Croton (Euphorbiaceae) of the Brazilian state of Paraná: an annotated checklist, species distribution, and identification key Pereira, Amanda Da Paixão Noronha Riina, Ricarda Caruzo, Maria Beatriz Rossi Phytotaxa 2022 2022-10-28 570 3 231 274 ZQH6 Baillon (1864: 332) Baillon 332 1864 [175,642,914,941] Magnoliopsida Euphorbiaceae Croton Plantae Malpighiales 22 253 Tracheophyta species macrobothrys    Lectotype(designated by Caruzo et al.2016): — BRAZIL. Rio de Janeiro: Serra dos Órgãos, 1833,  M. Vauthier92( P00633413!; isolectotypes F 0056158F!, G 00434542!, GH 00257954!, GDC 00311416!, GDC G00311415!, MPU 014844!, P 00633414!, P 00633415!). ( Fig. 13, J–L)    Croton macrobothrysis a large tree characterized by the presence of copious red latex when the bark is cut, ovatelanceolate leaves, stipitate acropetiolar glands, elongated inflorescences, and long pedicellate flowers. Caruzo et al.(2016)recognized two subspecies for  C. macrobothrys, with only the typical one present in Paraná(  C. macrobothryssubsp. macrobothrys). The species belongs to section  Cyclostigma( van Ee et al.2011).   Distribution and habitat:—It occurs in southern ( PR, SC, RS) and southeastern Brazil(RJ, SP) ( Caruzo et al.2016, 2020, Valduga et al.2021). In Paraná, the species is more common in the east of the state, on the slopes of Serra do Mar. It grows in moist forest, between 50 and 400 melevation. ( Fig. 12).   Phenology:—Flowering from November to December, and fruiting December, January and May.   Representative specimens:—  PARANÁ: Cerro Azul,  02 November 2006,  A .   Dunaiski3222( HFIE, MBM). Guaraqueçaba,  Salto Murato,  15 November 1993,  G .  Hatschbach& J. M.   Silva59776( MBM, RB, SPF, W). Guaratuba, Descampado,  15 January 1994,  G .  Hatschbach& J. M.   Silva59816( HUEFS, K, MBM, SPF). Morretes,  Colônia Limeira,  08 November 2007,  J. M .  Silva, J.  Cordeiro& C. B.   Poliquesi6139( HCF, MBM). Tunasdo  Paraná, Pacas,  15 December 1999,  J. M .  Silva& L. M.   Abe3116( MBM, SPF, W). 3948786382 P M. Vauthier Brazil Serra dos Orgaos 22 253 1 Rio de Janeiro lectotype 3948786592 [538,1081,1346,1375] 2006-11-02 A Cerro Azul 22 253 1 Parana 3948786546 HFIE, MBM Guaraquecaba Dunaiski 22 253 1 Parana 3948786709 [309,715,1382,1409] 1993-11-15 G Murato 22 253 1 Salto 3948786348 [725,938,1382,1409] J, M Hatschbach 22 253 1 Salto 3948786606 1994-01-15 MBM, RB, SPF, W, G Guaratuba & Descampado Silva 22 253 1 Salto 3948786336 [546,770,1418,1445] J, M Hatschbach 22 253 1 Salto 3948786740 [783,1348,1418,1445] HUEFS, K, MBM, SPF Morretes Silva 22 253 1 Salto 3948786743 2007-11-08 J, M Limeira 22 253 1 Colonia 3948786577 [522,608,1454,1480] J Silva 22 253 1 Colonia 3948786737 [617,799,1454,1481] C, B Cordeiro 22 253 1 Colonia 3948786489 [809,1250,1454,1481] HCF, MBM Tunas Poliquesi 22 253 1 Colonia 3948786373 1999-12-15 J, M Pacas 22 253 1 Parana 3948786562 [385,526,1490,1517] L, M Silva 22 253 1 Parana 3948786600 [536,836,1490,1516] MBM, SPF, W Abe 22 253 1 Parana