Revision ofEutyphlusLeConte (Coleoptera: Staphylinidae: Pselaphinae), with Description of a New Species and Phylogenetic Placement within the Tribe Trichonychini
Owens, Brittany E.
Carlton, Christopher E.
The Coleopterists Bulletin
2016
2016-03-31
70
1
1
29
3DF7C
O. Park
1956
[310,521,778,799]
Insecta
Staphylinidae
Eutyphlus
Animalia
Coleoptera
27
Arthropoda
species
thoracicus
Material Examined.( n= 150) *TENNESSEE: Sevier Co. GSMNP, Trillium Gap Tr. @ 35°39.9′N 83°26.2′W 1400m. Forest litter. 29 July 2004. A.Tishechkin/ LSAM0080787 ( LSAM)(slide mounted male)(description of male based on this specimen). *TENNESSEE:Sevier Co.; GSMNP, Trillium Gap Tr. @; 35°39.9′N 83°26.2′W; 1400m. Forest litter. 29 July; 2004. A.Tishechkin ( LSAM)(slide mounted female)(description of female based on this specimen). *Rocky Creek Gap Smoky Mts. Nat. Pk. Tenn. 1X:14:53, SA-HD/ Field Mus. Nat. Hist. Orlando Park Pselaphidae Colln./ (orange label) HOLOTYPE/ Eutyphlus thoracicus. Park, nsp. ( FMNH)(1M; HOLOTYPE). *same locality/ (orange label) PARATYPE. ( FMNH) (2F). *Chimney Cp. Gr Smoky Mts. N. P. Tenn.16: IX:53 Auerbach #5/ Field Mus. Nat. Hist Orlando Park Pselaphidae Colln./ (orange label) PARATYPE( FMNH)(1F). *Chimney Cp. Gr Smoky Mts. N. P. Tenn.16:IX:53 Auerbach #5/ Field Mus. Nat. Hist Orlando Park Pselaphidae Colln./ (orange label) PARATYPE( FMNH)(1F). *Cloudland Canyon S. Pk. Dade Co., GA. 14. VI.62 Forest floor debris/ H. R. Steeves, Jr. J.D.Patrick Jr. Collectors/ H. R. Steeves Jr. Collection ( FMNH)(5F). *N CAROLINA: Haywood Co. GSMNP, Caldwell Fork Tr. At UTM 30897 E 3940883 N. Moist forest Berlese. 3 August 2002. C.Carlton & N. Lowe ( LSAM)(18F; 2M) * USA: NCHaywood Co. GSMNP McKee Branch Tr. 35°35′6″N 83°05′W/ 1370m 16Jul–12Aug2002FIT C.Carlton & S.Gil ( LSAM)(1M). *N CAROLINA: Haywood Co. GSMNP, Mt. Sterling Gap. 35°42.0′N 83°5.9′W 1190m. Forest litter. 29 July 2004. R. T.Allen/ LSAM0095711 ( LSAM)(1F). *same locality ( LSAM)(16F; 1M). * USA: NC: Haywood Co. GSMNP Purchase Knob, berlese/ 313124E 3939640N 5089ft. 20 Jul 2002CCarlton (3F; 1M). * USA: NC: Macon Co. 3 miNW Highlands v-29- 1983, sift rhododendron litter, DSChandler ( DSC) (7F). * USA: NC: Swain Co GSMNP Appalachian Tr. 1.5 miNE Newfound Gap/ 281639E 3944038N ridgetop berlese 18 Jul 2002CCarlton ( LSAM)(3F). *N CAROLINA: Swain Co. Appalachian Tr. ∼ 1.5miE Newfound Gap. 35°36.96′N 83°24.66′W 1730m. Forest litter; 18 July 2002. C.Carlton ( LSAM)(1F). * U.S.A., NC, Swain Co.; GSMNP, Beech Gap Tr.; 83°12′32″W, 35°38′23″N/ elv. 1400m, 20, Oct. 2001; N. hardwood berlese; LSAM, C.Carlton ( LSAM)(2F). *N CAROLINA: Swain Co. GSMNP, jct. of Beech Gap & Hyatt Ridge Trails 35°37.82′N 83°42.07′W 1495m. Forest litter. 2 Oct 2005. W.D. Merritt ( LSAM)(3F). * USAN.C. Swain Co. Dirt Ri. From Heintooga Overlook to Cherokee/ Lot #76-102 Oct. 14, 1976Berlesate 5000′- 4500′ R.Chenowith & R.Tallen ( LSAM)(7F). * U.S.A., NC, Swain Co. GSMNP, Flat Creek Trail 83°10′21″W, 35°33′1″N/ elv. 1500m, leaf litter Berlese, 31 July 2001. A. Tishechkin ( LSAM) (1M). *N CAROLINA: Swain Co. GSMNP, Jenkins Ridge Tr. ∼ 1mifrom Appalachian Tr. 35°33.75′N 83°43.2′W 1540m. Forest litter. 