Chen & Toda, 1997: 787 Chen & Toda, 1997: 789 Revision of the subgenus Phortica (sensu stricto) (Diptera, Drosophilidae) from East Asia, with assessment of species delimitation using DNA barcodes Huang, Jia Gong, Lu Tsaur, Shun-Chern Zhu, Lin An, Keying Chen, Hongwei Zootaxa 2019 2019-09-30 4678 1 1 75 4GS4L (Chen & Toda, 1997) Chen & Toda 1997 [151,740,1485,1512] Insecta Drosophilidae Phortica Animalia Diptera 32 33 Arthropoda species foliata Phortica     Amiota( Phortica) foliata  Chen & Toda, 1997: 787.    Amiota( Phortica) antillaria  Chen & Toda, 1997: 789. syn. nov.   Diagnosis.Surstylus somewhat oblong; vertical process of gonopod dilated laterally, with 1 or 2 small projection(s) mediodorsally or apically; aedeagal outer membrane mostly with numerous spinules (after Chen & Toda 1997).   Remarks. Chen & Toda (1997)described two variable characters as the diagnosis of  P. antillaria, but we have not yet found sufficient morphological or molecular evidence in supporting  P. antillariaas a different species compared with  P. foliata.  Specimens examined.  CHINA: 1♂( SCAU), Lijia, Taitung, Taiwan, alt.  1300m, 28.iv.1994, S.W. Chang;  3♂( KIZ), Jiaoye Park, Kunming, Yunnan, 25°04'13''N, 102°37'13''E, alt.  2100m, 22.vii.2004, J.J. Gao;  2♂( KIZ), Yixiang, Pu'er, Yunnan, 22°44'19''N, 101°07'01''E, alt.  1400m, 6.xii.2000, J.J. Gao;  24♂( SCAU), Nanling, Ruyuan, Guangdong, 24°05'22''N, 113°04'47''E, alt.  550–1500m, 5♂on 23.vii.2002, H. Takamori, 6♂on 2–5.v.2004, H.W. Chen, 11♂on 8–10.ix.2017, L. Gong, N.N. Wang, 2♂on 28.v.2018, H.W. Chen, L. Gong;  2♂( SCAU), Menglun South 5km, Mengla, Yunnan, 21°59'57''N, 101°02'50''E, alt.  820–890m, 8.iii.2003, J.J. Gao;  1♂( KIZ), Xujiaba, Jingdong, Yunnan, 24°31'04''N, 101°00'46''E, alt.  1700m, 14.xi.2003, B.C. Wang;  17♂, Diaoluoshan, Lingshui, Hainan, 18°43'51''N, 109°51'28''E, alt.  980m,  2♂( SCAU) on  3.xii.2003, H.W. Chen,  5♂( SCAU) on  22–24.v.2004, H.W. Chen, 6♂on 17.v.2005, H.L. Cao, H.W. Chen, M.F. Xu,  4♂( KIZ) on  23.iv.2008, X.P. Chen, J.J. Gao;  2♂( SCAU), Wuyishan, Yanshan, Jiangxi, 27°59'38'', 117°49'04''E, alt.  1200m, 15.vi.2004, H.W. Chen, J.J. Gao;  2♂( SCAU), Mangshan, Yizhang, Hunan, 24°56'53''N, 112°54'01''E, alt.  1200–1300m, 25–27.iv.2005, H.L. Cao, M.F. Xu;  2♂( SCAU), Pingzhena, Chiayi, Taiwan, alt.  500m, 20.v.2005, C.F. Lee;  13♂( SCAU), Renai, Nantou, Tai- wan, alt.  2100m, 20.ix.2005, S.W. Chang;  6♂( SCAU), Jizushan, Binchuan, Yunnan, 25°56'54''N, 100°23'58''E, alt.  2100m, 21.viii.2006, H.L. Cao, H.Z. Cao, T. Li;  3♂( SCAU), Grand Canyon, Maoershan, Xing'an, Guangxi, 25°51'28''N, 110°25'01''E, alt.  380–580m, 22.ix.2006, J.J. Jiang, F. Zhao;  1♂( SCAU), E'meishan, Leshan, Sich- uan, 29°36'14''N, 103°28'56''E, alt.  700m, 7.vii.2007, H.Z. Cao;  15♂( 8♂in KIZ, 7♂in SCAU), Jianfengling, Le- dong, Hainan, 18°41'50''N, 108°50'04''E, alt.  700–860m, 17.iv.2008, X.P. Chen, J.J. Gao;  3♂( KIZ), Damingshan, Shanglin, Guangxi, 23°41'01''N, 108°30'33''E, alt.  900m, 15.iii.2009, J.J. Gao;  9♂( SCAU), Dongyanshan, Taoyu- an, Taiwan,  5.i.2010, no other data;  1♂( TMNS), Tongmai, Bomi, Xizang, 30°06'00''N, 95°05'00''E, alt.  2000m, 11.x.2010, L. Wang;  68♂, Hesong, Menghai, Yunnan, 21°50'17''N, 100°05'48''E, alt.  1700–1900m,  43♂( SCAU) on  6.iv.2011, J.M. Lu, Z.F. Shao, Y.R. Su,  25♂( 15♂in KIZ, 10♂in SCAU) on 7,12.  