Tetyra poecila Berg, 1879:22–24
Kernegaster gen. nov. (Heteroptera, Scutelleridae, Pachycorinae) and six new species based on genital and polychromatic synapomorphies
Schmitz, Luís Ricardo
Barcellos, Aline
Eger, Joseph
Bianchi, Filipe Michels
Zoologischer Anzeiger
2025
2025-05-11
316
97
123
(Berg, 1879)
(Berg, 1879)
2025
Berg
2025
[849,1156,1138,1157]
Insecta
Scutelleridae
Kernegaster
Animalia
Hemiptera
16
113
Arthropoda
species
poecilus
comb. nov.
( Fig. 1E; Fig. 4D–F; Fig. 7B; Fig. 8B; Fig. 9B; Fig. 10B; Fig. 12B, K; Fig. 13D and E)
Tetyra poecila Berg, 1879:22–24(description of adult and immature, host plants); Bertkau, 1879:376; Lethierry and Severin, 1893:31; Montandon, 1895:2; Schouteden, 1904:47, plate 3; Kirkaldy, 1909:284–285; Pennington, 1920:127; Bosq, 1937:127(distribution); Coscar´on and Froeschner, 2000:31(types); Mendonça et al., 2009:123; Barcellos et al., 2014:410,413,415; Carpintero et al., 2014:76(faunistic); Coscar´ on et al., 2015:94 (types); Eger et al., 2015a:766; Coscar´on, 2017:329; Brugnera et al., 2023:614( Fig. 4s).
Type material: Tetyra poecilatype material is deposited in the collection of Divisi´on Entomología, Museo de La Plata: Eight syntypes: 1–5. Argentina: Buenos Aires (1339); 6–8. Uruguai: Banda Oriental (1339) ( Coscar´on et al., 2015: Fig. 73: M, L). Here we designate as lectotype the first specimen of the serie, a male labelled as follows (each number corresponds to a different label): (1) Buenos Aires, (2) Typus, (3) T. poecilaBerg., (4) 1339/1 [green circle].
Material examined (209): BRAZIL, Bahia: ( 1F), 6.XII.1908, no collector ( CMNH 3702); Minas Gerais: ( 5 miS Prata), ( 1F), 25.I.1980, Thomas D.B. ( DBTC); Mato Grosso do Sul: Dourados( Fazenda Leiteiro), ( 2F), 9.II.1997, no collector ( UFRG); S˜ao Paulo: Guatapar´a, ( 2F, 3M), I.1945, Carrera M. ( MCNZ 2253, IBSP 7167, IBSP 7168, JEEC); S˜ao Bento do Sapucaí, ( 1F), I.1953, Buckup L. ( MCNZ 2240); S˜ao Jose´dos Campos, ( 2M), XI.1935, D. B. ( MCNZ 2244, MCNZ 2245); S˜ao Paulo, ( 1M), XI.1941, no collector ( MCNZ 2264); ibid., ( 1F), V.1926, no collector ( IBSP 7173); Sorocaba, ( 1F), X.1935, no collector ( IBSP 6810); Parana´: Curitiba, On Schinus terebinthifolius( 2F), 3.II.1988, no collector; Campo do Tenente, ( 1M), 10.X.1973, Mielke ( DZUP 217364, JEEC); Santa Catarina: Corup´a, ( 1F), XI.1954, Keller A. ( DZUP 217376); Jaguaruna, ( 2F), 18.X.2011, Perin C.; Ararangu´a, ( 1M), X.2008– V.2009, Daminelli M. ( UFRG); Rio Grande do Sul: Derrubadas( Parque Estadual do Turvo), 22.XII.1982, Bonatto S. ( UFRG); Caxias do Sul( Vila Oliva), ( 1M), 21.VII.1947, no collector ( MCNZ 2265); Canela, ( 1F), 2.III.1976, Lise A. ( MCNZ 45165); S˜ao Francisco de Paula (FLONA), ( 1F), 13. I.2007, Schwertner C.F. ( UFRG); Pareci Novo, ( 2M), XI.1946, no collector ( MCNZ 2235, MCNZ 2241); Torres, ( 1M), 28.II.1991, Moura L. ( MCNZ 50372); ibid.