Pheidole oswaldi r. bessonii Forel, 1891: 176
The ant genus Pheidole Westwood, 1839 (Hymenoptera: Formicidae) in Madagascar-taxonomic revision of the bessonii species-group
Salata, Sebastian
Fisher, Brian L.
Zootaxa
2020
2020-08-31
4843
1
1
64
4FY85
Forel, 1891
Forel
1891
[151,509,1050,1077]
Insecta
Formicidae
Pheidole
Animalia
Hymenoptera
16
17
Arthropoda
species
bessonii
Figs 11A–F, 27C, 28C
Pheidole oswaldir. bessonii Forel, 1891: 176, pl. 5, fig. 3 (w.). Forel, 1905: 162(s.).
Status as species: Dalla Torre, 1893: 88. LECTOTYPE[designated here]: 1w., top specimen of the pin, Madagascar, Fianarantsoa, coll. Besson, ANTC3392, CASENT0101838 ( MHNG). PARALECTOTYPES: 1w., bottom specimen, the same pin as lectotype; 3w., the same data as lectotype (CASENT0923201).
Other material. Madagascar. Antananarivo: 11w., Ambohitantely1410, -18.2253 47.28683, 1410 m, Fisher et al. leg. ( CASC). Fianarantsoa: 1w., 9.0 km NE Ivohibe, -22.42667 46.93833, 900 m, Fisher et al. leg. ( CASC); 12w., Belle Vuetrail, Ranomafana National Park, Fianarantsoa Prov., -21.2665 47.42017, 1020 m, R. Harin’Halaleg. ( CASC); 4w., dry wash, 1 kmE of Isalo National Park Interpretive Center, Fianarantsoa Prov., -22.62667 45.35817, 885 m, Irwin et al. leg ( CASC); 5w., Fitovavy Fitovinany Region, District of Ifanadiana Belle vue area 1200 mS of Ranomafana National Parkentrance, -21.2665 47.42017, 1018 m, M. Rin’haleg.( CASC); 9w.,2s., Forêt d’Analalava, 29.6 km280° W Ranohira, -22.59167 45.12833, 700 m, Fisher et al. leg. ( CASC); 4w., JIRAMA water works near river, Ranomafana National Park, Fianarantsoa Prov., -21.2485 47.45217, 690 m, R. Harin’Halaleg. ( CASC); 1w., MampiarikaIV Non Protected Area, 27.98 kmSW Ambositra, -20.73528 47.08382, 1486 m, A. Ravelomananaleg. ( CASC); 8w., 4s., Parc National d’Isalo, 9.1 km354° N Ranohira, -22.48167 45.46167, 725 m, Fisher et al. leg. ( CASC); 1w., Parc National d’Isalo, Ambovo Springs, 29.3 km4° N Ranohira, -22.29833 45.35167, 990 m, Fisher et al. leg. ( CASC); 3w., 1s., Parc National d’Isalo, Sahanafa River, 29.2 km351° N Ranohira, -22.31333 45.29167, 500 m, Fisher et al. leg. ( CASC); 1w., Parc National de Ranomafana, Vatoharanana River, 4.1 km231° SW Ranomafana, -21.29 47.43333, 1100 m, Fisher et al. leg. ( CASC); 4w., R.S. Ivohibe8.0 km E Ivohibe, -22.48333 46.96833, 1200 m, Fisher et al. leg. ( CASC); 2w., R.S. Ivohibe, 7.5 kmENE Ivohibe, -22.47 46.96, 900 m, Fisher et al. leg. ( CASC); 19w., radio tower, Ranomafana National Park, Fianarantsoa Prov., -21.25833 47.40717, 1130 m, Irwin et al. leg. ( CASC); 1s., Ranomafana National Park, Talatakely; Sahambavy; Fianarantsoa Rural, -21.451179 47.3023894, 1139 m, Lee& Ribardoleg. ( CASC); 2w., stream area, 900 mE of Isalo National Park Interpretive Center, Fianarantsoa Prov., -22.62667 45.35817, 750 m, R. Harin’Halaleg. ( CASC); 12w., Vohiparara broken bridge, Fianarantsoa Prov., -21.22617 47.36983, 1110 m, R. Harin’Halaleg. ( CASC); 1w., AmpangabeI Non Protected Area, 21.4 kmW Itremo, -20.61111 46.60688, 1414 