Arvelius edessoides SPINOLA, 1837:347 .
Taurocerus hector STÅL, 1862:108
WALKER, 1867:391
DALLAS, 1851:302
DISTANT, 1880:82
Stål, 1872:47
LETHIERRY & SEVERIN, 1893:185
KIRKALDY, 1909:150
Revision of Taurocerus (Heteroptera, Pentatomidae, Pentatomini)
Jocélia Grazia
Aline Barcellos
Iheringia, Sér. Zool.
2005
2005-06-30
95
2
173
181
edessoides (Spinola, 1837)
edessoides (Spinola
1837
[314,736,218,242]
Insecta
Pentatomidae
Taurocerus
GBIF
Animalia
Hemiptera
6
179
Arthropoda
species
edessoides
( Figs. 6, 9, 12, 15, 18, 21, 24, 26, 28)
Arvelius edessoides SPINOLA, 1837:347. Taurocerus hector STÅL, 1862:108; 1872:47; WALKER, 1867:391(list). Taurocerus edessoides; DALLAS, 1851:302(list); DISTANT, 1880:82, pl. 7, fig. 25 (list); Stål, 1872:47; LETHIERRY & SEVERIN, 1893:185(cat.); KIRKALDY, 1909:150(cat.).
Type-locality. Brazil.
Typematerial. Lectotypeand paralectotype, here designated. The material belongs to Spinola’s collection, at present deposited at MRSN. A green label in the box, containing the following data: Arvelius edessoidesSpin., G. taurocerus, A. et S., D. Bouquet, Brésil. Typematerial of T. hectorexamined. Stål described this species based on two males from Mexico and one female from unknown locality. We received from NHRS three males labeled as “ typus” (loan number 1068/99) and “ paratypus” (numbers 1069/99 and 1070/99), the last one collected in Bogotá [Colombia] by Alexander Lindig. We believe that the specimen 1070/99 is not actually a syntypeof T. hectorand the female mentioned in the Stål’s original description is missed. Then, we here designate as lectotypeand paralectotype, respectively: 1, labeled: a) Vera Cruz [handwritten]; b) HectorTyp. Stål [handwritten]; c) Sallé; d) Typus[red label]; e) Naturhistoriska Riksmuseet Stockholm Loan no 1068/99; 1, labeled: a) Mexico; b) Stål; c) HectorTyp. Stål [handwriting]; d) Paratypus; e) Naturhistoriska Riksmuseet Stockholm Loan number 1069/99. Description. Dorsal surface reddish-castaneous, abdominal venter reddish. Apex of humeri yellow. Rostrum reaching the fourth abdomninal sternite. Vshaped callous of scutellum, in the most of the specimens examined, with broad and closed arms, becoming cordiform in appearance; in some specimens, however, the callous is faded. Male. Measurements (n = 10). Total length 15.56 (14.63-16.50) ± 0.73; head length 2.96 (2.69-3.16) ± 0.13; width 3.27 (3.06-3.50) ± 0.15; eye width 0.74 (0.63-0.81) ± 0.06; interocular distance 1.80 (1.69-1.94) ± 0.08; anteocular length 1.47 (1.38-1.56) ± 0.07; antennal segments I, 0.88 (0.81-1.06) ± 0.08; II, 1.47 (1.25-1.63) ± 0.10; III, 1.94 (1.63-2.44) ± 0.22; IV, 3.13 (2.78-3.56) ± 0.22; V, 3.24 (3.06-3.44) ± 0.14; pronotal length 3.81 (3.63- 4.25) ± 0.24; width 13.62 (11.88-15.00) ± 1.19; scutellum length 6.70 (6.25-7.25) ± 0.36; width 5.98 (5.50-6.50) ± 0.35; corium length 9.34 (8.50-10.00) ± 0.50; abdominal width 9.24 (8.75-9.63) ± 0.35. Genitalia. Pygophore