Revision ofEutyphlusLeConte (Coleoptera: Staphylinidae: Pselaphinae), with Description of a New Species and Phylogenetic Placement within the Tribe Trichonychini Owens, Brittany E. Carlton, Christopher E. The Coleopterists Bulletin 2016 2016-03-31 70 1 1 29 3DF7C O. Park 1956 [310,521,778,799] Insecta Staphylinidae Eutyphlus Animalia Coleoptera 27 Arthropoda species thoracicus   Material Examined.( n= 150) *TENNESSEE: Sevier Co. GSMNP, Trillium Gap Tr. @ 35°39.9′N 83°26.2′W 1400m. Forest litter. 29 July 2004. A.Tishechkin/ LSAM0080787 ( LSAM)(slide mounted male)(description of male based on this specimen). *TENNESSEE:Sevier Co.; GSMNP, Trillium Gap Tr. @; 35°39.9′N 83°26.2′W; 1400m. Forest litter. 29 July; 2004. A.Tishechkin ( LSAM)(slide mounted female)(description of female based on this specimen). *Rocky Creek Gap Smoky Mts. Nat. Pk. Tenn. 1X:14:53, SA-HD/ Field Mus. Nat. Hist. Orlando Park Pselaphidae Colln./ (orange label) HOLOTYPE/ Eutyphlus thoracicus. Park, nsp. ( FMNH)(1M; HOLOTYPE). *same locality/ (orange label) PARATYPE. ( FMNH) (2F). *Chimney Cp. Gr Smoky Mts. N. P. Tenn.16: IX:53 Auerbach #5/ Field Mus. Nat. Hist Orlando Park Pselaphidae Colln./ (orange label) PARATYPE( FMNH)(1F). *Chimney Cp. Gr Smoky Mts. N. P. Tenn.16:IX:53 Auerbach #5/ Field Mus. Nat. Hist Orlando Park Pselaphidae Colln./ (orange label) PARATYPE( FMNH)(1F). *Cloudland Canyon S. Pk. Dade Co., GA. 14. VI.62 Forest floor debris/ H. R. Steeves, Jr. J.D.Patrick Jr. Collectors/ H. R. Steeves Jr. Collection ( FMNH)(5F). *N CAROLINA: Haywood Co. GSMNP, Caldwell Fork Tr. At UTM 30897 E 3940883 N. Moist forest Berlese. 3 August 2002. C.Carlton & N. Lowe ( LSAM)(18F; 2M) * USA: NCHaywood Co. GSMNP McKee Branch Tr. 35°35′6″N 83°05′W/ 1370m 16Jul–12Aug2002FIT C.Carlton & S.Gil ( LSAM)(1M). *N CAROLINA: Haywood Co. GSMNP, Mt. Sterling Gap. 35°42.0′N 83°5.9′W 1190m. Forest litter. 29 July 2004. R. T.Allen/ LSAM0095711 ( LSAM)(1F). *same locality ( LSAM)(16F; 1M). * USA: NC: Haywood Co. GSMNP Purchase Knob, berlese/ 313124E 3939640N 5089ft. 20 Jul 2002CCarlton (3F; 1M). * USA: NC: Macon Co. 3 miNW Highlands v-29- 1983, sift rhododendron litter, DSChandler ( DSC) (7F). * USA: NC: Swain Co GSMNP Appalachian Tr. 1.5 miNE Newfound Gap/ 281639E 3944038N ridgetop berlese 18 Jul 2002CCarlton ( LSAM)(3F). *N CAROLINA: Swain Co. Appalachian Tr. ∼ 1.5miE Newfound Gap. 35°36.96′N 83°24.66′W 1730m. Forest litter; 18 July 2002. C.Carlton ( LSAM)(1F). * U.S.A., NC, Swain Co.; GSMNP, Beech Gap Tr.; 83°12′32″W, 35°38′23″N/ elv. 1400m, 20, Oct. 2001; N. hardwood berlese; LSAM, C.Carlton ( LSAM)(2F). *N CAROLINA: Swain Co. GSMNP, jct. of Beech Gap & Hyatt Ridge Trails 35°37.82′N 83°42.07′W 1495m. Forest litter. 2 Oct 2005. W.D. Merritt ( LSAM)(3F). * USAN.C. Swain Co. Dirt Ri. From Heintooga Overlook to Cherokee/ Lot #76-102 Oct. 14, 1976Berlesate 5000′- 4500′ R.Chenowith & R.Tallen ( LSAM)(7F). * U.S.A., NC, Swain Co. GSMNP, Flat Creek Trail 83°10′21″W, 35°33′1″N/ elv. 1500m, leaf litter Berlese, 31 July 2001. A. Tishechkin ( LSAM) (1M). *N CAROLINA: Swain Co. GSMNP, Jenkins Ridge Tr. ∼ 1mifrom Appalachian Tr. 35°33.75′N 83°43.2′W 1540m. Forest litter. 