m, A. Ravelomananaleg. ( CASC); 1w., AmpangabeII Non Protected Area, 21.29 kmW Itremo, -20.61139 46.60809, 1402 m, A. Ravelomananaleg. ( CASC); 2w., AntapiaIII Non Protected Area, 26.43 kmSW Ambositra, -20.72 47.08785, 1494 m, A. Ravelomananaleg. ( CASC); 1w., AntohatsahombyII Non Protected Area, 23.38 kmNW Itremo, -20.55444 46.58438, 1640 m, A. Ravelomananaleg. ( CASC); 3w., Forêt d’Analalava, 29.6 km280° W Ranohira, -22.59167 45.12833, 700 m, Fisher et al. leg. ( CASC); 1w., MampiarikaII Non Protected Area, 28.08 kmSW Ambositra, -20.73556 47.08384, 1464 m, A. Ravelomananaleg. ( CASC); 3w., MampiarikaIII Non Protected Area, 28.93 kmSW Ambositra, -20.73583 47.08399, 1487 m, A. Ravelomananaleg. ( CASC); 1w., 2s., MampiarikaIV Non Protected Area, 27.98 kmSW Ambositra, - 20.73528 47.08382, 1486 m, A. Ravelomananaleg. ( CASC); 1w., ManandrianaII Non Protected Area, 27.12 kmSW Ambositra, -20.7325 47.09461, 1589 m, A. Ravelomananaleg. ( CASC); 1s., Parc National d’Isalo, 9.1 km354° N Ranohira, -22.48167 45.46167, 725 m, Fisher et al. leg. ( CASC); 1w., Parc National d’Isalo, Sahanafa River, 29.2 km351° N Ranohira, -22.31333 45.29167, 500 m, Fisher et al. leg. ( CASC); 1w., SoanierenanaII Non Protected Area, 25.61 kmSW Ambositra, -20.72194 47.10896, 1732 m, A. Ravelomananaleg. ( CASC). Mahajanga: 9w., 5s., Forêt Ambohimanga, 26.1 km314° Mampikony, -15.96267 47.43817, 250 m, Fisher et al. leg. ( CASC); 1w., Mahavavy River, 6.2 km145° SE Mitsinjo, -16.05167 45.90833, 20 m, Fisher et al. leg. ( CASC); 4w., 8s., 1q., Parc National d’Ankarafantsika, Ampijoroa StationForestière, 5.4 km331° NW Andranofasika, -16.29889 46.813, 70 m, Fisher et al. leg. ( CASC); 2w., Parc National d’Ankarafantsika, Ampijoroa StationForestière, 40 km306° NW Andranofasika, -16.32083 46.81067, 130 m, Fisher et al. leg. ( CASC); 8w., Parc National d’Ankarafantsika, Forêt de Tsimaloto, 18.3 km46° NE de Tsaramandroso, -16.22806, 47.14361, 135 m, Fisher et al. leg. ( CASC); 7w., 5s., Parc National de Baie de Baly, 12.4 km337° NNW Soalala, -16.01 45.265, 10 m, Fisher et al. leg. ( CASC); 12w., 3s., Parc National de Namoroka, 16.9 km317° NW Vilanandro, -16.40667 45.31, 100 m, Fisher et al. leg. ( CASC); 2w., 1s., Parc National de Namoroka, 9.8 km300° WNW Vilanandro, -16.46667 45.35, 140 m, Fisher et al. leg. ( CASC); 1w., Parc National Tsingy de Bemaraha, 10.6 kmESE 123° Antsalova, -18.70944 44.71817, 150 m, Fisher et al. leg. ( CASC); 4w., 1s., Parc National Tsingy de Bemaraha, 2.5 km62° ENE Bekopaka, Ankidrodroa River, -19.13222 44.81467, 100 m, Fisher et al. leg. ( CASC); 6w., 1s., Parc National Tsingy de Bemaraha, 3.4 km93° E Bekopaka, Tombeau Vazimba, -19.14194 44.828, 50 m, Fisher et al. leg. ( CASC); 1w, 1s., Réserve d’Ankoririka, 10.6 km13° NE de Tsaramandroso, -16.26722 47.04861, 210 m, Fisher et al. leg. ( CASC); 2w., 1s., Réserve Spéciale de Bemarivo, 23.8 km223° SW Besalampy, -16.925 44.36833, 30 m, Fisher et al. leg. ( CASC); 1w., Réserveforestière Beanka, 50.2 kmE Maintirano, -18.02649 