Pheidole oswaldi r. bessonii Forel, 1891: 176 The ant genus Pheidole Westwood, 1839 (Hymenoptera: Formicidae) in Madagascar-taxonomic revision of the bessonii species-group Salata, Sebastian Fisher, Brian L. Zootaxa 2020 2020-08-31 4843 1 1 64 4FY85 Forel, 1891 Forel 1891 [151,509,1050,1077] Insecta Formicidae Pheidole Animalia Hymenoptera 16 17 Arthropoda species bessonii   Figs 11A–F, 27C, 28C      Pheidole oswaldir. bessonii Forel, 1891: 176, pl. 5, fig. 3 (w.).  Forel, 1905: 162(s.).  Status as species: Dalla Torre, 1893: 88.   LECTOTYPE[designated here]: 1w., top specimen of the pin, Madagascar, Fianarantsoa, coll. Besson, ANTC3392, CASENT0101838 ( MHNG). PARALECTOTYPES: 1w., bottom specimen, the same pin as lectotype;  3w., the same data as lectotype (CASENT0923201).   Other material.  Madagascar. Antananarivo: 11w., Ambohitantely1410, -18.2253 47.28683,  1410 m, Fisher et al. leg. ( CASC).  Fianarantsoa: 1w., 9.0 km NE Ivohibe, -22.42667 46.93833,  900 m, Fisher et al. leg. ( CASC);  12w., Belle Vuetrail, Ranomafana National Park,  Fianarantsoa Prov., -21.2665 47.42017,  1020 m, R. Harin’Halaleg. ( CASC);  4w., dry wash, 1 kmE of Isalo National Park Interpretive Center,  Fianarantsoa Prov., -22.62667 45.35817,  885 m, Irwin et al. leg ( CASC);  5w., Fitovavy Fitovinany Region, District of Ifanadiana Belle vue area  1200 mS of Ranomafana National Parkentrance, -21.2665 47.42017,  1018 m, M. Rin’haleg.( CASC);  9w.,2s., Forêt d’Analalava,  29.6 km280° W Ranohira, -22.59167 45.12833,  700 m, Fisher et al. leg. ( CASC);  4w., JIRAMA water works near river, Ranomafana National Park,  Fianarantsoa Prov., -21.2485 47.45217,  690 m, R. Harin’Halaleg. ( CASC);  1w., MampiarikaIV Non Protected Area,  27.98 kmSW Ambositra, -20.73528 47.08382,  1486 m, A. Ravelomananaleg. ( CASC);  8w., 4s., Parc National d’Isalo,  9.1 km354° N Ranohira, -22.48167 45.46167,  725 m, Fisher et al. leg. ( CASC);  1w., Parc National d’Isalo, Ambovo Springs, 29.3 km4° N Ranohira, -22.29833 45.35167,  990 m, Fisher et al. leg. ( CASC);  3w., 1s., Parc National d’Isalo, Sahanafa River,  29.2 km351° N Ranohira, -22.31333 45.29167,  500 m, Fisher et al. leg. ( CASC);  1w., Parc National de Ranomafana, Vatoharanana River,  4.1 km231° SW Ranomafana, -21.29 47.43333,  1100 m, Fisher et al. leg. ( CASC);  4w., R.S. Ivohibe8.0 km E Ivohibe, -22.48333 46.96833,  1200 m, Fisher et al. leg. ( CASC);  2w., R.S. Ivohibe,  7.5 kmENE Ivohibe, -22.47 46.96,  900 m, Fisher et al. leg. ( CASC);  19w., radio tower, Ranomafana National Park,  Fianarantsoa Prov., -21.25833 47.40717,  1130 m, Irwin et al. leg. ( CASC); 1s., Ranomafana National Park, Talatakely; Sahambavy;  Fianarantsoa Rural, -21.451179 47.3023894,  1139 m, Lee& Ribardoleg. ( CASC);  2w., stream area,  900 mE of Isalo National Park Interpretive Center,  Fianarantsoa Prov., -22.62667 45.35817,  750 m, R. Harin’Halaleg. ( CASC); 12w., Vohiparara broken bridge,  Fianarantsoa Prov., -21.22617 47.36983,  1110 m, R. Harin’Halaleg. ( CASC);  1w., AmpangabeI Non Protected Area,  21.4 kmW Itremo, -20.61111 46.60688,  1414 m, A. Ravelomananaleg. ( CASC);  1w., AmpangabeII Non Protected Area,  21.29 kmW Itremo, -20.61139 46.60809,  1402 m, A. Ravelomananaleg. ( CASC);  2w., AntapiaIII Non Protected Area,  26.43 kmSW Ambositra, -20.72 47.08785,  1494 m, A. Ravelomananaleg. ( CASC);  