30 Jul 2004. A.K. Tishechkin ( LSAM)(3F). *N CAROLINA: Swain Co. GSMNP, upper Jenkins Ridge Tr. At 35°33.75′N 83°43.2′W. 1540m. Leaf litter 31 Jul 2004. A.K. Tishechkin ( LSAM)(1F). *N. CAR. Swain Co. Mt. Collins 5900′ v.21.1977 A.Newton, M.Thayer ( DENH)(2F). *N CAROLINA: Swain Co. GSMNP, Noland Divide Tr. at 35°33′57″N 83°28′36″W 1770m.Leaf/moss mat litter 19 July 2003. A. Tishechkin ( LSAM)(2F). * USA: NC. Yancey Co. Black Mtn. info cntr v-21-1991sift fir-oak S.O’ Keefe collr ( DSC) (3M). * USA: NC, Yancy Co. B. R. Pkwy. 37.4 mi.SW intersection US221 elv. 4600′ log litter 2.vi.1991C.E.Carlton ( LSAM)(1F). * SC: Oconee Co. 6.7 miNNW Walhalla 1.vii 19831760′ hdwd forest litter J. Pakaluk #148 ( DSC)(1M). * TENNESSEE: Blount Co. GSMNP, App. Tr. ∼ 0.5miE Thunderhead Mt. summit 35°34.05′N 83°43.05′W 1510m. Forest litter. 30 Jul 2004A.K.Tishechkin ( LSAM)(5F). * TENNESSEE: Blount Co. GSMNP, App. Tr. ∼ 0.5kmW Mt. Thunderhead summit at 35°34.11′N 83°42.00′W 1585m. Forest litter sifting 13 April 2006. A.K. Tishechkin ( LSAM)(4F). * USA: TN, Blount Co. 5.8mi.ECling. Dome 1.vi.1991C.E.Carlton ( LSAM)(2F). * TENNESSEE: Blount Co. GSMNP, Gregory Cave entrance. 35°33.74′N 83°51.624′W. Litter. 28 July 2004. V.Bayless, C.Carlton & A.Tishechkin ( LSAM)(1F). * TENNESSEE: Blount Co. GSMNP, lower Gregory Ridge Tr. @ 35°33.5′N 83°30.5′W.  630m.For. Litter 28 Jul 2004. A.K. Tishechkin ( LSAM)(3F). * TENNESSEE: Cocke Co. GSMNP, Albright Grove Loop Tr. Ca.  950m.Berlese near rotten longs 19 July 2003. S.O’ Keefe ( LSAM)(2F). * USATenn Cocke Co. GSMNP Cosly Creek Trail/ Lot#76-105 Oct. 15, 1976Berlesate R.Chenowith& R. T.Allen ( LSAM)(4F). * TENNESSEE: Sevier Co. GSMNP, App. Tr. Just W of Sweat Heifer Cr. Trhd. 1720m 35°37.3′N 83°24.35′W. Forest litter. 14 April 2006. C.Carlton ( LSAM)(1F). * TENNESSEE: Sevier Co. GSMNP, Indian Head Tr. 35.60944°N 83.44659°WSift litter. 5290′ 20 July 2003. S.O’ Keefe ( LSAM) (2F; 2M). *USA-TN: Sevier Co. GSMNP. Mount LeConte, Alum Cave Tr. at 35°39.33′N 83°26.75′W, 1920m, spruce-fir litter berlese Sokolov I.M 6 Oct 2008( LSAM)(3F; 3M). * TENNESSEE: Sevier Co. GSMNP, Porters Creek Tr. Sift litter near rotten logs Beetle Blitz- 2003 18 July 2003. S.O’ Keefe ( LSAM)(9F). * U.S.A., TN, Sevier Co. GSMNP, Porters Crk. Tr. 83°23′52″W, 35°40′13″N/ elv. 870 m, 18 Oct. 2001old growth berlese A.Tishechkin, A. Cline ( LSAM)(2F; 4M). * TENNESSEE: Sevier Co. GSMNP, Road Prong Trail 35°36′36″N 83°27′3″W 1580m. Leaf/moss mat litter. 20 July 2003A. Tishechkin ( LSAM)(3F; 1M). * USA: TN: Sevier Co. GSMNP, Roaring Fork Area, Rainbow Falls Tr UTM 275540 E 3949710 N, Moist forest Berlese 1Aug2002CEC ( LSAM)(6F). * TENNESSEE: Sevier Co. Trillium Gap Tr. @ 35°39.9′N 83°26.2′W 1400m. Forest litter. 29 July; 2004. A.Tishechkin ( LSAM)(1F). * USATenn.