44.05051, 250 m, Fisher et al. leg. ( CASC). Toamasina: 1w., 1s., 6.3 kmS Ambanizana, Andranobe, -15.6813 49.958, 25 m, Fisher et al. leg. ( CASC). Toliara: 1w., 50 kmN Morondava, - 20.06667, 44.58333, A. Paulyleg. ( CASC); 1w., 1s., Anosy Region, Parc National d’Andohahela, Forêt de Manatalinjo, -24.82505 46.57811, 90 m, Fisher et al. leg. ( CASC); 1w., Antafoky, -23.48778 44.0775, 60 m, Fisher et al. leg. ( CASC); 1w., 1s., Beza-Mahafaly, 27 kmE Betioky, -23.65 44.63333, 135 m, Fisher et al. leg. ( CASC); 1s., Fiherenana, -23.17694 43.96083, 100 m, Fisher et al. leg. ( CASC); 1w., Fiherenana, -23.23528 43.87083, 50 m, Fisher et al. leg. ( CASC); 1w., Forêt Classée d’Analavelona, 29.2 km343° NNW Mahaboboka, -22.675 44.19, 1100 m, Fisher et al. leg. ( CASC); 3w., 3s., Forêt de Beroboka, 5.9 km131° SE Ankidranoka, -22.23306 43.36633, 80 m, Fisher et al. leg. ( CASC); 4w., Forêt de Kirindy, 15.5 km64° ENE Marofandilia, -20.045 44.66222, 100 m, Fisher et al. leg. ( CASC); 7w., Forêt de Kirindy, 15.5 km64° ENE Marofandilia, -20.06855 44.6595667, 30 m, Fisher et al. leg. ( CASC); 4w., Forêt de Mahavelo, Isantoria River, -24.75833 46.15717, 110 m, Fisher et al. leg. ( CASC); 1w., Forêt de Mahavelo, Isantoria River, 5.5 km37° NE Ifotaka, -24.75361 46.1515, 115 m, Fisher et al. leg. ( CASC); 3w., 3s., Forêt de Mite, 20.7 km29° WNW Tongobory, -23.52417 44.12133, 75 m, Fisher et al. leg. ( CASC); 1w., Forêt Ivohibe55.0 km N Tolagnaro, -24.569 47.204, 200 m, Fisher et al. leg. ( CASC); 1w., Forêt Vohidava 88.9 kmN Amboasary, -24.24067 46.28783, 500 m, Fisher et al. leg. ( CASC); 1w., 1s., Forêt Vohidava 89.2 kmN Amboasary, -24.239 46.28233, 850 m, Fisher et al. leg. ( CASC); 2w., Ivahona, -23.45591 46.17376, 820 m, Fisher et al. leg. ( CASC); 1w., Makay Mts., -21.21836 45.3106, 510 m, Fisher et al. leg. ( CASC); 2w., Makay Mts., -21.30997, 45.12946, 590 m, Fisher et al. leg. ( CASC); 3w., Manderano, -23.52722 44.0875, 70 m, Fisher et al. leg. ( CASC); 1w., Manombo, -22.81092 43.7344, 177 m, Fisher et al. leg. ( CASC); 1w., Manombo, -22.8123 43.73932, 165 m, Fisher et al. leg. ( CASC); 1w., Parc National d’Andohahela, Col du Sedro, 3.8 km113° ESE Mahamavo, 37.6 km341° NNW Tolagnaro, -24.76389 46.75167, 900 m, Fisher et al. leg. ( CASC); 6w., Parc National d’Andohahela, Forêt d’Ambohibory, 1.7 km61° ENE Tsimelahy, 36.1 km308° NW Tolagnaro, -24.93 46.6455, 300 m, Fisher et al. leg. ( CASC); 3w., Parc National d’Andohahela, Forêt de Manatalinjo, 33.6 km63° ENE Amboasary, 7.6 km99° E Hazofotsy, -24.81694 46.61, 150 m, Fisher et al. leg. ( CASC); 1w., Parc National de Kirindy Mite, 16.3 km127° SE Belo sur Mer, -20.79528 44.147, 80 m, Fisher et al. leg. ( CASC); 4w., Parc National de Zombitse, 19.8 km84° E Sakaraha, -22.84333 44.71, 770 m, Fisher et al. leg. ( CASC); 11w., 5s., 3m., Réserve Privé Berenty, Forêt de Bealoka, Mandraré River, 14.6 km329° NNW Amboasary, -24.95694 46.2715, 35 m, Fisher et al. leg. ( CASC); 5w., Réserve Spéciale d’Ambohijanahary, Forêt d’Ankazotsihitafototra, 