Oxylymma rileyi, Heffern & Botero & Santos-Silva, 2020

Heffern, Daniel, Botero, Juan Pablo & Santos-Silva, Antonio, 2020, Synonymies, new records, and new species in Cerambycinae (Coleoptera Cerambycidae), Zootaxa 4747 (1), pp. 177-190 : 187-189

Oxylymma rileyi sp. nov.

( Figs. 31–35 )

Description. Male. Head mostly orangish-yellow; antennal tubercles with transverse brown macula; with large dark brown macula (almost black on some areas, more light brown toward prothorax) on each side of vertex, covering area close to upper eye lobes, mostly of area between posterior margin of eyes and prothorax, extended toward area behind eyes (gradually narrowed toward middle of lower eye lobes); remaining surface of vertex yellowish-brown (this area noticeably narrowed centrally after posterior margin of eyes, then gradually widened toward prothorax); apex of anteclypeus and base of labrum brownish; mandibles black from about middle to apex; last maxillary and labial palpomeres mostly brown; scape with two longitudinal dark brown bands (almost black), one on outer side, another ventrally, and remaining surface yellowish; outer side of pedicel and antennomeres III–IV dark brown (almost black), and remaining surface yellowish; antennomere V entirely yellowish; base of antennomere VI yellowish, and remaining surface dark brown (almost black); antennomeres VII–XI dark brown; prothorax orange, more orangish-yellow on part of ventral surface, and posterior area of dorsal and lateral surfaces; ventral surface of mesothorax orangish-yellow centrally, gradually more orange toward sides, except dark-brown on mesanepisternum; metanepisternum dark brown, except yellowish distal area; metaventrite with large triangular macula laterally, dark brown close to metanepisternum, gradually yellowish-brown toward central area, and remaining surface yellowish, irregularly stained by yellowish-brown and brownish areas; scutellum orange; elytra yellowish-white, except orangish or brownish punctures; pro- and mesofemora yellowish, except nearly black longitudinal band dorsally, only on femoral club in profemora, and from middle of peduncle in mesofemora; metafemora yellowish, except small dark brown macula on apex of dorsal surface; tibiae black dorsally, orangish-yellow on remaining surface; tarsomeres orangish-brown, except dark brown posterior area; abdominal ventrites yellowish, gradually lighter toward apex of ventrite V.

Head. Anterior area of head forming distinct rostrum. Frons finely, sparsely punctate (punctures slightly coars- er laterally); glabrous, with three long, erect, dark setae on each side near eyes. Area between antennal tubercles and posterior margin of upper eye lobes flattened, obliquely sulcate close to eyes and centrally, somewhat rugose on remaining surface; with moderately short, erect, sparse yellowish-white setae; area close to upper eye lobes moderately finely, shallowly punctate, with a few short, erect yellowish-white setae; area of vertex close to prothorax moderately finely, punctate (somewhat rugose), glabrous.Area behind eyes tumid close to eye, smooth behind upper eye lobes, striate behind and close to lower eye lobe; area close to prothorax moderately finely, abundantly punctate (punctures transverse); with a few long, erect yellowish-white setae close to upper eye lobe, and short, erect setae of same color on middle of tumid area behind lower eye lobe. Antennal tubercles nearly contiguous, smooth and glabrous. Genae 0.7 times length of lower eye lobe, finely, sparsely punctate; with sparse, moderately long, erect yellowish-white setae, and one long, erect dark seta inferiorly. Median groove distinct from near clypeus to area between upper eye lobes (carina-shaped between antennal tubercles and upper eye lobes). Postclypeus nearly smooth centrally, with a few fine punctures laterally; with moderately short, erect yellowish-white setae on punctate area, and a very long, erect seta of same color on each side. Labrum with long, erect, moderately abundant yellowishwhite setae on anterior half, longer laterally. Gulamentum smooth, glabrous on posterior half, transversely striate, with moderately long, erect, sparse yellowish-white setae on anterior half; area between lower eye lobes and gulamentum with long, erect, sparse dark setae. Eyes protuberant; distance between upper eye lobes 0.77 times length of scape (0.46 times distance between outer margins of eyes); in frontal view, distance between lower eye lobes 0.69 times length of scape (0.41 times distance between outer margins of eyes). Antennae 2.0 times elytral length, reaching elytral apex near middle of antennomere IX. Scape and pedicel finely, sparsely punctate, with long, erect, sparse brownish setae throughout. Antennomeres III–XI elongated, cylindrical; antennomeres III–IV with long, erect dark setae ventrally, with long, erect yellowish setae interspersed, and moderately short, erect, sparse yellowish setae dorsally (III with a few longer dark setae interspersed); remaining antennomeres with moderately long, erect yellowish setae throughout. Antennal formula (ratio) based on length of antennomere III: scape = 0.65; pedicel = 0.16; IV = 0.53; V = 0.64; VI = 0.61; VII = 0.64; VIII = 0.58; IX = 0.57; X = 0.51; XI = 0.54.