1w., AntohatsahombyII Non Protected Area,  23.38 kmNW Itremo, -20.55444 46.58438,  1640 m, A. Ravelomananaleg. ( CASC);  3w., Forêt d’Analalava,  29.6 km280° W Ranohira, -22.59167 45.12833,  700 m, Fisher et al. leg. ( CASC);  1w., MampiarikaII Non Protected Area,  28.08 kmSW Ambositra, -20.73556 47.08384,  1464 m, A. Ravelomananaleg. ( CASC);  3w., MampiarikaIII Non Protected Area,  28.93 kmSW Ambositra, -20.73583 47.08399,  1487 m, A. Ravelomananaleg. ( CASC);  1w., 2s., MampiarikaIV Non Protected Area,  27.98 kmSW Ambositra, - 20.73528 47.08382,  1486 m, A. Ravelomananaleg. ( CASC);  1w., ManandrianaII Non Protected Area,  27.12 kmSW Ambositra, -20.7325 47.09461,  1589 m, A. Ravelomananaleg. ( CASC); 1s., Parc National d’Isalo,  9.1 km354° N Ranohira, -22.48167 45.46167,  725 m, Fisher et al. leg. ( CASC);  1w., Parc National d’Isalo, Sahanafa River,  29.2 km351° N Ranohira, -22.31333 45.29167,  500 m, Fisher et al. leg. ( CASC);  1w., SoanierenanaII Non Protected Area,  25.61 kmSW Ambositra, -20.72194 47.10896,  1732 m, A. Ravelomananaleg. ( CASC).  Mahajanga: 9w., 5s., Forêt Ambohimanga, 26.1 km314° Mampikony, -15.96267 47.43817,  250 m, Fisher et al. leg. ( CASC);  1w., Mahavavy River,  6.2 km145° SE Mitsinjo, -16.05167 45.90833,  20 m, Fisher et al. leg. ( CASC);  4w., 8s., 1q., Parc National d’Ankarafantsika, Ampijoroa StationForestière,  5.4 km331° NW Andranofasika, -16.29889 46.813, 70 m, Fisher et al. leg. ( CASC);  2w., Parc National d’Ankarafantsika, Ampijoroa StationForestière,  40 km306° NW Andranofasika, -16.32083 46.81067,  130 m, Fisher et al. leg. ( CASC);  8w., Parc National d’Ankarafantsika, Forêt de Tsimaloto,  18.3 km46° NE de Tsaramandroso, -16.22806, 47.14361,  135 m, Fisher et al. leg. ( CASC);  7w., 5s., Parc National de Baie de Baly,  12.4 km337° NNW Soalala, -16.01 45.265, 10 m, Fisher et al. leg. ( CASC);  12w., 3s., Parc National de Namoroka,  16.9 km317° NW Vilanandro, -16.40667 45.31,  100 m, Fisher et al. leg. ( CASC);  2w., 1s., Parc National de Namoroka,  9.8 km300° WNW Vilanandro, -16.46667 45.35,  140 m, Fisher et al. leg. ( CASC);  1w., Parc National Tsingy de Bemaraha, 10.6 kmESE 123° Antsalova, -18.70944 44.71817,  150 m, Fisher et al. leg. ( CASC);  4w., 1s., Parc National Tsingy de Bemaraha,  2.5 km62° ENE Bekopaka, Ankidrodroa River, -19.13222 44.81467,  100 m, Fisher et al. leg. ( CASC);  6w., 1s., Parc National Tsingy de Bemaraha,  3.4 km93° E Bekopaka, Tombeau Vazimba, -19.14194 44.828, 50 m, Fisher et al. leg. ( CASC); 1w, 1s., Réserve d’Ankoririka,  10.6 km13° NE de Tsaramandroso, -16.26722 47.04861,  210 m, Fisher et al. leg. ( CASC);  2w., 1s., Réserve Spéciale de Bemarivo,  23.8 km223° SW Besalampy, -16.925 44.36833,  30 m, Fisher et al. leg. ( CASC);  1w., Réserveforestière Beanka,  50.2 kmE Maintirano, -18.02649 44.05051,  250 m, Fisher et al. leg. ( CASC).  Toamasina: 1w., 1s.,  6.3 kmS Ambanizana, Andranobe, -15.6813 49.958, 25 m, Fisher et al. leg. ( CASC).  