30 Jul 2004. A.K. Tishechkin ( LSAM)(3F). *N CAROLINA: Swain Co. GSMNP, upper Jenkins Ridge Tr. At 35°33.75′N 83°43.2′W. 1540m. Leaf litter 31 Jul 2004. A.K. Tishechkin ( LSAM)(1F). *N. CAR. Swain Co. Mt. Collins 5900′ v.21.1977 A.Newton, M.Thayer ( DENH)(2F). *N CAROLINA: Swain Co. GSMNP, Noland Divide Tr. at 35°33′57″N 83°28′36″W 1770m.Leaf/moss mat litter 19 July 2003. A. Tishechkin ( LSAM)(2F). * USA: NC. Yancey Co. Black Mtn. info cntr v-21-1991sift fir-oak S.O’ Keefe collr ( DSC) (3M). * USA: NC, Yancy Co. B. R. Pkwy. 37.4 mi.SW intersection US221 elv. 4600′ log litter 2.vi.1991C.E.Carlton ( LSAM)(1F). * SC: Oconee Co. 6.7 miNNW Walhalla 1.vii 19831760′ hdwd forest litter J. Pakaluk #148 ( DSC)(1M). * TENNESSEE: Blount Co. GSMNP, App. Tr. ∼ 0.5miE Thunderhead Mt. summit 35°34.05′N 83°43.05′W 1510m. Forest litter. 30 Jul 2004A.K.Tishechkin ( LSAM)(5F). * TENNESSEE: Blount Co. GSMNP, App. Tr. ∼ 0.5kmW Mt. Thunderhead summit at 35°34.11′N 83°42.00′W 1585m. Forest litter sifting 13 April 2006. A.K. Tishechkin ( LSAM)(4F). * USA: TN, Blount Co. 5.8mi.ECling. Dome 1.vi.1991C.E.Carlton ( LSAM)(2F). * TENNESSEE: Blount Co. GSMNP, Gregory Cave entrance. 35°33.74′N 83°51.624′W. Litter. 28 July 2004. V.Bayless, C.Carlton & A.Tishechkin ( LSAM)(1F). * TENNESSEE: Blount Co. GSMNP, lower Gregory Ridge Tr. @ 35°33.5′N 83°30.5′W. 630m.For. Litter 28 Jul 2004. A.K. Tishechkin ( LSAM)(3F). * TENNESSEE: Cocke Co. GSMNP, Albright Grove Loop Tr. Ca. 950m.Berlese near rotten longs 19 July 2003. S.O’ Keefe ( LSAM)(2F). * USATenn Cocke Co. GSMNP Cosly Creek Trail/ Lot#76-105 Oct. 15, 1976Berlesate R.Chenowith& R. T.Allen ( LSAM)(4F). * TENNESSEE: Sevier Co. GSMNP, App. Tr. Just W of Sweat Heifer Cr. Trhd. 1720m 35°37.3′N 83°24.35′W. Forest litter. 14 April 2006. C.Carlton ( LSAM)(1F). * TENNESSEE: Sevier Co. GSMNP, Indian Head Tr. 35.60944°N 83.44659°WSift litter. 5290′ 20 July 2003. S.O’ Keefe ( LSAM) (2F; 2M). *USA-TN: Sevier Co. GSMNP. Mount LeConte, Alum Cave Tr. at 35°39.33′N 83°26.75′W, 1920m, spruce-fir litter berlese Sokolov I.M 6 Oct 2008( LSAM)(3F; 3M). * TENNESSEE: Sevier Co. GSMNP, Porters Creek Tr. Sift litter near rotten logs Beetle Blitz- 2003 18 July 2003. S.O’ Keefe ( LSAM)(9F). * U.S.A., TN, Sevier Co. GSMNP, Porters Crk. Tr. 83°23′52″W, 35°40′13″N/ elv. 870 m, 18 Oct. 2001old growth berlese A.Tishechkin, A. Cline ( LSAM)(2F; 4M). * TENNESSEE: Sevier Co. GSMNP, Road Prong Trail 35°36′36″N 83°27′3″W 1580m. Leaf/moss mat litter. 20 July 2003A. Tishechkin ( LSAM)(3F; 1M). * USA: TN: Sevier Co. GSMNP, Roaring Fork Area, Rainbow Falls Tr UTM 275540 E 3949710 N, Moist forest Berlese 1Aug2002CEC ( LSAM)(6F). * TENNESSEE: Sevier Co. Trillium Gap Tr. @ 35°39.9′N 83°26.2′W 1400m. Forest litter. 29 July; 2004. A.Tishechkin ( LSAM)(1F). * USATenn.-N.Car. Sevier Co. GSMNP, Newfound Gap to Clingman’ s Dome/ Lot #76-107 Oct. 11, 1976Berlesate R.Chenowith & R. T.Allen ( LSAM)(2F). * USA: VA, Wash. Co. E. of Damascus leaf litter 3.vi.1991C.E.Carlton ( LSAM)(1M). * USA: WV. Pocahontas Co. 6 miSE Marltinon Co. forest trail #407, off Hwy 92, v-16- 1991, sift oak litter, S.O’ Keefe ( DSC)(1F). APPENDIX 2 List of taxa (with specimen label data) included in the phylogenetic analysis. SUPERTRIBE BATRISITAE TRIBE BATRISINI Reitter Arthmius bulbiferCasey, 1983: USA: AR, Montgomery Co.approx. 