-N.Car. Sevier Co. GSMNP, Newfound Gap to Clingman’ s Dome/ Lot #76-107 Oct. 11, 1976Berlesate R.Chenowith & R. T.Allen ( LSAM)(2F). * USA: VA, Wash. Co. E. of Damascus leaf litter 3.vi.1991C.E.Carlton ( LSAM)(1M). * USA: WV. Pocahontas Co. 6 miSE Marltinon Co. forest trail #407, off Hwy 92, v-16- 1991, sift oak litter, S.O’ Keefe ( DSC)(1F).  APPENDIX 2 List of taxa (with specimen label data) included in the phylogenetic analysis.   SUPERTRIBE BATRISITAE   TRIBE  BATRISINI Reitter  Arthmius bulbiferCasey, 1983: USA: AR, Montgomery Co.approx. 5 mi.W. of  Little Mo.Fallsrec. Area MVLT  25.vii.1991 C.E. Carlton   SUPERTRIBE EUPLECTITAE   TRIBE  BYTHINOPLECTINI Schaufuss  Hendecameros panamenseComellini, 1985: PANAMA: ColonPr., San Lorenzo Forest. 9°17′N 79°58′W. Flight interceptFIT-C1-5. 27-28.ix.2003 A. Tishechkin– IBISCA’03   TRIBE  DIMERINIRaffray  Barroeuplectoidessp. Park, 1942:   BELIZE: Orange Walk Dist. Rio Bravo Conservation Area La Milpa Ruins  17.IV.1995 P.W. Kovarikrot. Log. Berlesedet. C. Carlton1995   TRIBE  EUPLECTINIStreubel   Euplectus confluensLeConte, 1849:   USA, AR,  Newton Co.Buf. Nat. Riv. Kyles LandingHdwd. Berlesate  10. Mar. 1988 C.Carlton   Leptoplectus pertenuis(Casey, 1884):  USA, Ark, Polk Co, Shady Lake Rec. Area, 1200ft.Hdwd-pine berlese. 13 Oct. 1974A. Newton   Pycnoplectus linearis(LeConte, 1849):  FLORIDA: Alachua Co.29°34½′N, 82°29′W  18-X-1994 Randall W. Lundgren Flightbarrier trapin hardwood hammock det. C. Carlton1996   TRIBE  JUBINIRaffray   Jubussp. Schaufuss, 1872:   ECUADOR, AzuayPr. 45 kmNE Cuenca, 195 elv.  2850 m,  1 Jan 1992cloud forest Berlese C. Carlton, R. Leschen   Sebagasp. Raffray, 1891:   BELIZE: Orange Walk Dist. Rio Bravo Cons. Area Research Station  15-20 IV 1995, F.I. T. P.W. Kovarik   TRIBE  MAYETIINIWinkler  Mayetia domesticaSchuster, Marsh, and Park, 1959: USA: MO: Taney Co., Ozark Underground Lab 36.5789°N, 92.8641°W. 19.X.2007, soil under fungi, #998. L.F. Watrous   TRIBE  METOPIASINIRaffray   Metopiassp. Gory, 1832:   ECUADOR: Depto. Orellana, Res. Ethnica Waorani,  1 kmS Onkone Gare Camp, Trans. Ent. 0°39′10″S 76°26′W.   220m.  5 February 1996 T. L. Erwinet al. collectors Insecticidalfogging of mostly bare green leaves, some with covering of lichenous or bryophytic plants in terra firme forest. Project MAXUS Lot1427 Trans.3 Sta.7  Subtribe  RhinoscepsinaBowman 1934   Rhinoscepsis bistriatusLeConte, 1878: Crescent City, Fla. Coll Hubbard & Schwarz   TRIBE  TRICHONYCHINIReitter  Subtribe  TrichonychinaReitter   Foveoscapha terracola Park and Wagner, 1962: USA: Washington,  Pierce Co. Mt RainierNP,  1000ftN Green Lake, 12.3 mi( 19.8 km) SE WilkesonUTM: 10 Z, 586840mE, 5203500mN elev:  3360ft  27Oct1996 E.A. Lisowski Berleseof rotting wood and forest litter #105-01  Subtribe  PanaphantinaJeannel   Bibloplectus cherokeeChandler, 1990:   USA: LA: Bossier Par BodcauWMA. UVLT  22 May 1996 C.Carlton/ V. Moseley. Det. CECARLTON 2002   Eutyphlus dybasiPark, 1956:   USA: TN: Cocke Co.GSMNP Snake Den Ridge Trailbtw 299077 3957780 & 299657 3957885 CWright  June 2005   Eutyphlus spiralis Owens and Carlton, new species: USA: NC: Macon Co.  10 miSW Franklin; Backcountry info center;  V-22-1991; Berlese oak & Rhododendron; S.O’ Keefe   Eutyphlus prominensPark, 1956:   TENNESSEE: Sevier Co.GSMNP, Trillium Gap Tr.