Exocelina ketembang (Balke, 1998)

Shaverdo, Helena, Sumoked, Bob & Balke, Michael, 2017, Descriptions of two new species and one new subspecies from the Exocelinaokbapensis-group, and notes on the E. aipo-group (Coleoptera, Dytiscidae, Copelatinae), ZooKeys 715, pp. 17-37 : 27-31

3. Exocelina ketembang (Balke, 1998) Figs 8, 10

Copelatus (Papuadytes) ketembang Balke, 1998: 332; Nilsson 2001: 77 (catalogue).

Papuadytes ketembang (Balke, 1998): Nilsson and Fery 2006: 56 (comb.n.).

Exocelina ketembang (Balke, 1998): Nilsson 2007: 34 (comb.n.).

Exocelina ketembang (Balke, 1998): MB0680: Toussaint et al. 2014: Supplementary figs 1-4, tab. 2.

Type locality.

Papua: Pegunungan Bintang Regency, Borme, according to the original description.

Type material studied.

Holotype: The holotype and 46 paratypes from the same locality have not been found. According to the original description ( Balke 1998), they were deposited at the NHMW and had labels with the following data: Irian Jaya, Jayawijaya Province, Borme, 900 m, 18.viii.1992, IR 92#17, M. Balke leg. No beetles with such label data have been found in the collections of NHMW, ZSM, CGW, or in London. Moreover, the label data of this locality (17 from 1992) are wrongly given in the original descriptions not only for this species but also for Copelatus takime Balke, 1998: locality 17 is not Borme but Kali Takime, as the labels of the type specimens of C. takime state. There are four males (three in NHMW and one in CGW) with the geographical labels "IRIAN JAYA Zentralmassive 140°25'E, 04°24'S " and "14./17.viii.1992 Borme, 1900 m leg. Balke (11)", one of which has a red holotype label. Therefore, perhaps, the holotype label was simply misquoted in the original description. However, since the type series, indicated in the original description, is very large (47 specimens), investigations on this subject are ongoing.

Paratypes: 2 males "IRIAN JAYA: Borme, Tarmlu, 1500 m 6.9.1993", "ca. 140°25'E, 04°24'S leg. M. Balke (4)", "Paratypus Copelatus ketembang Balke des. 1997" [red], one of them with an additional green label "M. Balke 3284" (NHMW). 3 males "IRIAN JAYA: Borme, Tarmlu, 1500 m, 6.ix.1993", "ca. 140°25'E, 04°24'S leg. M. Balke (4-6)", "Paratypus Copelatus ketembang Balke des. 1997" [red], one of them with two additional labels "M. Balke 3285" [green] and "M. Balke 6410" [green text] (NHMW). 4 males "IRIAN JAYA Zentralmassive 140°25'E, 04°24'S ", "14./17.8.1992 Borme, 1900 m leg. Balke (11)", "Paratypus Copelatus ketembang Balke des. 1997" [red] (CGW, NHMW). 1 male "IRIAN JAYA: 1.x.1993 Eme Gebiet Okloma, 1500 m", "ca. 139°55'E, 04°14'S leg. M. Balke (28)", "Paratypus Copelatus ketembang Balke des. 1997" [red] (NHMW). 5 males, 4 females "IRIAN JAYA, 7.9.1992 Kono, 1800 m 139°47'E, 04°21'S, leg. Balke (41)" (CGW, NHMW); the females might belong to two species, Exocelina ketembang and E. damantiensis (Balke, 1998).

Additional material.

Indonesia: Papua Province: Pegunungan Bintang Regency: 1 female "IRIAN JAYA Zentralmassive 140°25'E, 04°24'S ", "14./17.8.1992 Borme, 1900 m leg. Balke (11)" (NHMW). 1 male, 1 female "IRIAN JAYA Zentralmassive 140°25'E, 04°24'S ", "Borme, 1800 m 16.8.1992 leg. Balke (12, 12 A)" (NHMW). Papua New Guinea: Sandaun Province: 1 male "M. Balke 670", "Papua New Guinea: Sandaun, Mekil (WB106), 14.x.2003, K.Sagata, M. Balke: MB 670" (ZSM). 1 male "M. Balke 680", "Papua New Guinea: Sandaun, Mekil (WB106), 14.x.2003, K. Sagata, M. Balke: MB 680" [04° 48.742S 141.39.075E] (ZSM). 1 male "M. Balke 3727" [green], "Papua New Guinea: Sandaun, Ofektaman, 820 m, 17.x.2008, 5.04.113S 141.35.841E, Ibalim (PNG 190)" (ZSM).

Females of doubtful identity.

Indonesia: Papua Province: Pegunungan Bintang Regency: 1 female "IR 92#17B: West New Guinea, Tanime, 1600 m, 20.viii.1992, Balke" (NHMW); this female might belong to three species: Exocelina ketembang , E. danae (Balke, 1998) and E. damantiensis . 1 female "IRIAN JAYA, 7.ix.1992 Kono, 1800m 139°47'E, 04°21'S, leg. Balke (41) (CGW), the female might belong to two species: Exocelina ketembang and E. damantiensis . Papua New Guinea: Sandaun Province: 3 females "Papua New Guinea: Sandaun, MekilK [sic!], 1718 m, 14.x.2003, 4 48.742S 141 39.075E, K. Sagata (WB 106)" (ZSM); these females might belong to two species: E. sandaunensis Shaverdo & Balke, 2014 and E. ketembang . 7 females "Papua New Guinea: Sandaun, Ofektaman, 820 m, 17.x.2008, 5.04.113S 141.35.841E, Ibalim (PNG 190)" (ZSM); these females might belong to three species: E. sandaunensis , E. aipomek (Balke, 1998), and E. ketembang .

Diagnosis.

Beetle medium-sized (TL-H 3.9-4.35 mm); oblong-oval; dark brown to piceous, sometimes with reddish brown pronotal sites and head anteriorly; shiny, with very fine punctation and weakly impressed microreticulation; pronotum with distinct lateral bead; male antennae simple (Fig. 8); male protarsomere 4 with large, thick, strongly curved anterolateral hook-like seta; male protarsomere 5 long and narrow, without concavity, with anterior row of 21 and posterior row of six relatively short setae; median lobe strongly curved, sharply pointed in lateral view, evenly tapering, with narrowly rounded apex in ventral view; paramere with notch on its dorsal side and small, less evidently separated subdistal part with a tuft of dense setae; proximal setae inconspicuous (Fig. 10 A–D). For complete description, see Balke (1998).

Distribution.

Papua: Pegunungan Bintang Regency and PNG: Sandaun Province (Fig. 12).