5 mi.W. of Little Mo.Fallsrec. Area MVLT 25.vii.1991 C.E. Carlton SUPERTRIBE EUPLECTITAE TRIBE BYTHINOPLECTINI Schaufuss Hendecameros panamenseComellini, 1985: PANAMA: ColonPr., San Lorenzo Forest. 9°17′N 79°58′W. Flight interceptFIT-C1-5. 27-28.ix.2003 A. Tishechkin– IBISCA’03 TRIBE DIMERINIRaffray Barroeuplectoidessp. Park, 1942: BELIZE: Orange Walk Dist. Rio Bravo Conservation Area La Milpa Ruins 17.IV.1995 P.W. Kovarikrot. Log. Berlesedet. C. Carlton1995 TRIBE EUPLECTINIStreubel Euplectus confluensLeConte, 1849: USA, AR, Newton Co.Buf. Nat. Riv. Kyles LandingHdwd. Berlesate 10. Mar. 1988 C.Carlton Leptoplectus pertenuis(Casey, 1884): USA, Ark, Polk Co, Shady Lake Rec. Area, 1200ft.Hdwd-pine berlese. 13 Oct. 1974A. Newton Pycnoplectus linearis(LeConte, 1849): FLORIDA: Alachua Co.29°34½′N, 82°29′W 18-X-1994 Randall W. Lundgren Flightbarrier trapin hardwood hammock det. C. Carlton1996 TRIBE JUBINIRaffray Jubussp. Schaufuss, 1872: ECUADOR, AzuayPr. 45 kmNE Cuenca, 195 elv. 2850 m, 1 Jan 1992cloud forest Berlese C. Carlton, R. Leschen Sebagasp. Raffray, 1891: BELIZE: Orange Walk Dist. Rio Bravo Cons. Area Research Station 15-20 IV 1995, F.I. T. P.W. Kovarik TRIBE MAYETIINIWinkler Mayetia domesticaSchuster, Marsh, and Park, 1959: USA: MO: Taney Co., Ozark Underground Lab 36.5789°N, 92.8641°W. 19.X.2007, soil under fungi, #998. L.F. Watrous TRIBE METOPIASINIRaffray Metopiassp. Gory, 1832: ECUADOR: Depto. Orellana, Res. Ethnica Waorani, 1 kmS Onkone Gare Camp, Trans. Ent. 0°39′10″S 76°26′W. 220m. 5 February 1996 T. L. Erwinet al. collectors Insecticidalfogging of mostly bare green leaves, some with covering of lichenous or bryophytic plants in terra firme forest. Project MAXUS Lot1427 Trans.3 Sta.7 Subtribe RhinoscepsinaBowman 1934 Rhinoscepsis bistriatusLeConte, 1878: Crescent City, Fla. Coll Hubbard & Schwarz TRIBE TRICHONYCHINIReitter Subtribe TrichonychinaReitter Foveoscapha terracola Park and Wagner, 1962: USA: Washington, Pierce Co. Mt RainierNP, 1000ftN Green Lake, 12.3 mi( 19.8 km) SE WilkesonUTM: 10 Z, 586840mE, 5203500mN elev: 3360ft 27Oct1996 E.A. Lisowski Berleseof rotting wood and forest litter #105-01 Subtribe PanaphantinaJeannel Bibloplectus cherokeeChandler, 1990: USA: LA: Bossier Par BodcauWMA. UVLT 22 May 1996 C.Carlton/ V. Moseley. Det. CECARLTON 2002 Eutyphlus dybasiPark, 1956: USA: TN: Cocke Co.GSMNP Snake Den Ridge Trailbtw 299077 3957780 & 299657 3957885 CWright June 2005 Eutyphlus spiralis Owens and Carlton, new species: USA: NC: Macon Co. 10 miSW Franklin; Backcountry info center; V-22-1991; Berlese oak & Rhododendron; S.O’ Keefe Eutyphlus prominensPark, 1956: TENNESSEE: Sevier Co.GSMNP, Trillium Gap Tr.@ 35°39.9′N 83°26.2′W 1400m. Forestlitter. 