Amphineurus (Rhamphoneurus) anfractus, Santos & Santos & Ribeiro, 2022

Santos, Daubian, Santos, Rodrigo dos Reis & Ribeiro, Guilherme Cunha, 2022, Systematics of Amphineurus (Rhamphoneurus Alexander) (Diptera: Tipuloidea: Limoniidae), Arthropod Systematics & amp; Phylogeny 80, pp. 439-494 : 439

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Amphineurus (Rhamphoneurus) anfractus

sp. nov.

3.3.4. Amphineurus (Rhamphoneurus) anfractus sp. nov.

Material examined.

Holotype: ♂, Chile, Arauco, Nahuelbuta, Butamalal [37°49′S 73°14′W], 1100-1400 m, 23/ 31-I-1954, Peña (USNM) GoogleMaps *. Paratypes: 1 [sex unknown], Chile, Arauco, Nahuelbuta [37°46′S 72°59′W], 5/ 10-II-1953, Peña (USNM) GoogleMaps ; 1 ♂, Chile, Arauco, Nahuelbuta, Butamalal [37°49′S 73°14′W], 1100-1400 m, 23/ 31-I-1954, Peña (USNM) GoogleMaps *.


The word Amphineurus anfractus in Latin is for “winding”. The term refers to the winding or sinuous aspect of the anal vein and crossvein m-cu.


This species is distinguished by sinuose crossvein m-cu and anal vein. Furthermore, this species is characterized by the male terminalia with a folded medial branch of the clasper of gonostylus and a cylindrical left mesal lobe.


Wing length 7.84 mm, width 2.48 mm. - Coloration: General coloration dark brown. Thorax dark brown, pleura brown. Scutum with one black stripe. Halter pale with knobs ochreous. Coxae brownish yellow and legs dark brown. - Head (dorsal view Fig. 11B View Figure 11 , ventral view Fig. 11A View Figure 11 ): Rostrum bifurcated with long appendices; first palpal segment shorter than terminal segment; scape as thick as pedicel, pedicel constricted at base; terminal flagellomere shorter than penultimate, antenna with long setae. - Thorax (lateral view Fig. 11E View Figure 11 , dorsal view Fig. 11F View Figure 11 ). Division line of laterotergite very sinuous; anatergite shorter than katatergite. Halter with dilated knob. Wing (Fig. 11D View Figure 11 ) with several marks, including faint markings on A1 and near tip of R3, marking highlighting m-cu, markings on Rs, R2+3+4 and R2+3, small, strong marking at tip of R1 and large, strong marking covering R2; veins near fork of bM (Fig. 11C View Figure 11 ) faded; A1 sinuose; M1+2 longer than basal deflection of M1; R2+3+4 longer than R2+3. - Male terminalia (Fig. 11G View Figure 11 ): Male tergite IX darkened along posterior margin; posterior margin flat, with short median U-shaped notch; posterolateral lobes nearly rectangular with slightly visible shallow emarginations. Ventral branch of gonocoxite wide. Dorsal branch of gonocoxite long, almost as long as ventral branch, with long setae on distal tip. Lobe of gonostylus with rounded lobule; lobule wider than long, shorter than half-length of lobe of gonostylus, twice as wide as stem. Clasper of gonostylus with differently shaped branches: lateral branch straight; medial branch curved, club-shaped. Mesal lobes of gonocoxite asymmetrical, both similarly shaped with flat posterior margins; left mesal lobe narrower. Sheath of aedeagus slightly curved on distal half, with tip narrowed, darkened almost along entire length.


Some specimens of this new species were previously identified by C.P. Alexander as A. (R.) glabristylatus Alexander and some as A. (R.) nothofagetorum Alexander. This species resembles A. (R.) chiloeanus but differs mainly in the form of the anal vein, shape of male tergite IX, sheath of aedeagus and mesal lobe.