in particular, spine rows on all urosomites complete dorsally. Anal somite, operculum, and caudal rami as in female.

Antennule ( Fig. 6 View FIGURE 6 A) apparently of 9 segments, segments 4 and 9 each with aesthetasc.

Antenna and mouthparts as in female.

Legs 14, coxopodites, basipodites, exopodites, endopodite of leg 1, and couplers as in female; spine on mediodistal corner of basipodite of leg 1 also similar to that of female. Legs 24 endopodites 2segmented, all dimorphic. Leg 2 endopodite ( Fig. 6 View FIGURE 6 B, C), segment 1 with 1 spinulose seta on medial margin and 2 stout lateral spines; segment 2 with 2 long slender plumose setae. Leg 3 endopodite ( Fig. 6 View FIGURE 6 D), segment 1 with 1 spinulose seta and 1 spine on medial margin, and 2 spines on lateral margin; segment 2 with stout apophysis having simple acute tip, and 2 terminal setae of which the medial seta is longer and spinulose. Leg 4 endopodite ( Fig. 6 View FIGURE 6 E, F, G) segment 1 with spine on medial surface, larger spine on laterodistal corner, and smooth slender seta on mediodistal corner. Leg 4 segment 2, lateral margin with 1 or 2 spines; apex with doubly recurved, corkscrewshaped spine; and medial margin with 3 or 4 setae, of which 1 seta is at least twice as long as the others, relatively stiff, and spinulose, plus 1 tiny, apparently unsocketed setiform process near base of apical spine.

Leg 5 ( Fig. 6 View FIGURE 6 H), baseoendopodites partly fused; medial lobe bearing 2 stout, spinulose setae; exopodite with 5 setae.

Leg 6 ( Fig. 5 View FIGURE 5 A, C) a broad plate bearing at its outer corner 2 setae and 1 medial spine. Etymology: The species appellation honors the collector, Patrick L. Hudson, in appreciation of his many contributions toward understanding the ecology and taxonomy of the fauna of the Laurentian Great Lakes.