Toliara: 1w.,  50 kmN Morondava, - 20.06667, 44.58333, A. Paulyleg. ( CASC);  1w., 1s., Anosy Region, Parc National d’Andohahela, Forêt de Manatalinjo, -24.82505 46.57811,  90 m, Fisher et al. leg. ( CASC);  1w., Antafoky, -23.48778 44.0775,  60 m, Fisher et al. leg. ( CASC);  1w., 1s., Beza-Mahafaly,  27 kmE Betioky, -23.65 44.63333,  135 m, Fisher et al. leg. ( CASC); 1s., Fiherenana, -23.17694 43.96083,  100 m, Fisher et al. leg. ( CASC);  1w., Fiherenana, -23.23528 43.87083,  50 m, Fisher et al. leg. ( CASC);  1w., Forêt Classée d’Analavelona,  29.2 km343° NNW Mahaboboka, -22.675 44.19,  1100 m, Fisher et al. leg. ( CASC);  3w., 3s., Forêt de Beroboka,  5.9 km131° SE Ankidranoka, -22.23306 43.36633,  80 m, Fisher et al. leg. ( CASC);  4w., Forêt de Kirindy,  15.5 km64° ENE Marofandilia, -20.045 44.66222,  100 m, Fisher et al. leg. ( CASC);  7w., Forêt de Kirindy,  15.5 km64° ENE Marofandilia, -20.06855 44.6595667,  30 m, Fisher et al. leg. ( CASC);  4w., Forêt de Mahavelo, Isantoria River, -24.75833 46.15717,  110 m, Fisher et al. leg. ( CASC);  1w., Forêt de Mahavelo, Isantoria River,  5.5 km37° NE Ifotaka, -24.75361 46.1515,  115 m, Fisher et al. leg. ( CASC);  3w., 3s., Forêt de Mite,  20.7 km29° WNW Tongobory, -23.52417 44.12133,  75 m, Fisher et al. leg. ( CASC);  1w., Forêt Ivohibe55.0 km N Tolagnaro, -24.569 47.204,  200 m, Fisher et al. leg. ( CASC);  1w., Forêt Vohidava  88.9 kmN Amboasary, -24.24067 46.28783,  500 m, Fisher et al. leg. ( CASC);  1w., 1s., Forêt Vohidava  89.2 kmN Amboasary, -24.239 46.28233,  850 m, Fisher et al. leg. ( CASC);  2w., Ivahona, -23.45591 46.17376,  820 m, Fisher et al. leg. ( CASC);  1w., Makay Mts., -21.21836 45.3106,  510 m, Fisher et al. leg. ( CASC);  2w., Makay Mts., -21.30997, 45.12946,  590 m, Fisher et al. leg. ( CASC);  3w., Manderano, -23.52722 44.0875,  70 m, Fisher et al. leg. ( CASC);  1w., Manombo, -22.81092 43.7344,  177 m, Fisher et al. leg. ( CASC);  1w., Manombo, -22.8123 43.73932,  165 m, Fisher et al. leg. ( CASC);  1w., Parc National d’Andohahela, Col du Sedro,  3.8 km113° ESE Mahamavo,  37.6 km341° NNW Tolagnaro, -24.76389 46.75167,  900 m, Fisher et al. leg. ( CASC);  6w., Parc National d’Andohahela, Forêt d’Ambohibory,  1.7 km61° ENE Tsimelahy,  36.1 km308° NW Tolagnaro, -24.93 46.6455,  300 m, Fisher et al. leg. ( CASC);  3w., Parc National d’Andohahela, Forêt de Manatalinjo,  33.6 km63° ENE Amboasary, 7.6 km99° E Hazofotsy, -24.81694 46.61,  150 m, Fisher et al. leg. ( CASC);  1w., Parc National de Kirindy Mite,  16.3 km127° SE Belo sur Mer, -20.79528 44.147, 80 m, Fisher et al. leg. ( CASC);  4w., Parc National de Zombitse, 19.8 km84° E Sakaraha, -22.84333 44.71,  770 m, Fisher et al. leg. ( CASC);  11w., 5s.,  3m., Réserve Privé Berenty, Forêt de Bealoka, Mandraré River,  14.6 km329° NNW Amboasary, -24.95694 46.2715,  35 m, Fisher et al. leg. ( CASC);  5w., Réserve Spéciale d’Ambohijanahary, Forêt d’Ankazotsihitafototra,  34.6 km314° NW Ambaravaranala, -18.26 45.41833,  1100 m, Fisher et al. leg. ( CASC);  2w., Réserve Spéciale d’Ambohijanahary, Forêt d’Ankazotsihitafototra,  35.2 km312° NW Ambaravaranala, -18.26667 45.40667,  1050 m, Fisher et al. leg. ( CASC);  1w., Sakaraha, -22.91233 44.53283,  470 m, Fisher et al. leg. ( CASC);  1w., Sept Lacs, -23.52833, 44.15556,  80 m, Fisher et al. leg. ( CASC);  1w., southern Isoky-Vohimena Forest,  59 kmNE Sakaraha, -22.46667 44.85,  730 m, Fisher et al. leg. ( CASC);  14w., 9s., Forêt Classée d’Analavelona,  29.2 km343° NNW Mahaboboka, -22.675 44.19,  1100 m, Fisher et al. leg. ( CASC);  1w., Forêt Classée d’Analavelona,  29.4 km343° NNW Mahaboboka, - 22.675 44.18667,  1050 m, Fisher et al. leg. ( CASC); 1s., Forêt Vohidava  89.2 kmN