34.6 km314° NW Ambaravaranala, -18.26 45.41833, 1100 m, Fisher et al. leg. ( CASC); 2w., Réserve Spéciale d’Ambohijanahary, Forêt d’Ankazotsihitafototra, 35.2 km312° NW Ambaravaranala, -18.26667 45.40667, 1050 m, Fisher et al. leg. ( CASC); 1w., Sakaraha, -22.91233 44.53283, 470 m, Fisher et al. leg. ( CASC); 1w., Sept Lacs, -23.52833, 44.15556, 80 m, Fisher et al. leg. ( CASC); 1w., southern Isoky-Vohimena Forest, 59 kmNE Sakaraha, -22.46667 44.85, 730 m, Fisher et al. leg. ( CASC); 14w., 9s., Forêt Classée d’Analavelona, 29.2 km343° NNW Mahaboboka, -22.675 44.19, 1100 m, Fisher et al. leg. ( CASC); 1w., Forêt Classée d’Analavelona, 29.4 km343° NNW Mahaboboka, - 22.675 44.18667, 1050 m, Fisher et al. leg. ( CASC); 1s., Forêt Vohidava 89.2 kmN Amboasary, -24.239 46.28233, 850 m, Fisher et al. leg. ( CASC); 2w., Makay Mts., -21.21761, 45.33917, 500 m, Fisher et al. leg. ( CASC); 2w., Makay Mts., -21.22336 45.32628, 480 m, Fisher et al. leg. ( CASC); 4w., Makay Mts., -21.30997 45.12946, 590 m, Fisher et al. leg. ( CASC); 2w., 2m., Makay Mts., -21.31664 45.1296, 620 m, Fisher et al. leg. ( CASC); 3w., 2s., Parc National de Zombitse, 19.8 km84° E Sakaraha, -22.84333 44.71, 770 m, Fisher et al. leg. ( CASC); 1w., Réserve Privé Berenty, Forêt de Malaza, Mandraré River, 8.6 km314° NW Amboasary, -25.00778 46.306, 40 m, Fisher et al. leg. ( CASC). Geographic range. Madagascar, widely distributed on the island except Antsiranana.
Diagnosis. Major workers.Head in full-face view sub-rectangular, not or slightly widening posteriorly, lateral margin slightly convex with sparse, long, decumbent to subdecumbent pilosity; medial frons with moderately thick to thin and dense costulae that are on the posterior part; interspaces between costulae smooth to indistinctly punctate; frons laterally with sparser and thinner rugocostulae; interspaces between rugocostulae distinctly punctate; occipital lobes punctate, sometimes with additional network of sparse and thin rugae, sculpture weakening posteriorly; scape, when laid back, exceeding the midlength of head by two-fifths of its length; inner hypostomal tooth absent or indistinct, bulge-like; outer hypostomal tooth lobe-like, high and wide, directed posteriorly; median tooth absent; promesonotum short, angular, and moderately low; posterior mesonotum moderately steep; mesonotal process distinct and tubercle-like; mesosoma microreticulate; promesonotum with additional thin, sparse, and transverse rugae; gaster shagreened; body brown to blackish brown. Minor workers.Occiput elongated; nuchal collar distinct; head sculpture smooth; anterior frons with sparse puncta; scape, when laid back, exceeding the posterior head margin by two-fifths of its length; promesonotum low, long, and arched; promesonotal groove present; metanotal groove distinct; propodeal spines very small with relatively wide base; promesonotum mostly smooth with sparse microreticulae laterally; mesopleuron and propodeum with sparse microreticulae; body bright brown to brown.