29 July 2004. A. Tishechkin Eutyphlus schmittiRaffray, 1904: USA: PA: Jeffer.Co., Cook State Forest 0.4 miS Cooksburg Mohawk Trail. VI-6-1997, DSChandler hardwood leaf litt. Eutyphlus similisLeConte, 1880: USA: Tenn. Cocke Co.GSMNP Colby CreekTrail Lot#76-106 Oct 15, 1976Berlesate R. Chenowith & R. T. Allen Eutyphlus thoracicusPark, 1956: N CAROLINA: Haywood Co. GSMNP, Caldwell Fork Tr. At UTM 30897 E 3940883 N. Moist forest Berlese. 3 August 2002. C.Carlton & N.Lowe Pseudactium carolinaeCasey, 1908: USA, TX, Sabine Co. 9 mi.E Hemphill Beech Bottom 11 May 1988beech, magnolia sifter R. Anderson Thesiastessp. Casey, 1894: Falls Church 6:iv:1919 Va EA Chapin Collector Thesiumsp. Casey, 1884: USAArk. Scot Co. 6.7 Mi. S. Inters Hwys. 270 & 71 on 71 Lot #77-103 July 25, 1977Berlesate Hardwood-Pine Chenowith, Carlton, Heiss Trimioplectus australisChandler, 1990: USA, AR, Polk Co. Rich Mt., Eagleton Overlook Berlesate R. T. Allen, 17 Feb. 1988 Subtribe TrimiinaBowman Actiastes globiferum(LeConte, 1849): Nashville Tenn H Soltau Collection Dalmosanus quercavum(Chandler, 1990): USAArk. St. Francis Co. 6.5 Mi. NW. Inters Hwys 79 & 38 on 38 Lot #77-23 March 10, 1977 Berlesate Oak RChenowith Dalmosella tenuisCasey, 1878: USA, AR, Cross Co., Village Crk. St.Pk. Camploop A Berlesate 7 Feb. 1989 C. E. Carlton Hanfordiasp. Park, 1960: BELIZE: Orange Walk Dist. Rio Bravo Conservation Area La Milparuins, 18.IV.1995 Attadebris pile, P.W. Kovarikdet. C. Carlton1995 Melba clypeata(Reitter, 1883): Virgin IS: St Croix Est. Rust-up-Twist North Shore Rd. 01Oct1987, M.A. Ivie under bark Trimiomelba dubia(LeConte, 1849): USA: TX Brewster Co.Big Bend N.P., Cattail Falls, elv. 1310m 6 Sept 1988hdwd. Berlesate R. Anderson Trimium brevicorne(Reichenbach, 1816): PQ# A20402DC 9.XII. 32 inmoss, taking from Czechoslovzk USNM Quadrelbasp. Park, 1942: VIRGIN IS: St. John Bordeaux Mt. Elv. 1150 ft. 13-AUG 1980M A Ivie Colr. Rotten Log TRIBE TROGASTRINIJeannel Subtribe TrogastrinaJeannel Conoplectus acornusCarlton, 1983: USA Ark. Fulton Co.1.4 Mi. N. Inters. Hwys.62& 289 on 289 Lot#77-186 Oct. 18, 1977 Berlesate Hardwoods R. Chenowith Subtribe RhexiinaPark Rhexiussp. LeConte, 1849: USA, Ark., Desha Co. 1.9mi.W. intersec. Hwy. 65+I59 on I59. Bottomland Hdwd. Mar. 8, 1977Chenowith 77-19 SUPERTRIBE FARONITAE Sonoma baylessaeFerro and Carlton, 2010: USATenn-N. Car. Sevier Co. GSMNP, Newfound Gap to Mt. Le Conte Lot #77- 74 June 22, 1977 Berlesate 5000′–6300′ R. Chenowith & J. Heiss SUPERTRIBE GONIACERITAE TRIBE BRACHYGLUTINIRaffray Reichenbachia howardiPark, 1958: Gueydan La Aug 7 1925EKalmbach At lights TRIBE BYTHININIRaffray Pselaptrichus perfidus Schuster and Marsh, 1956: USA: Washington, King Co. Mt.Baker- Snoqualmie Nat. Forest. 0.8 miSSW Mount Defiancesummit 11.3 miESE North BendUTM: 10Z, 607800mE, 5253180mN Elev: 3200ft, 1-ix-1996 E.A. Lisowski#523-01 berlese of moss and liverworts on wood TRIBE TYCHINIRaffray Lucifotychus cognatus(LeConte, 1874): USA: WASHINGTON, Yakima Co.#567-1 ravine north of Wildrose Campground 3.8mi( 6.1km) ENERimrock UTM: 10 Z, 649460mE, 5170760mN Elev: 2650ft, 780m 15-II-1998, EA Lisowski Berlesecreek litter
[364,428,1336,1357]
1991-07-25
AR, MVLT
Little Mo. & Area & C. E. Carlton
United States of America
Montgomery Co.