quadrangular, globose in caudal view, little open dorsally. Dorsal rim slightly sinuated. Proctiger trapezoidal, wider basally, with 1 + 1 tumid areas at the distal half ( fig. 6). Parameres distally thin ( fig. 15), apex strongly curved latero-posteriorly ( fig. 9). Posterolateral angles with a small, acute tubercle. Ventral rim: median process acute; lateromedian processes small, conical, delimiting a median V-shaped excavation ( fig. 12). Phallus( figs. 18, 21, 24): basal plate welldeveloped, half as long as phallotheca.Visible part of processus phallothecaelong, more than two thirds as long as phallotheca. Vesica well-developed with one median, ventral process. Ductus seminis distalislong, surpassing conjunctiva. Conjunctiva well-developed, with 1 + 1 dorsal processes (prcj1); prcj2 absent. Female. Measurements (n = 10). Total length 15.56 (14.25-17.00) ± 0.98; head length 2.87 (2.66-3.19) ± 0.13; width 3.28 (3.13-3.44) ± 0.10; eye width 0.77 (0.69-0.81) ± 0.04; interocular distance 1.80 (1.69-1.88) ± 0.08; anteocular length 1.43 (1.31-1.56) ± 0.07; antennal segments I, 0.92 (0.81-1.06) ± 0.07; II, 1.40 (1.25-1.56) ± 0.10; III, 1.83 (1.63-2.00) ± 0.11; IV, 3.06 (2.81-3.25) ± 0.15; V, 3.18 (3.06-3.25) ± 0.10; pronotal length 3.81 (3.56-4.00) ± 0.17; width 13.41 (12.13-14.75) ± 0.78; scutellum length 6.73 (6.25-7.19) ± 0.30; width 6.01 (5.75-6.38) ± 0.21; corium length 9.31 (8.88-10.00) ± 0.42; abdominal width 9.29 (8.88-9.75) ± 0.32. Genitalia. Gonocoxites 8 triangular, about one and a half wider than long ( fig. 26). Sutural borders of gonocoxites 8 convergent, contiguous at apical half. Posterior borders concave near laterotergites 9. Laterotergites 9 narrow at apex. Secondary thickenings of gonapophysis 9 ear-like ( fig. 28).
Material examined. MEXICO, Veracruz: playa escondida 28 Km NE of Catemaco, 50 m),, 30.IX.[19]74, M. Madisoncol. ( AMNH); Los Tuxtlas(“ camino a Balzapotl”),, 11.IX.[19]86, L. Cervantescol.(10:45 80, Siparuna) ( AMNH); idem,, 15.VI.[19]86, idem (13:20, Siparuna), idem ( AMNH); GUATEMALA, Alta Verapaz: San Juan,, Championcol. ( NHMW); COSTA RICA, Guanacaste: Santa Rosa (PN Guanacaste),, 20.X.[19]96, F. Fernándezcol. (IAvH 05098); PANAMA, Panama: Distr. Chame, Cerro Campana( 800 m),, 24.VII.[19]82, H.D. Englemancol. ( AMNH); idem,, 3.V.[19]81, idem ( AMNH); idem,, 10.V.[19]81, E. Giesbertcol. ( AMNH); Canal Zone: Barro Colorado,, II-III.1936, W.J. Gertschcol. ( AMNH); COLÔMBIA, Meta: Caño Grande( 450 m),, II.[19]48, Richtercol. ( UNCBHe 00435); Amazonas: Araracuara,,, 21.VIII.[19]77, R. Restrepocol. ( UNCB He 00376, He 00377); ECUADOR, Napo: Limoncocha( 00 o 24’S, 76 o 36’W),, 7.VIII.1972, Peter L. Kazancol. ( AMNH); GUYANA, Cuyuni- Mazaruni: Bartica,, VIII, Pomeratcol. ( AMNH); BRAZIL, C. Samuel, Jamari,, III.1944, Parkocol. ( UFRG); Amazonas: Benjamin Constant( Rio Javari),, III.1942, August Rabautcol. ( AMNH); idem,, 25.I-15.II.1942( AMNH); Querari( Pelotão), 01 o 05’N/ 69 o 51’W,, 20-21.IV.1993, C. Motta, R. Ferreira, J. Vidal, B. Matteocol. (“ luz mista de mercúrio e luz negra BL e BLB”) ( INPA); Pará: Belém( Utinga),, 20.I.1977, N. Guimarãescol. ( MPEG); Acre: Feijó,, 2, XI.1956( MZSP); idem,, XII.1956( MZSP); Mato Grosso: Chapada [dos Guimarães],, IX, acc.13739 ( AMNH); idem,, VIII, acc. 23739 ( AMNH); Rondônia: Porto Velho( Terr. Guaporé),, X.