Description. Major workers.Measurements (n=10): HL: 1.78–2.13 (1.97); HW: 1.74–2.01 (1.89); SL: 1.13–1.2 (1.17); EL: 0.22–0.27 (0.25); WL: 1.61–1.78 (1.68); PSL: 0.26–0.34 (0.3); MTL: 1.24–1.38 (1.3); PNW: 0.66–0.75 (0.7); PTW: 0.22–0.31 (0.28); PPW: 0.62–0.81 (0.7); CI: 99.7–106.5 (104.3); SI: 57.8–65.4 (61.7); PSLI: 13.9–16.6 (15.3); PPI: 32.1–47.0 (40.4); PNI: 34.2–39.1 (37.0); MTI: 64.8–74.8 (69.1). Head.In full-face view sub-rectangular, slightly widening posteriorly with sides slightly convex ( Fig. 11B); in lateral view sub-oval, not depressed posteriorly; ventral and dorsal margins convex; inner hypostomal tooth not visible. Sides of head with sparse, long, decumbent to subdecumbent pilosity; head dorsum with relatively dense, long, decumbent to erect pilosity. Medial frons with moderately thick to thin and dense costulae on the posterior part; interspaces between costulae smooth to indistinctly punctate. Frons laterally with sparser and thinner rugocostulae; interspaces between rugocostulae distinctly punctate. Occipital lobes punctate, sometimes with additional network of sparse and thin rugae; sculpture weakening posteriorly. Gena with dense, moderately thick to thin costulae; interspaces between costulae distinctly punctate. Sides posterolateral from eyes densely punctate to microreticulate, sometimes with additional network of thin and sparse rugulae. Center of clypeus shiny and with a few rugulae, lateral sides with distinct rugulae; median notch present, moderately wide, and shallow; median longitudinal carina present; lateral longitudinal carinae present. Scape, when laid back, exceeding the midlength of head by two-fifths of its length; pilosity subdecumbent to erect ( Fig. 11B, D). Inner hypostomal tooth absent or indistinct, bulge-like; outer hypostomal tooth lobe-like, high, and wide,, with top directed posteriorly; median tooth absent ( Fig. 27C). Mesosoma.In lateral view, promesonotum short, angular, and moderately low; posterior mesonotum moderately steep; mesonotal process distinct and tubercle-like; promesonotal groove weakly developed; metanotal groove present; propodeal spines moderately long with wide base and acute top; humeral tubercle weakly produced ( Fig. 11D). Surface shiny and microreticulate; promesonotum with additional thin, sparse, and transverse rugae. Pilosity moderately dense, long, and erect ( Fig. 11D, F). Gaster.Dull and shagreened; pilosity moderately sparse, long, and erect ( Fig. 11D, F). Color.Brown to blackish brown, antennae and legs yellowish brown, mandibles ochraceous ( Fig. 11D, F). FIGURE 11. Pheidole bessonii, full-face view (A), profile (C), and dorsal view (E) of minor worker (CASENT0496974) and full-face view (B), profile (D), and dorsal view (F) of major worker (CASENT0496976). Description. Minor workers.Measurements (n=10): HL: 0.69–0.82 (0.77); HW: 0.52–0.61 (0.57); SL:0.9– 1.09 (1.0); EL: 0.13–0.19 (0.16); WL: 0.96–1.12 (1.04); PSL: 0.07–0.12 (0.1); MTL: 0.78–0.95 (0.88); PNW: 0.33–0.41 (0.37); PTW: 0.09–0.12 (0.1); PPW: 0.17–0.21 (0.18); CI: 131.4–137.6 (133.5); SI: 169.3–183.2 (175.0); PSLI: 10.2–15.3 (13.3); PPI: 48.7–60.0 (55.3); PNI: 61.0–68.7 (65.2); MTI: 144.7–158.1 (152.8). Head.In full-face view oval, posterior region elongated; nuchal collar distinct ( Fig. 11A). Pilosity relatively sparse, moderately long, subdecumbent to erect. Sculpture shiny and smooth; anterolateral frons with sparse puncta. Clypeus with median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, exceeding the posterior head margin by two-fifths of its length; pilosity dense, subdecumbent to erect ( Fig. 11A, C). Mesosoma.In lateral view, promesonotum low, long, and arched; promesonotal groove present; metanotal groove distinct; propodeal spines very small, triangular, and with relatively wide base ( Fig. 11C). Promesonotum mostly smooth with sparse microreticulae on lateral sides; katepisternum, anepisternum, and propodeum with sparse microreticulae. Pilosity very sparse, moderately long, and erect ( Fig. 11C, E). Gaster.With sparse and erect pilosity ( Fig. 11C, E). Color.Bright brown to brown, legs, mandibles, and antennae yellowish ( Fig. 11C, E).
Biology.The species was collected between 30–1732 melevation, in Uapacawoodland, montane rainforest, tropical dry forest, savanna and woodland, gallery forest and rainforest. Nests were located in rotten logs, in the soil, and under stones. Workers were collected from sifted litter.
Comments. Pheidole bessoniiis probably one of the most widespread Pheidolespecies known from Madagascar. Morphologically it is most similar to P. voreiosand P. atsirakambiaty, and its major workers can be separated from both species by punctate occipital lobes (which sometimes have additional thin and very sparse rugulae that arch posterolaterally), and sparsely rugocostulate lateral frons with distinctly punctate interspaces between rugocostulae. In contrast, majors of P. voreiosand P. atsirakambiatyalways have distinctly rugulate to costulate occipital lobes and frons. Minors of P. bessoniican be separated from P. voreiosand P. atsirakambiatyby the less elongated posterior part of the neck, a mostly smooth promesonotum with sparse microreticulae on its sides, and very small propodeal spines. Pheidole voreiosand P. atsirakambiatyhave a more elongated posterior part of the neck, mostly or entirely sculptured promesonotum, and larger propodeal spines. In addition, majors and minors of P. bessoniican be distinguished from P. voreiosby the brown body.