v.2012, H.W. Chen, J.J. Gao;  25♂, Baihualing, Baoshan, Yunnan, 25°17'50''N, 98°48'12''E, alt.  1400m,  10♂( KIZ) on  12.vi.2011, H.W. Chen, J.J. Gao,  15♂( SCAU) on  24.viii.2013, Q.S. Gao, K. Liu;  5♂( SCAU), Langping, Tianlin, Guangxi, 24°27'43''N, 106°22'52''E, alt.  1490m, 30.vii.2016, H.W. Chen, L. Gong, Y.Q. Liu;  6♂( SCAU), Banpo, Lüchun, Yunnan, 22°34'50''N, 102°34'50''E, alt.  560–860m, 27.x.2016, H.W. Chen, J.J. Gao;  1♂( SCAU), Dawuling, Xinyi, Guang- dong, 22°16'29''N, 111°11'39''E, alt.  570m, 22.vi.2017, H.W. Chen;  2♂( SCAU), Zhongshan, Mangshi, Yunnan, 24°07'49''N, 98°31'27''E, alt.  1480m, 3.xi.2017, H.W. Chen, B.X. Li;  2♂( SCAU), Moli ForestPark, Ruili, Yunnan, 24°07'05''N, 97°58'51''E, alt.  920m, 8.xi.2017, B.X. Li;  2♂( SCAU), Muyiji Park, Ximeng, Yunnan, 22°37'15''N, 99°35'42''E, alt.  1200m, 16.iv.2018, L. Gong.   Distribution. China( Jiangxi  *, Hunan  *, Taiwan, Guangdong, Hainan, Guangxi, Yunnan  *, Guizhou, Xizang  *). 2423595489 [463,1106,1834,1860] SCAU Taitung & Taiwan China 1300 Lijia 32 33 1 1 2423595239 KIZ China 2100 25.070278 Kunming 21 102.62028 Jiaoye Park 32 33 3 3 Yunnan 2423595006 KIZ China 1400 22.738611 Pu'er 21 101.11694 Yixiang 32 33 2 2 Yunnan 2423595336 SCAU China 1025 24.089445 Ruyuan 21 113.07972 Nanling 32 33 24 24 Guangdong 2423595285 SCAU China 855 21.999166 Mengla 21 101.04722 Menglun South 32 33 2 2 Yunnan 2423595136 KIZ China 1700 33 34 24.517778 Jingdong 21 101.01278 Xujiaba 32 33 1 1 Yunnan 2423594889 China 980 18.730833 Lingshui 21 109.85777 Diaoluoshan 33 34 17 17 Hainan 2423595062 [662,1091,189,215] 2003-12-03 SCAU Chen China 33 34 2 2 2423595548 2004-05-22 2004-05-24 2004-05-22 SCAU Chen China 33 34 5 5 2423595002 [874,1388,225,251] 2008-04-23 KIZ X. P. Chen & Gao China 33 34 4 4 2423594950 SCAU China 1200 Yanshan 117.81778 Wuyishan 33 34 2 2 Jiangxi 2423595247 SCAU China 1250 24.948055 Yizhang 21 112.900276 Mangshan 33 34 2 2 Hunan 2423595311 [203,789,332,359] SCAU Taiwan China 500 Chiayi Pingzhena 33 34 2 2 2423595551 SCAU China 2100 Nantou Renai 33 34 13 13 2423594904 SCAU China 2100 25.948332 Binchuan 20 100.399445 Jizushan 33 34 6 6 Yunnan 2423595334 SCAU China Grand Canyon 480 25.857779 Xing'an 20 110.41694 Maoershan 33 34 3 3 Guangxi 2423595510 SCAU China 700 29.60389 Leshan 20 103.48222 33 34 1 1 2423594981 KIZ, SCAU China 780 18.697222 Jianfengling 21 108.83445 33 34 30 30 Hainan 2423595529 KIZ China 900 23.68361 Shanglin 21 108.50916 Damingshan 33 34 3 3 Guangxi 2423595074 2010-01-05 SCAU China Taiwan Dongyanshan 33 34 9 9 2423594962 [568,1432,584,611] TMNS China 2000 30.1 Bomi 20 95.083336 Tongmai 33 34 1 1 Xizang 2423595061 [392,1272,620,647] China 1800 21.838057 Menghai 21 100.09667 Hesong 33 34 68 68 Yunnan 2423595231 2011-04-06 SCAU J. M. Lu & Z. F. Shao & Su China 33 34 43 43 2423595014 [629,1432,656,683] 2012-05 KIZ, SCAU H. W. Chen & Gao China 33 34 50 50 2423595267 [151,984,692,719] China 1400 25.297222 Baoshan 21 98.80334 Baihualing 33 34 25 25 Yunnan 2423595491 2011-06-12 KIZ H. W. Chen & Gao China 33 34 10 10 2423595277 [258,803,728,755] 2013-08-24 SCAU Q. S. Gao & Liu China 33 34 15 15 2423594900 SCAU China 1490 24.461945 Tianlin 21 106.38111 Langping 33 34 5 5 Guangxi 2423595283 SCAU China 710 22.580557 Luchun 21 102.58055 Banpo 33 34 6 6 Yunnan 2423594912 SCAU China 570 22.274721 Xinyi 21 111.19417 Dawuling 33 34 1 1 2423595086 SCAU China 1480 24.130278 Mangshi 21 98.52417 Zhongshan 33 34 2 2 Yunnan 2423594928 SCAU China 920 24.118055 Ruili 21 97.980835 Moli Forest 33 34 2 2 Yunnan 2423594905 SCAU China 1200 22.620834 Ximeng 21 99.595 Muyiji Park 33 34 2 2 Yunnan