Amboasary, -24.239 46.28233,  850 m, Fisher et al. leg. ( CASC);  2w., Makay Mts., -21.21761, 45.33917,  500 m, Fisher et al. leg. ( CASC);  2w., Makay Mts., -21.22336 45.32628,  480 m, Fisher et al. leg. ( CASC);  4w., Makay Mts., -21.30997 45.12946,  590 m, Fisher et al. leg. ( CASC);  2w.,   2m., Makay Mts., -21.31664 45.1296,  620 m, Fisher et al. leg. ( CASC);  3w., 2s., Parc National de Zombitse, 19.8 km84° E Sakaraha, -22.84333 44.71,  770 m, Fisher et al. leg. ( CASC);  1w., Réserve Privé Berenty, Forêt de Malaza, Mandraré River,  8.6 km314° NW Amboasary, -25.00778 46.306, 40 m, Fisher et al. leg. ( CASC).  Geographic range. Madagascar, widely distributed on the island except Antsiranana.   Diagnosis. Major workers.Head in full-face view sub-rectangular, not or slightly widening posteriorly, lateral margin slightly convex with sparse, long, decumbent to subdecumbent pilosity; medial frons with moderately thick to thin and dense costulae that are on the posterior part; interspaces between costulae smooth to indistinctly punctate; frons laterally with sparser and thinner rugocostulae; interspaces between rugocostulae distinctly punctate; occipital lobes punctate, sometimes with additional network of sparse and thin rugae, sculpture weakening posteriorly; scape, when laid back, exceeding the midlength of head by two-fifths of its length; inner hypostomal tooth absent or indistinct, bulge-like; outer hypostomal tooth lobe-like, high and wide, directed posteriorly; median tooth absent; promesonotum short, angular, and moderately low; posterior mesonotum moderately steep; mesonotal process distinct and tubercle-like; mesosoma microreticulate; promesonotum with additional thin, sparse, and transverse rugae; gaster shagreened; body brown to blackish brown. Minor workers.Occiput elongated; nuchal collar distinct; head sculpture smooth; anterior frons with sparse puncta; scape, when laid back, exceeding the posterior head margin by two-fifths of its length; promesonotum low, long, and arched; promesonotal groove present; metanotal groove distinct; propodeal spines very small with relatively wide base; promesonotum mostly smooth with sparse microreticulae laterally; mesopleuron and propodeum with sparse microreticulae; body bright brown to brown.   Description. Major workers.Measurements (n=10): HL: 1.78–2.13 (1.97); HW: 1.74–2.01 (1.89); SL: 1.13–1.2 (1.17); EL: 0.22–0.27 (0.25); WL: 1.61–1.78 (1.68); PSL: 0.26–0.34 (0.3); MTL: 1.24–1.38 (1.3); PNW: 0.66–0.75 (0.7); PTW: 0.22–0.31 (0.28); PPW: 0.62–0.81 (0.7); CI: 99.7–106.5 (104.3); SI: 57.8–65.4 (61.7); PSLI: 13.9–16.6 (15.3); PPI: 32.1–47.0 (40.4); PNI: 34.2–39.1 (37.0); MTI: 64.8–74.8 (69.1).  Head.In full-face view sub-rectangular, slightly widening posteriorly with sides slightly convex ( Fig. 11B); in lateral view sub-oval, not depressed posteriorly; ventral and dorsal margins convex; inner hypostomal tooth not visible. Sides of head with sparse, long, decumbent to subdecumbent pilosity; head dorsum with relatively dense, long, decumbent to erect pilosity. Medial frons with moderately thick to thin and dense costulae on the posterior part; interspaces between costulae smooth to indistinctly punctate. Frons laterally with sparser and thinner rugocostulae; interspaces between rugocostulae distinctly punctate. Occipital lobes punctate, sometimes