Little Mo. Falls
28
1
[469,565,1522,1543]
Supertribe
Flight & A. Tishechkin
Panama
9.283334
San Lorenzo Forest
1300
-79.96667
28
1
Colon
1995-04-17
P. W. Kovarik
Belize
Log. Berlese & C. Carlton
Rio Bravo Conservation Area La Milpa Ruins
28
1
Orange Walk
1988-03-10
AR
Newton Co. Buf. Nat. Riv. Kyles Landing & C. Carlton
United States of America
Newton Co.
Newton Co.
28
1
1994-10-18
Randall W.
Alachua Co.
C. Carlton
Alachua Co.
-82.48333
28
1
Florida
1992-01-01
R
Berlese C. Carlton & Leschen
Ecuador
2850
Cuenca
28
1
Azuay
1995-04-15
1995-04-20
1995-04-15
T
P. W. Kovarik
Belize
Rio Bravo Cons. Area Research Station
28
1
Orange Walk
1996-02-05
T, MAXUS
L. Erwin & Insecticidal & Project & Lot & Trans. & Sta.
Ecuador
220
-0.65277773
Trans. Ent.
925
-76.433334
28
220
Orellana
[939,1168,1549,1570]
1996-10-27
Pierce Co. Mt Rainier NP & SE Wilkeson & E. A. Lisowski Berlese
United States of America
Pierce Co.
305
Green Lake
Mt Rainier
28
1
Washington
1996-05-22
UVLT, V
C. Carlton & Moseley. Det. CE
United States of America
Bossier Par Bodcau
29
1
2005-06
TN
Snake Den Ridge Trail
United States of America
Cocke Co.
Cocke Co.
29
1
[362,537,433,454]
1991-05-22
NC
United States of America
Macon Co.
Back
29
1
2004-07-29
Trillium Gap Tr. & Forest & A. Tishechkin
Sevier Co.
1400
35.665
Sevier Co.
119
-83.43667
29
1
Tennessee
1997-06-06
PA
United States of America
Jeffer. Co.
Cook State Forest
29
1
1976-10-15
R, T
United States of America
Tenn. Cocke Co.
Colby Creek Trail Lot
Colby Creek
29
1
1988-05-11
TX
United States of America
Sabine Co.
Sabine Co.
29
1
[277,398,1071,1092]
R
United States of America
Anderson
29
1
AR
United States of America
Polk Co. Rich Mt.
29
1
[415,659,1336,1357]
1988-02-17
R, T
United States of America
29
1
1989-02-07
AR
Village Crk. St. Pk. Camp & A Berlesate & C. E. Carlton
United States of America
Cross Co.
Village Crk. St.
St. Pk. Camp
29
1
1995-04-18
Atta
Belize
P. W. Kovarik & C. Carlton
Rio Bravo Conservation Area La Milpa
29
1
Orange Walk
1988-09-06
TX
Brewster Co. Big Bend N. P.
United States of America
Brewster Co.
1310
Cattail Falls
29
1
[755,875,407,428]
R
Anderson
United States of America
29
1
1977-10-18
R
Mi. N. Inters. Hwys. & Lot & Berlesate Hardwoods
United States of America
Ark. Fulton Co.
Ark. Fulton Co.
29
1
[944,1190,1326,1347]
1996-09-01
Tribe
E. A. Lisowski
United States of America
King Co. Mt.
975
Elev
Snoqualmie Nat. Forest
29
1
Washington
United States of America
Yakima Co.
Wild
29
1
Washington
1998-02-15
EA Lisowski Berlese
United States of America
0
Lisowski Berlese creek
29
1
Z