[19]44, Parkocol. ( UFRG); Vilhena,, 11.XII.1986, C. Eliascol., Polonoroeste( DZUP); Rolim de Moura( Linha 204),, 31.X.1978, K. S. Brown Jr.col. (“ em mato aberto, terra roxa”) ( ZUEC 2544); Minas Gerais: C[armo do] R[io] Claro,, I.[19]45, Carvalhocol. ( MNRJ); Espírito Santo: Baixo Guandu,, 23-30.XI.1970, C. Eliascol. ( DZUP); Santa Leopoldina,, 3.IX.1964, Claudionor Eliascol. ( DZUP); São Paulo: Tupã,, V.1947, Fritz Plaumanncol., Ex. Coll. H. Ruckes ( AMNH); Severinia,, XII.1940, A.G. Silvacol. ( MCNZ 003616); Osasco,, XII.1957( MZSP); PERU, Loreto: Iquitos,, no date, F6062, H. Bassler Collection, Acc. 33591 ( AMNH); Upper Rio Maranon,, 20.IV.[19]24, F6029, idem ( AMNH); idem,, 5.X.[19]24, F6103, H. Bassler Collection, Acc. 33591 ( AMNH); idem, 3, 11.X.[19]24, F6102, idem ( AMNH); idem,, 11.X.[19]24, F6102, idem ( AMNH); Amazonas: Rio Santiago,, 06.XI.1928, F619F, H. Bassler Collection, Acc. 33591 ( AMNH); idem,, 27.XI.[19]24, F6012, idem ( AMNH); idem,, 23.XI.[19]24, F6012, idem ( AMNH); Rio Ucayali,,, 27.IX.[19]23, F6004, idem ( AMNH); Ucayali: Pucallpa, Rio Ucayali, 200m,, no date, Diringscol., Coll. Dirings ( MCNZ 003585); Huánuco: Tingo María,, 18-23.IX.[19]76, Jeff Baiercol. ( AMNH); Cuzco: Pilcopata,, 11.X.68, (“arbusto, 1604”) ( UFRG); BOLIVIA, Cochabamba: Chaparé( 400 m),, 12.XII.1949, Diringscol. ( MZSP).
Comments. The color of the specimens varies from reddish to dark reddish-castaneous; the length of humeri is also variable, but the apices are always yellowish. The size and the triangular shape of gonocoxites 8, combined with the color of humeri distinguish T. edessoidesfrom T. achillesand T. abruptus.
2561150655
30. IX. [19] 74
AMNH
M. Madison
Mexico
50
Veracruz
Veracruz
6
179
1
1
2561150645
11. IX. [19] 86
AMNH
L. Cervantes
Mexico
camino a Balzapotl
Los Tuxtlas
6
179
1
1
2561150631
[861,1455,959,978]
15. VI. [19] 86
AMNH
Mexico
Siparuna
6
179
1
1
2561150638
NHMW
Champion
Guatemala
San Juan,
7
180
1
1
Alta Verapaz
2561150670
20. X. [19] 96
PN
F. Fernandez
Costa Rica
Santa Rosa
7
180
1
1
Guanacaste
2561150669
24. VII. [19] 82
AMNH
H. D. Engleman
Panama
800
Distr. Chame
Distr. Chame
7
180
1
1
2561150658
3. V. [19] 81
AMNH
H. D. Engleman
Panama
800
Cerro Campana
Distr. Chame
7
180
1
1
2561150635
[278,722,1888,1907]
10. V. [19] 81
AMNH
E. Giesbert
Panama
800
Cerro Campana
Distr. Chame
7
180
1
1
2561150672
II-III. 1936
AMNH
W. J. Gertsch
Panama
Barro Colorado
Barro Colorado
7
180
1
1
Canal Zone
2561150660
II. [19] 48
UNCB
Richter
Colombia
450
Cano Grande
Cano Grande
7
180
1
1
Meta
2561150650