with additional network of sparse and thin rugae; sculpture weakening posteriorly. Gena with dense, moderately thick to thin costulae; interspaces between costulae distinctly punctate. Sides posterolateral from eyes densely punctate to microreticulate, sometimes with additional network of thin and sparse rugulae. Center of clypeus shiny and with a few rugulae, lateral sides with distinct rugulae; median notch present, moderately wide, and shallow; median longitudinal carina present; lateral longitudinal carinae present. Scape, when laid back, exceeding the midlength of head by two-fifths of its length; pilosity subdecumbent to erect ( Fig. 11B, D). Inner hypostomal tooth absent or indistinct, bulge-like; outer hypostomal tooth lobe-like, high, and wide,, with top directed posteriorly; median tooth absent ( Fig. 27C).  Mesosoma.In lateral view, promesonotum short, angular, and moderately low; posterior mesonotum moderately steep; mesonotal process distinct and tubercle-like; promesonotal groove weakly developed; metanotal groove present; propodeal spines moderately long with wide base and acute top; humeral tubercle weakly produced ( Fig. 11D). Surface shiny and microreticulate; promesonotum with additional thin, sparse, and transverse rugae. Pilosity moderately dense, long, and erect ( Fig. 11D, F). Gaster.Dull and shagreened; pilosity moderately sparse, long, and erect ( Fig. 11D, F). Color.Brown to blackish brown, antennae and legs yellowish brown, mandibles ochraceous ( Fig. 11D, F).   FIGURE 11.  Pheidole bessonii, full-face view (A), profile (C), and dorsal view (E) of minor worker (CASENT0496974) and full-face view (B), profile (D), and dorsal view (F) of major worker (CASENT0496976).  Description. Minor workers.Measurements (n=10): HL: 0.69–0.82 (0.77); HW: 0.52–0.61 (0.57); SL:0.9– 1.09 (1.0); EL: 0.13–0.19 (0.16); WL: 0.96–1.12 (1.04); PSL: 0.07–0.12 (0.1); MTL: 0.78–0.95 (0.88); PNW: 0.33–0.41 (0.37); PTW: 0.09–0.12 (0.1); PPW: 0.17–0.21 (0.18); CI: 131.4–137.6 (133.5); SI: 169.3–183.2 (175.0); PSLI: 10.2–15.3 (13.3); PPI: 48.7–60.0 (55.3); PNI: 61.0–68.7 (65.2); MTI: 144.7–158.1 (152.8).  Head.In full-face view oval, posterior region elongated; nuchal collar distinct ( Fig. 11A). Pilosity relatively sparse, moderately long, subdecumbent to erect. Sculpture shiny and smooth; anterolateral frons with sparse puncta. Clypeus with median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, exceeding the posterior head margin by two-fifths of its length; pilosity dense, subdecumbent to erect ( Fig. 11A, C).  Mesosoma.In lateral view, promesonotum low, long, and arched; promesonotal groove present; metanotal groove distinct; propodeal spines very small, triangular, and with relatively wide base ( Fig. 11C). Promesonotum mostly smooth with sparse microreticulae on lateral sides; katepisternum, anepisternum, and propodeum with sparse microreticulae. Pilosity very sparse, moderately long, and erect ( Fig. 11C, E). Gaster.With sparse and erect pilosity ( Fig. 11C, E). Color.Bright brown to brown, legs, mandibles, and antennae yellowish ( Fig. 11C, E).   Biology.The species was collected between 30–1732 melevation, in  Uapacawoodland, montane rainforest, tropical dry forest, savanna and woodland, gallery forest and rainforest. Nests were located in rotten logs, in the soil, and under stones. Workers were collected from sifted litter.   