3027954670
MHNG
Besson
Madagascar
Fianarantsoa
16
17
1
Fianarantsoa
lectotype
3027954712
MHNG
Besson
Madagascar
Fianarantsoa
16
17
3
Fianarantsoa
paralectotype
3027954322
CASC
Fisher
Madagascar
1410
-18.2253
Ambohitantely
5
47.28683
16
17
11
Antananarivo
3027954597
[337,1366,1431,1457]
CASC
NE Ivohibe & Fisher
Madagascar
900
-22.42667
Fianarantsoa
1
46.93833
16
17
1
Fianarantsoa
3027954369
Belle Vue
Madagascar
Ranomafana National Park
16
17
12
3027954573
CASC
R. Harin'Hala
Madagascar
1020
-21.2665
Fianarantsoa Prov.
5
47.42017
16
17
1
Fianarantsoa
3027954663
[257,972,1503,1529]
Madagascar
Isalo National Park Interpretive Center
16
17
4
3027954642
CASC
Irwin
Madagascar
885
-22.62667
Fianarantsoa Prov.
1
45.35817
16
17
1
Fianarantsoa
3027954381
CASC
M. Rin'ha
Madagascar
1018
-21.2665
Fitovavy Fitovinany Region
5
47.42017
16
17
5
3027954773
CASC
Fisher
Madagascar
700
-22.59167
Foret d'Analalava
1
45.12833
16
17
9
3027954441
Madagascar
Ranomafana National Park
16
17
4
3027954536
[535,1378,1646,1673]
CASC
R. Harin'Hala
Madagascar
690
-21.2485
Fianarantsoa Prov.
5
47.45217
16
17
1
Fianarantsoa
3027954460
CASC
A. Ravelomanana
Madagascar
1486
-20.73528
Non Protected Area
1
47.08382
Mampiarika
16
17
1
3027954777
CASC
Fisher
Madagascar
725
-22.48167
Parc National d'Isalo
1
45.46167
16
17
8
3027954315
CASC
Fisher
Madagascar
Parc National d'Isalo
990
-22.29833
Ranohira
1
45.35167
Ambovo Springs
16
17
1
3027954708
CASC
Fisher
Madagascar
500
-22.31333
Sahanafa River
1
45.29167
Parc National d'Isalo
16
17
3
3027954511
CASC
Fisher
Madagascar
1100
Vatoharanana River
Parc National de Ranomafana
16
17
1
3027954492
CASC
Fisher
Madagascar
1200
-22.48333
1
46.96833
16
17
4
3027954446
CASC
R. S. Ivohibe & Fisher
Madagascar
900
7.5 km ENE Ivohibe
16
17
2
3027954433
[366,876,1935,1961]
Madagascar
Ranomafana National Park
16
17
19
3027954651
CASC
Irwin
Madagascar
1130
-21.25833
Fianarantsoa Prov.
1
47.40717
16
17
1
Fianarantsoa
3027954512
[439,1070,1971,1997]
Madagascar
Ranomafana National Park
16
17
1
3027954342
CASC
Lee & Ribardo
Madagascar
1139
-21.45118
Rural
1
47.30239
16
17
1
Fianarantsoa
3027954371
Madagascar
900
17
18
Isalo National Park Interpretive Center
16
17
2
3027954434
[243,1138,150,177]
CASC
R. Harin'Hala
Madagascar
750
-22.62667
Fianarantsoa Prov.
1
45.35817
17
18
1
Fianarantsoa
3027954443
[237,1109,187,213]
CASC
R. Harin'Hala
Madagascar
1110
-21.22617
Fianarantsoa Prov.