@ 35°39.9′N 83°26.2′W  1400m. Forestlitter.  29 July 2004. A. Tishechkin   Eutyphlus schmittiRaffray, 1904:   USA: PA: Jeffer.Co., Cook State Forest  0.4 miS Cooksburg Mohawk Trail.  VI-6-1997, DSChandler hardwood leaf litt.   Eutyphlus similisLeConte, 1880:   USA: Tenn. Cocke Co.GSMNP  Colby CreekTrail Lot#76-106  Oct 15, 1976Berlesate R. Chenowith & R. T. Allen   Eutyphlus thoracicusPark, 1956: N CAROLINA: Haywood Co. GSMNP, Caldwell Fork Tr. At UTM 30897 E 3940883 N. Moist forest Berlese. 3 August 2002. C.Carlton & N.Lowe   Pseudactium carolinaeCasey, 1908:   USA, TX, Sabine Co.  9 mi.E Hemphill Beech Bottom  11 May 1988beech, magnolia sifter  R. Anderson   Thesiastessp. Casey, 1894: Falls Church 6:iv:1919 Va EA Chapin Collector   Thesiumsp. Casey, 1884:  USAArk. Scot Co. 6.7 Mi. S. Inters Hwys. 270 & 71 on 71 Lot #77-103 July 25, 1977Berlesate Hardwood-Pine Chenowith, Carlton, Heiss   Trimioplectus australisChandler, 1990:   USA, AR, Polk Co. Rich Mt., Eagleton Overlook Berlesate  R. T. Allen,  17 Feb. 1988  Subtribe  TrimiinaBowman   Actiastes globiferum(LeConte, 1849): Nashville Tenn H Soltau Collection   Dalmosanus quercavum(Chandler, 1990): USAArk. St. Francis Co. 6.5 Mi. NW. Inters Hwys 79 & 38 on 38 Lot #77-23 March 10, 1977 Berlesate Oak RChenowith   Dalmosella tenuisCasey, 1878:   USA, AR, Cross Co.,  Village Crk. St.Pk. Camploop A Berlesate  7 Feb. 1989 C. E. Carlton   Hanfordiasp. Park, 1960:   BELIZE: Orange Walk Dist. Rio Bravo Conservation Area La Milparuins,  18.IV.1995 Attadebris pile, P.W. Kovarikdet. C. Carlton1995   Melba clypeata(Reitter, 1883): Virgin IS: St Croix Est. Rust-up-Twist North Shore Rd. 01Oct1987, M.A. Ivie under bark  Trimiomelba dubia(LeConte, 1849):   USA: TX  Brewster Co.Big Bend N.P., Cattail Falls, elv.  1310m  6 Sept 1988hdwd. Berlesate  R. Anderson   Trimium brevicorne(Reichenbach, 1816): PQ# A20402DC 9.XII. 32 inmoss, taking from Czechoslovzk USNM   Quadrelbasp. Park, 1942: VIRGIN IS: St. John Bordeaux Mt. Elv. 1150 ft. 13-AUG 1980M A Ivie Colr. Rotten Log   TRIBE  TROGASTRINIJeannel  Subtribe  TrogastrinaJeannel   Conoplectus acornusCarlton, 1983:   USA Ark. Fulton Co.1.4 Mi. N. Inters. Hwys.62& 289 on 289 Lot#77-186  Oct. 18, 1977 Berlesate Hardwoods R. Chenowith  Subtribe  RhexiinaPark   Rhexiussp. LeConte, 1849:  USA, Ark., Desha Co. 1.9mi.W. intersec. Hwy. 65+I59 on I59. Bottomland Hdwd. Mar. 8, 1977Chenowith 77-19   SUPERTRIBE FARONITAE  Sonoma baylessaeFerro and Carlton, 2010: USATenn-N. Car. Sevier Co. GSMNP, Newfound Gap to Mt. Le Conte Lot #77- 74 June 22, 1977 Berlesate 5000′–6300′ R. Chenowith & J. Heiss   SUPERTRIBE GONIACERITAE TRIBE  BRACHYGLUTINIRaffray   Reichenbachia howardiPark, 1958: Gueydan La Aug 7 1925EKalmbach At lights   TRIBE  BYTHININIRaffray   Pselaptrichus perfidus Schuster and Marsh, 1956: USA: Washington, King Co. Mt.Baker- Snoqualmie Nat. Forest.  