Comments.  Pheidole bessoniiis probably one of the most widespread  Pheidolespecies known from Madagascar. Morphologically it is most similar to  P. voreiosand  P. atsirakambiaty, and its major workers can be separated from both species by punctate occipital lobes (which sometimes have additional thin and very sparse rugulae that arch posterolaterally), and sparsely rugocostulate lateral frons with distinctly punctate interspaces between rugocostulae. In contrast, majors of  P. voreiosand  P. atsirakambiatyalways have distinctly rugulate to costulate occipital lobes and frons. Minors of  P. bessoniican be separated from  P. voreiosand  P. atsirakambiatyby the less elongated posterior part of the neck, a mostly smooth promesonotum with sparse microreticulae on its sides, and very small propodeal spines.  Pheidole voreiosand  P. atsirakambiatyhave a more elongated posterior part of the neck, mostly or entirely sculptured promesonotum, and larger propodeal spines. In addition, majors and minors of  P. bessoniican be distinguished from  P. voreiosby the brown body. 3027954670 MHNG Besson Madagascar Fianarantsoa 16 17 1 Fianarantsoa lectotype 3027954712 MHNG Besson Madagascar Fianarantsoa 16 17 3 Fianarantsoa paralectotype 3027954322 CASC Fisher Madagascar 1410 -18.2253 Ambohitantely 5 47.28683 16 17 11 Antananarivo 3027954597 [337,1366,1431,1457] CASC NE Ivohibe & Fisher Madagascar 900 -22.42667 Fianarantsoa 1 46.93833 16 17 1 Fianarantsoa 3027954369 Belle Vue Madagascar Ranomafana National Park 16 17 12 3027954573 CASC R. Harin'Hala Madagascar 1020 -21.2665 Fianarantsoa Prov. 5 47.42017 16 17 1 Fianarantsoa 3027954663 [257,972,1503,1529] Madagascar Isalo National Park Interpretive Center 16 17 4 3027954642 CASC Irwin Madagascar 885 -22.62667 Fianarantsoa Prov. 1 45.35817 16 17 1 Fianarantsoa 3027954381 CASC M. Rin'ha Madagascar 1018 -21.2665 Fitovavy Fitovinany Region 5 47.42017 16 17 5 3027954773 CASC Fisher Madagascar 700 -22.59167 Foret d'Analalava 1 45.12833 16 17 9 3027954441 Madagascar Ranomafana National Park 16 17 4 3027954536 [535,1378,1646,1673] CASC R. Harin'Hala Madagascar 690 -21.2485 Fianarantsoa Prov. 5 47.45217 16 17 1 Fianarantsoa 3027954460 CASC A. Ravelomanana Madagascar 1486 -20.73528 Non Protected Area 1 47.08382 Mampiarika 16 17 1 3027954777 CASC Fisher Madagascar 725 -22.48167 Parc National d'Isalo 1 45.46167 16 17 8 3027954315 CASC Fisher Madagascar Parc National d'Isalo 990 -22.29833 Ranohira 1 45.35167 Ambovo Springs 16 17 1 3027954708 CASC Fisher Madagascar 500 -22.31333 Sahanafa River 1 45.29167 Parc National d'Isalo 16 17 3 3027954511 CASC Fisher Madagascar 1100 Vatoharanana River Parc National de Ranomafana 16 17 1 3027954492 CASC Fisher Madagascar 1200 -22.48333 1 46.96833 16 17 4 3027954446 CASC R. S. Ivohibe & Fisher Madagascar 900 7.5 km ENE Ivohibe 16 17 2 3027954433 [366,876,1935,1961] Madagascar Ranomafana National Park 16 17 19 3027954651 CASC Irwin Madagascar 1130 -21.25833 Fianarantsoa Prov. 1 47.40717 16 17 1 Fianarantsoa 3027954512 [439,1070,1971,1997] Madagascar Ranomafana National Park 16 17 1 3027954342 CASC Lee & Ribardo Madagascar 1139 -21.45118 Rural 1 47.30239 16 17 1 Fianarantsoa 3027954371 Madagascar 900 17 18 Isalo National Park Interpretive Center 16 17 2 3027954434 [243,1138,150,177] CASC