Thorax. Prothorax slightly wider than long, with distinct anterior and posterior constriction; posterior margin widely, slightly concave; anterior margin straight; sides rounded between constrictions. Pronotum distinctly depressed in posterior third, with noticeable, strongly elevated, laterally flattened tubercle placed centrally after middle; coarsely, moderately abundantly punctate on somewhat V-shaped anterior area (centrally reaching central tubercle), smooth on remaining surface; with long, erect, moderately sparse yellowish-white setae. Sides of prothorax coarsely, abundantly punctate on anterior 2/3, sparsely on posterior third; with long, erect, sparse yellowish-white setae. Prosternum with large, nearly triangular, slightly depressed, coarsely, partially confluently punctate area on each side of posterior 2/3; central area transversely striate from anterior constriction to prosternal process; with a few short, erect yellowish-white setae. Central area of prosternal process distinctly narrowed. Ventral surface of meso- and metathorax with whitish pubescence laterally, not obscuring integument, nearly absent centrally; metaventrite with long, erect, sparse yellowish setae throughout. Scutellum rectangular, longitudinally concave centrally, smooth, glabrous. Elytra. Parallel-sided in anterior 3/4, gradually narrowed toward apex in posterior quarter; coarsely, abundantly punctate on anterior third, gradually finely punctate on remaining surface, especially on posterior half; apex triangular-shaped; with long, erect yellowish-white setae throughout, and a few long, erect dark setae anteriorly and posteriorly. Legs. Femora distinctly pedunculate-clavate; with long, erect, sparse yellow- ish setae. Protibiae with long, erect yellowish setae dorsally, and sides of posterior half; remaining surface with dense, bristly yellowish-white setae. Meso- and metatibiae with long, erect, moderately sparse yellowish setae, and short, moderately abundant, bristly yellowish setae on posterior quarter of ventral surface.

Abdomen. Ventrites finely, sparely punctate; with long, erect, sparse yellowish setae, denser on posterior third of ventrite V; apex of ventrite V truncate.

Dimensions (mm). Total length, 8.75; prothoracic length, 1.70; anterior prothoracic width, 1.30; posterior prothoracic width, 1.70; maximum prothoracic width, 1.85; humeral width, 2.40; elytral length, 5.25.

Type material. Holotype male from PANAMA, Bocas del Toro: Fortuna Cabins , forest trail (8.7814ºN / 82.1909ºW), 16-20.V.2018, E.G. Riley col. ( TAMU).

Remarks. Oxylymma rileyi sp. nov. belongs to the group of species with longer and filiform antennae. It differs from all other species of the genus by the antennae distinctly longer than the body, and by the noticeably elevated tubercle of the pronotum ( Fig. 34 ). In the other species of the genus the antennae are distinctly shorter than body, and there is no strongly elevated tubercle in the pronotum.

Etymology. This species is named for our friend and colleague, Edward G. Riley, of College Station, Texas, who began collecting Coleoptera in Panama at a young age before eventually working for over a quarter of a century as the curator for the Texas A&M Department of Entomology’s insect collection. His recent collecting trips to Panama have yielded a treasure trove of specimens that will be appreciated by many future generations of researchers.

Figures

FIGURES 31–35. Oxylymma rileyi, holotype male: 31) Dorsal habitus; 32) Ventral habitus; 33) Lateral habitus; 34), Oblique habitus; 35) Head, frontal view.