0.8 miSSW Mount Defiancesummit  11.3 miESE North BendUTM: 10Z, 607800mE, 5253180mN Elev:  3200ft,  1-ix-1996 E.A. Lisowski#523-01 berlese of moss and liverworts on wood   TRIBE  TYCHINIRaffray   Lucifotychus cognatus(LeConte, 1874):   USA: WASHINGTON, Yakima Co.#567-1 ravine north of Wildrose Campground 3.8mi( 6.1km) ENERimrock UTM: 10  Z, 649460mE, 5170760mN Elev:  2650ft, 780m  15-II-1998, EA Lisowski Berlesecreek litter [364,428,1336,1357] 1991-07-25 AR, MVLT Little Mo. & Area & C. E. Carlton United States of America Montgomery Co. Little Mo. Falls 28 1 [469,565,1522,1543] Supertribe Flight & A. Tishechkin Panama 9.283334 San Lorenzo Forest 1300 -79.96667 28 1 Colon 1995-04-17 P. W. Kovarik Belize Log. Berlese & C. Carlton Rio Bravo Conservation Area La Milpa Ruins 28 1 Orange Walk 1988-03-10 AR Newton Co. Buf. Nat. Riv. Kyles Landing & C. Carlton United States of America Newton Co. Newton Co. 28 1 1994-10-18 Randall W. Alachua Co. C. Carlton Alachua Co. -82.48333 28 1 Florida 1992-01-01 R Berlese C. Carlton & Leschen Ecuador 2850 Cuenca 28 1 Azuay 1995-04-15 1995-04-20 1995-04-15 T P. W. Kovarik Belize Rio Bravo Cons. Area Research Station 28 1 Orange Walk 1996-02-05 T, MAXUS L. Erwin & Insecticidal & Project & Lot & Trans. & Sta. Ecuador 220 -0.65277773 Trans. Ent. 925 -76.433334 28 220 Orellana [939,1168,1549,1570] 1996-10-27 Pierce Co. Mt Rainier NP & SE Wilkeson & E. A. Lisowski Berlese United States of America Pierce Co. 305 Green Lake Mt Rainier 28 1 Washington 1996-05-22 UVLT, V C. Carlton & Moseley. Det. CE United States of America Bossier Par Bodcau 29 1 2005-06 TN Snake Den Ridge Trail United States of America Cocke Co. Cocke Co. 29 1 [362,537,433,454] 1991-05-22 NC United States of America Macon Co. Back 29 1 2004-07-29 Trillium Gap Tr. & Forest & A. Tishechkin Sevier Co. 1400 35.665 Sevier Co. 119 -83.43667 29 1 Tennessee 1997-06-06 PA United States of America Jeffer. Co. Cook State Forest 29 1 1976-10-15 R, T United States of America Tenn. Cocke Co. Colby Creek Trail Lot Colby Creek 29 1 1988-05-11 TX United States of America Sabine Co. Sabine Co. 29 1 [277,398,1071,1092] R United States of America Anderson 29 1 AR United States of America Polk Co. Rich Mt. 29 1 [415,659,1336,1357] 1988-02-17 R, T United States of America 29 1 1989-02-07 AR Village Crk. St. Pk. Camp & A Berlesate & C. E. Carlton United States of America Cross Co. Village Crk. St. St. Pk. Camp 29 1 1995-04-18 Atta Belize P. W. Kovarik & C. Carlton Rio Bravo Conservation Area La Milpa 29 1 Orange Walk 1988-09-06 TX Brewster Co. Big Bend N. P. United States of America Brewster Co. 1310 Cattail Falls 29 1 [755,875,407,428] R Anderson United States of America 29 1 1977-10-18 R Mi. N. Inters. Hwys. & Lot & Berlesate Hardwoods United States of America Ark. Fulton Co. Ark. Fulton Co. 29 1 [944,1190,1326,1347] 1996-09-01 Tribe E. A. Lisowski United States of America King Co. Mt. 975 Elev Snoqualmie Nat. Forest 29 1 Washington United States of America Yakima Co. Wild 29 1 Washington 1998-02-15 EA Lisowski Berlese United States of America 0 Lisowski Berlese creek 29 1 Z