R. Harin'Hala Madagascar 750 -22.62667 Fianarantsoa Prov. 1 45.35817 17 18 1 Fianarantsoa 3027954443 [237,1109,187,213] CASC R. Harin'Hala Madagascar 1110 -21.22617 Fianarantsoa Prov. 1 47.36983 17 18 1 Fianarantsoa 3027954785 CASC A. Ravelomanana Madagascar 1414 -20.61111 Non Protected Area 1 46.60688 Ampangabe 17 18 1 3027954584 CASC A. Ravelomanana Madagascar 1402 -20.61139 Non Protected Area 1 46.60809 Ampangabe 17 18 1 3027954501 CASC A. Ravelomanana Madagascar 1494 Non Protected Area Antapia 17 18 2 3027954769 CASC A. Ravelomanana Madagascar 1640 -20.55444 Non Protected Area 1 46.58438 Antohatsahomby 17 18 1 3027954335 CASC Fisher Madagascar 700 -22.59167 Foret d'Analalava 1 45.12833 17 18 3 3027954340 CASC A. Ravelomanana Madagascar 1464 -20.73556 Non Protected Area 1 47.08384 Mampiarika 17 18 1 3027954747 CASC A. Ravelomanana Madagascar 1487 -20.73583 Non Protected Area 1 47.08399 Mampiarika 17 18 3 3027954365 CASC A. Ravelomanana Madagascar 1486 -20.73528 Non Protected Area 1 47.08382 Mampiarika 17 18 1 3027954766 CASC A. Ravelomanana Madagascar 1589 -20.7325 Non Protected Area 5 47.09461 Manandriana 17 18 1 3027954643 CASC Fisher Madagascar 725 -22.48167 Parc National d'Isalo 1 45.46167 17 18 1 3027954632 CASC Fisher Madagascar 500 -22.31333 Sahanafa River 1 45.29167 Parc National d'Isalo 17 18 1 3027954504 CASC A. Ravelomanana Madagascar 1732 -20.72194 Non Protected Area 1 47.10896 Soanierenana 17 18 1 3027954718 CASC Fisher Madagascar 250 -15.96267 Mampikony 1 47.43817 Foret Ambohimanga 17 18 9 Mahajanga 3027954789 CASC Fisher Madagascar 20 -16.05167 Mahavavy River 1 45.90833 17 18 1 3027954644 CASC Fisher Madagascar -16.29889 Ampijoroa Station 53 46.813 Parc National d'Ankarafantsika 17 18 5 3027954469 CASC Fisher Madagascar 130 -16.32083 Ampijoroa Station 1 46.81067 Parc National d'Ankarafantsika 17 18 2 3027954426 CASC Fisher Madagascar 135 -16.22806 Foret de Tsimaloto 1 47.14361 Parc National d'Ankarafantsika 17 18 8 3027954359 CASC Fisher Madagascar Parc National de Baie de Baly 17 18 7 3027954324 CASC Fisher Madagascar 100 Parc National de Namoroka 17 18 12 3027954751 CASC Fisher Madagascar 140 Parc National de Namoroka 17 18 2 3027954598 CASC Fisher Madagascar 150 -18.70944 Antsalova 1 44.71817 Parc National Tsingy de Bemaraha 17 18 1 3027954666 CASC Fisher Madagascar 100 -19.13222 Ankidrodroa River 1 44.81467 Parc National Tsingy de Bemaraha 17 18 4 3027954455 CASC Fisher Madagascar -19.14194 Tombeau Vazimba 52 44.828 Parc National Tsingy de Bemaraha 17 18 6 3027954507 CASC Fisher Madagascar 210 -16.26722 Reserve d'Ankoririka 1 47.04861 17 18 1 3027954544 CASC Fisher Madagascar 30 -16.925 Reserve Speciale de Bemarivo 55 44.36833 17 18 2 3027954605 CASC Fisher Madagascar 250 -18.02649 Beanka 1 44.05051 Reserve 17 18 1 3027954302 CASC Fisher Madagascar -15.6813 Andranobe 53 49.958 17 18 1 Toamasina 3027954429 CASC A. Pauly Madagascar -20.06667 50 km N Morondava 1 44.58333 17 18 1 Toliara 3027954382 CASC Fisher Madagascar Anosy Region 90 -24.82505 Foret de Manatalinjo 1 46.57811 Parc National d'Andohahela 17 18 1 3027954326 CASC Fisher Madagascar 60 -23.48778 Antafoky 5 44.0775 17 18 1 3027954612 [309,1384,1374,1401] CASC Fisher Madagascar 135 Beza-Mahafaly 17 18 1 3027954678 CASC Fisher Madagascar 100 -23.17694 Fiherenana 1 43.96083 17 18 1 3027954748 CASC Fisher