21. VIII. [19] 77
UNCB
R. Restrepo
Colombia
Araracuara
Araracuara
7
180
He 00376, He 00377
1
1
1
Amazonas
2561150632
[129,1162,1768,2027]
1972-08-07
AMNH
Ecuador
-0.4
Limoncocha
1308
-76.6
Napo
7
180
1
1
2561150663
AMNH
Pomerat
Guyana
Bartica
Cuyuni- Mazaruni
7
180
1
1
2561150633
1944-03
UFRG
Parko
Brazil
Jamari
Jamari
7
180
1
1
2561150680
1942-03
AMNH
August Rabaut
Brazil
Rio Javari
Benjamin Constant
7
180
1
1
Amazonas
2561150674
[861,1180,1864,1883]
1942-01-25
1942-02-15
1942-01-25
AMNH
August Rabaut
Brazil
Rio Javari
Benjamin Constant
7
180
1
1
2561150662
1993-04-20
1993-04-21
1993-04-20
luz mista de mercurio e luz negra BL e BLB
INPA
C. Motta & R. Ferreira & J. Vidal & B. Matteo
Brazil
1.0833334
Pelotao
1308
-69.85
Querari
7
180
1
1
2561150636
1977-01-20
MPEG
N. Guimaraes
Brazil
Utinga
Belem
7
180
1
1
Para
2561150664
[902,1270,1959,1978]
1956-11
MZSP
Brazil
Feijo
7
180
1
2
1
Acre
2561150657
1956-12
MZSP
Brazil
Feijo
7
180
1
1
Acre
2561150684
AMNH
Brazil
Chapada [dos Guimaraes]
Chapada [dos Guimaraes]
7
180
1
1
Mato Grosso
2561150644
[1008,1357,2007,2026]
AMNH
Brazil
Chapada [dos Guimaraes]
7
180
1
1
Mato Grosso
2561150673
X. [19] 44
UFRG
Parko
Brazil
Terr. Guapore
Porto Velho
8
181
1
1
Rondonia
2561150642
1986-12-11
DZUP
C. Elias
Brazil
Polonoroeste
Vilhena
8
181
1
1
2561150661
em mato aberto, terra roxa
1978-10-31
ZUEC
K. S. Brown Jr.
Brazil
Linha 204
Rolim de Moura
8
181
ZUEC 2544
1
1
2561150654
I. [19] 45
MNRJ
Carvalho
Brazil
C [armo do] R [io] Claro
Minas Gerais
8
181
1
1
2561150671
1970-11-23
1970-11-30
1970-11-23
DZUP
C. Elias
Brazil
Baixo Guandu
Baixo Guandu
8
181
1
1
Espirito Santo
2561150641
1964-09-03
DZUP
Claudionor Elias
Brazil
Santa Leopoldina
Santa Leopoldina
8
181
1
1
2561150665
1947-05
AMNH
Fritz Plaumann
Brazil
Tupa
Sao Paulo
8
181
1
1
2561150634
1940-12
MCNZ
A. G. Silva
Brazil
Severinia
Severinia
8
181
MCNZ 003616
1
1
2561150682
[487,751,434,453]
1957-12
MZSP
Brazil
Osasco
Osasco
8
181
1
1
2561150656
AMNH
H. Bassler Collection
Peru
Iquitos
8
181
1
1
Loreto
2561150678
20. IV. [19] 24
AMNH
Peru
Upper Rio Maranon
8
181
1
1
2561150691
5. X. [19] 24
AMNH
H. Bassler Collection
Peru
Upper Rio Maranon
8
181
1
1
2561150651
11. X. [19] 24
AMNH
Peru
Upper Rio Maranon
8
181
1
3
2561150676
[337,822,554,573]
11. X. [19] 24
AMNH
Peru
Upper Rio Maranon
8
181
1
1
2561150692
1928-11-06
AMNH
H. Bassler Collection
Peru
Rio Santiago
Rio Santiago
8
181
1
1
Amazonas
2561150637
27. XI. [19] 24
AMNH
Peru
Rio Santiago
8
181
1
1
Amazonas
2561150652
[381,821,626,645]
23. XI. [19] 24
AMNH
Peru
Rio Santiago
8
181
1
Amazonas
2561150688
[227,736,650,669]
27. IX. [19] 23
AMNH
Peru
Rio Ucayali
8
181
1
1
1
Amazonas
2561150639
MCNZ
Dirings
Ucayali
200
Rio Ucayali
Pucallpa
8
181
MCNZ 003585
1
1
Ucayali
2561150648
2018-09-23
AMNH
Jeff Baier
Peru
Tingo Maria
Huanuco
8
181
1
1
2561150681
1968-10-11
UFRG
Peru
Pilcopata
Cuzco
8
181
1
1
2561150659
1949-12-12
MZSP
Dirings
Bolivia
400
Chapare
8
181
1
1
Cochabamba