1
47.36983
17
18
1
Fianarantsoa
3027954785
CASC
A. Ravelomanana
Madagascar
1414
-20.61111
Non Protected Area
1
46.60688
Ampangabe
17
18
1
3027954584
CASC
A. Ravelomanana
Madagascar
1402
-20.61139
Non Protected Area
1
46.60809
Ampangabe
17
18
1
3027954501
CASC
A. Ravelomanana
Madagascar
1494
Non Protected Area
Antapia
17
18
2
3027954769
CASC
A. Ravelomanana
Madagascar
1640
-20.55444
Non Protected Area
1
46.58438
Antohatsahomby
17
18
1
3027954335
CASC
Fisher
Madagascar
700
-22.59167
Foret d'Analalava
1
45.12833
17
18
3
3027954340
CASC
A. Ravelomanana
Madagascar
1464
-20.73556
Non Protected Area
1
47.08384
Mampiarika
17
18
1
3027954747
CASC
A. Ravelomanana
Madagascar
1487
-20.73583
Non Protected Area
1
47.08399
Mampiarika
17
18
3
3027954365
CASC
A. Ravelomanana
Madagascar
1486
-20.73528
Non Protected Area
1
47.08382
Mampiarika
17
18
1
3027954766
CASC
A. Ravelomanana
Madagascar
1589
-20.7325
Non Protected Area
5
47.09461
Manandriana
17
18
1
3027954643
CASC
Fisher
Madagascar
725
-22.48167
Parc National d'Isalo
1
45.46167
17
18
1
3027954632
CASC
Fisher
Madagascar
500
-22.31333
Sahanafa River
1
45.29167
Parc National d'Isalo
17
18
1
3027954504
CASC
A. Ravelomanana
Madagascar
1732
-20.72194
Non Protected Area
1
47.10896
Soanierenana
17
18
1
3027954718
CASC
Fisher
Madagascar
250
-15.96267
Mampikony
1
47.43817
Foret Ambohimanga
17
18
9
Mahajanga
3027954789
CASC
Fisher
Madagascar
20
-16.05167
Mahavavy River
1
45.90833
17
18
1
3027954644
CASC
Fisher
Madagascar
-16.29889
Ampijoroa Station
53
46.813
Parc National d'Ankarafantsika
17
18
5
3027954469
CASC
Fisher
Madagascar
130
-16.32083
Ampijoroa Station
1
46.81067
Parc National d'Ankarafantsika
17
18
2
3027954426
CASC
Fisher
Madagascar
135
-16.22806
Foret de Tsimaloto
1
47.14361
Parc National d'Ankarafantsika
17
18
8
3027954359
CASC
Fisher
Madagascar
Parc National de Baie de Baly
17
18
7
3027954324
CASC
Fisher
Madagascar
100
Parc National de Namoroka
17
18
12
3027954751
CASC
Fisher
Madagascar
140
Parc National de Namoroka
17
18
2
3027954598
CASC
Fisher
Madagascar
150
-18.70944
Antsalova
1
44.71817
Parc National Tsingy de Bemaraha
17
18
1
3027954666
CASC
Fisher
Madagascar
100
-19.13222
Ankidrodroa River
1
44.81467
Parc National Tsingy de Bemaraha
17
18
4
3027954455
CASC
Fisher
Madagascar
-19.14194
Tombeau Vazimba
52
44.828
Parc National Tsingy de Bemaraha
17
18
6
3027954507
CASC
Fisher
Madagascar
210
-16.26722
Reserve d'Ankoririka
1
47.04861
17
18
1
3027954544
CASC
Fisher
Madagascar
30
-16.925
Reserve Speciale de Bemarivo
55
44.36833
17
18
2
3027954605
CASC
Fisher
Madagascar
250
-18.02649
Beanka
1
44.05051
Reserve
17
18
1
3027954302
CASC
Fisher
Madagascar
-15.6813
Andranobe
53
49.958
17
18
1
Toamasina
3027954429
CASC
A. Pauly
Madagascar
-20.06667
50 km N Morondava
1
44.58333
17
18
1
Toliara
3027954382
CASC
Fisher
Madagascar
Anosy Region
90
-24.82505
Foret de Manatalinjo
1
46.57811
Parc National d'Andohahela
17
18
1
3027954326
CASC
Fisher
Madagascar
60
-23.48778
Antafoky
5
44.0775
17
18
1
3027954612
[309,1384,1374,1401]
CASC
Fisher
Madagascar
135
Beza-Mahafaly
17
18
1
3027954678
CASC
Fisher
Madagascar
100
-23.17694
Fiherenana
1
43.96083
17
18
1
3027954748
CASC
Fisher
Madagascar
50
-23.23528
Fiherenana
1
43.87083
17
18
1
3027954477
CASC
Fisher
Madagascar
1100
Foret Classee d'Analavelona
17
18
1
3027954706
CASC
Fisher
Madagascar
80
-22.23306
Foret de Beroboka
1
43.36633
17
18
3
3027954427
CASC
Fisher
Madagascar
100
-20.045
Foret de Kirindy
55
44.66222
17
18
4
3027954571
CASC
Fisher
Madagascar
30
-20.06855
Foret de Kirindy
1
44.659565
17
18
7
3027954585
[346,1432,1590,1617]
CASC
Fisher
Madagascar
110
-24.75833
Isantoria River
1
46.15717
Foret de Mahavelo
17
18
4
3027954410
CASC
Fisher
Madagascar
115
-24.75361
Isantoria River
5
46.1515
Foret de Mahavelo
17
18
1
3027954715
CASC
Fisher
Madagascar
75
-23.52417
Foret de Mite
1
44.12133
17
18
3
3027954579
[260,1308,1698,1725]
CASC
N Tolagnaro & Fisher
Madagascar
200
-24.569
Ivohibe
74
47.204
17
18
1
3027954526
CASC
Fisher
Madagascar
500
-24.24067
Foret Vohidava
1
46.28783
17
18
1
3027954560
CASC
Fisher
Madagascar
850
-24.239
Foret Vohidava
55
46.28233
17
18
1
3027954368
CASC
Fisher
Madagascar
820
-23.45591
Ivahona
1
46.17376
17
18
2
3027954545
[486,1292,1806,1833]
CASC
Fisher
Madagascar
510
-21.21836
Makay Mts.