Madagascar 50 -23.23528 Fiherenana 1 43.87083 17 18 1 3027954477 CASC Fisher Madagascar 1100 Foret Classee d'Analavelona 17 18 1 3027954706 CASC Fisher Madagascar 80 -22.23306 Foret de Beroboka 1 43.36633 17 18 3 3027954427 CASC Fisher Madagascar 100 -20.045 Foret de Kirindy 55 44.66222 17 18 4 3027954571 CASC Fisher Madagascar 30 -20.06855 Foret de Kirindy 1 44.659565 17 18 7 3027954585 [346,1432,1590,1617] CASC Fisher Madagascar 110 -24.75833 Isantoria River 1 46.15717 Foret de Mahavelo 17 18 4 3027954410 CASC Fisher Madagascar 115 -24.75361 Isantoria River 5 46.1515 Foret de Mahavelo 17 18 1 3027954715 CASC Fisher Madagascar 75 -23.52417 Foret de Mite 1 44.12133 17 18 3 3027954579 [260,1308,1698,1725] CASC N Tolagnaro & Fisher Madagascar 200 -24.569 Ivohibe 74 47.204 17 18 1 3027954526 CASC Fisher Madagascar 500 -24.24067 Foret Vohidava 1 46.28783 17 18 1 3027954560 CASC Fisher Madagascar 850 -24.239 Foret Vohidava 55 46.28233 17 18 1 3027954368 CASC Fisher Madagascar 820 -23.45591 Ivahona 1 46.17376 17 18 2 3027954545 [486,1292,1806,1833] CASC Fisher Madagascar 510 -21.21836 Makay Mts. 5 45.3106 17 18 1 3027954521 CASC Fisher Madagascar 590 -21.30997 Makay Mts. 1 45.12946 17 18 2 3027954725 CASC Fisher Madagascar 70 -23.52722 Manderano 5 44.0875 17 18 3 3027954583 [346,1138,1879,1905] CASC Fisher Madagascar 177 -22.81092 Manombo 5 43.7344 17 18 1 3027954590 CASC Fisher Madagascar 165 -22.8123 Manombo 5 43.73932 17 18 1 3027954464 CASC Col du Sedro & Fisher Madagascar 900 -24.76389 Parc National d'Andohahela 1 46.75167 17 18 1 3027954378 CASC Fisher Madagascar 300 18 19 Foret d'Ambohibory Parc National d'Andohahela 17 18 6 3027954450 CASC Fisher Madagascar Parc National d'Andohahela 150 Hazofotsy 99.0 Foret de Manatalinjo 18 19 3 3027954396 CASC Fisher Madagascar -20.79528 Parc National de Kirindy Mite 51 44.147 18 19 1 3027954755 CASC Fisher Madagascar 770 Sakaraha 84.0 Parc National de Zombitse 18 19 4 3027954438 CASC Fisher Madagascar Reserve Prive Berenty 35 -24.95694 Mandrare River 5 46.2715 Foret de Bealoka 18 19 1 3027954437 CASC Fisher Madagascar 1100 Foret d'Ankazotsihitafototra Reserve Speciale d'Ambohijanahary 18 19 5 3027954418 CASC Fisher Madagascar 1050 -18.26667 Foret d'Ankazotsihitafototra 1 45.40667 Reserve Speciale d'Ambohijanahary 18 19 2 3027954582 [256,1016,438,465] CASC Fisher Madagascar 470 -22.91233 Sakaraha 1 44.53283 18 19 1 3027954379 CASC Fisher Madagascar 80 -23.52833 Sept Lacs 1 44.15556 18 19 1 3027954422 CASC Fisher Madagascar 730 southern Isoky-Vohimena Forest 18 19 1 3027954454 CASC Fisher Madagascar 1100 Foret Classee d'Analavelona 18 19 14 3027954470 CASC Fisher Madagascar 1050 -22.675 Foret Classee d'Analavelona 55 44.18667 18 19 1 3027954519 CASC Fisher Madagascar 850 -24.239 Foret Vohidava 55 46.28233 18 19 1 3027954639 [542,1377,618,645] CASC Fisher Madagascar 500 -21.21761 Makay Mts. 1 45.33917 18 19 2 3027954587 CASC Fisher Madagascar 480 -21.22336 Makay Mts. 1 45.32628 18 19 2 3027954494 CASC Fisher Madagascar 590 -21.30997 Makay Mts. 1 45.12946 18 19 4 3027954646 [440,1279,690,717] CASC Fisher Madagascar 2 -21.31664 Makay Mts. 5 45.1296 18 19 4 3027954316 CASC Fisher Madagascar 770 Sakaraha 84.0 Parc National de Zombitse 18 19 3 3027954770 CASC Fisher Madagascar Reserve Prive Berenty -25.00778 Mandrare River 50 46.306 Foret de Malaza 18 19 1