5
45.3106
17
18
1
3027954521
CASC
Fisher
Madagascar
590
-21.30997
Makay Mts.
1
45.12946
17
18
2
3027954725
CASC
Fisher
Madagascar
70
-23.52722
Manderano
5
44.0875
17
18
3
3027954583
[346,1138,1879,1905]
CASC
Fisher
Madagascar
177
-22.81092
Manombo
5
43.7344
17
18
1
3027954590
CASC
Fisher
Madagascar
165
-22.8123
Manombo
5
43.73932
17
18
1
3027954464
CASC
Col du Sedro & Fisher
Madagascar
900
-24.76389
Parc National d'Andohahela
1
46.75167
17
18
1
3027954378
CASC
Fisher
Madagascar
300
18
19
Foret d'Ambohibory
Parc National d'Andohahela
17
18
6
3027954450
CASC
Fisher
Madagascar
Parc National d'Andohahela
150
Hazofotsy
99.0
Foret de Manatalinjo
18
19
3
3027954396
CASC
Fisher
Madagascar
-20.79528
Parc National de Kirindy Mite
51
44.147
18
19
1
3027954755
CASC
Fisher
Madagascar
770
Sakaraha
84.0
Parc National de Zombitse
18
19
4
3027954438
CASC
Fisher
Madagascar
Reserve Prive Berenty
35
-24.95694
Mandrare River
5
46.2715
Foret de Bealoka
18
19
1
3027954437
CASC
Fisher
Madagascar
1100
Foret d'Ankazotsihitafototra
Reserve Speciale d'Ambohijanahary
18
19
5
3027954418
CASC
Fisher
Madagascar
1050
-18.26667
Foret d'Ankazotsihitafototra
1
45.40667
Reserve Speciale d'Ambohijanahary
18
19
2
3027954582
[256,1016,438,465]
CASC
Fisher
Madagascar
470
-22.91233
Sakaraha
1
44.53283
18
19
1
3027954379
CASC
Fisher
Madagascar
80
-23.52833
Sept Lacs
1
44.15556
18
19
1
3027954422
CASC
Fisher
Madagascar
730
southern Isoky-Vohimena Forest
18
19
1
3027954454
CASC
Fisher
Madagascar
1100
Foret Classee d'Analavelona
18
19
14
3027954470
CASC
Fisher
Madagascar
1050
-22.675
Foret Classee d'Analavelona
55
44.18667
18
19
1
3027954519
CASC
Fisher
Madagascar
850
-24.239
Foret Vohidava
55
46.28233
18
19
1
3027954639
[542,1377,618,645]
CASC
Fisher
Madagascar
500
-21.21761
Makay Mts.
1
45.33917
18
19
2
3027954587
CASC
Fisher
Madagascar
480
-21.22336
Makay Mts.
1
45.32628
18
19
2
3027954494
CASC
Fisher
Madagascar
590
-21.30997
Makay Mts.
1
45.12946
18
19
4
3027954646
[440,1279,690,717]
CASC
Fisher
Madagascar
2
-21.31664
Makay Mts.
5
45.1296
18
19
4
3027954316
CASC
Fisher
Madagascar
770
Sakaraha
84.0
Parc National de Zombitse
18
19
3
3027954770
CASC
Fisher
Madagascar
Reserve Prive Berenty
-25.00778
Mandrare River
50
46.306
Foret de Malaza
18
19
1