( 1F), 22.II.1991, Moura L. ( MCNZ 50373); ibid., ( 1M), I.1978, Mansa M.E.D. ( MCNZ 15100); S˜ao Francisco de Assis( Sanga da Areia), ( 1M), 10.III.2010, Barcellos A. & Ott R. ( MCNZ 181640); ibid., ( 1F), 27.V.2009, Barcellos A. & Ott R. ( MCNZ 181706); ibid., ( 2F), 20.VIII.2009, Barcellos A. & Ott R. ( MCNZ 181737, MCNZ 181738); Canoas, ( 1F), VII.1962, no collector ( MCNZ 1993); Imbe´( Santa Terezinha) ( 1M), 24.II.1973, Galileo M.H. ( MCNZ 8493); Tramandaí, ( 1M), IX.1955, Pintal D. ( MCNZ 2260); Triunfo, ( 1M), 12. V.1981, Becker C.J. ( MCNZ 16031); ibid., ( 1F), 23.I.1979, Gastal H.A. ( MCNZ 13004); ibid.,( 1M), 22.VI.1999, Bonaldo A. ( MCNZ 53169); ibid.( Parque COPESUL), ( 1M), 14.II.2008, Moraes R. ( MCNZ 180430); Gravataí, ( 1F), 19.XII.1966, no collector ( CIBM); Eldorado do Sul( Fazenda Kramm), ( 1M), 1.VIII.1999, Bonaldo A. & Silva A. ( MCNZ 53175); ibid., ( 1M), 19.I.1999, Moura L. ( MCNZ); ibid., ( Fazenda Sao ˜ Jos´e), ( 1M), 5–7.I.2000, Bonaldo A. & Silva A. ( MCNZ 53190); Porto Alegre, ( 1M), 20.V.1953, no collector ( MCNZ 15081); ibid., ( 1M), no date, Buck P. ( MCNZ 2263); ibid., ( 1F), 25.II.1964, Buck P. ( MCNZ 15070); ibid., ( 1F), 23.V.1951, no collector ( MCNZ 15080); ibid., ( 1F), II.1955, Buckup L. ( MCNZ 2259); ibid., ( 1M), I.1959, Buck P. ( MCNZ 15064); ibid., ( 1M), 15.IX.1956, Buck P. ( CIBM 6246); ibid., ( 1F), no date, Corsevel E. ( MCNZ 2940); ibid.( Ilha das Flores), ( 1F), 15.IV.1999, Moura L. ( MCNZ 53171); ibid., ( 1F), XI.1956, Palcya M. ( MCNZ 2256); ibid., ( Lami), ( 1F), 25.I.1979, Gastal H.A. ( MCNZ 13005); ibid., ( Morro do Sabi´a), ( 1F), 14.I.1964, no collector ( MCNZ 15065); ibid., ( Morro Santana), ( 1M), 13.VIII.1982, Bonatto S. ( UFRG 970); Guaíba, ( 1F), 28–29.I.1980, Gastal H.A. ( MCNZ 14080); ibid.( Arroio Petim), ( 5F), 19. I.1981, Gastal H.A. ( MCNZ 16996, MCNZ 45002, MCNZ 45016, MCNZ 45022, MCNZ 45032); ibid.( Florida), ( 1M), 15.III.1955, Buckup L. ( MCNZ 2257); Cidreira, ( 1F), 1936, no collector ( CIBM); Viam˜ao, ( 1M), IX.1952, Buckup L. & E. ( MCNZ 2237); ibid., ( 1F), IV.1956, Buckup L. ( MCNZ 2266); ibid., ( 2M, 3F), V.1957, Buckup L. & E. ( MCNZ 2254, MCNZ 2255, MCNZ 2258, MCNZ 2262, MCNZ 2267); ibid., ( 1M), 9. XI.1976, Becker C.J. ( MCNZ 15095); ibid., ( 1M), 13.II.1983, Becker C.J. ( MCNZ 47962); ibid., ( 1F, 1M), 11–14.IV.1983, Lise A. ( MCNZ 47916, MCNZ 47920); ibid., ( 2M), 12.IV.1983, Becker C.J. ( MCNZ 47905, MCNZ 47907); ibid., ( 1F), 13.IV.1983, Becker C.J. ( MCNZ 47964); ibid., ( 1F, 1M), 10.V.1983, Becker C.J. ( MCNZ 47975, MCNZ 47977); ibid.( Morro do Coco), ( 1M), 20.X.2018, Schmitz L.R.( MCNZ); Palmares do Sul( Ilha Grande), ( 1F, 2M), 10.IV.2003, Equipe Probio ( MCNZ 53717, MCNZ 53718, MCNZ 53877); Barra do Ribeiro( Fazenda Boa Vista), ( 1F, 1M), 13.V.2003, Equipe Probio ( MCNZ 53969, MCNZ 53998); ibid., ( 1F), 14.V.2003, Equipe Probio ( MCNZ 54261); ibid., ( 2F, 2M), 16. V.2003, Equipe Probio ( MCNZ 54079, MCNZ 54086, MCNZ 176143, MCNZ 176235); Tapes, ( 3F, 4M), XI.1963, Buck P. ( MCNZ 1945; MCNZ 1946; MCNZ 1988; MCNZ 1991; MCNZ 1992; MCNZ 1996; MCNZ 1998); ibid., ( 1M), 1.IV.1983, Hennig J.E. ( MCNZ 47886); ibid.( Fazenda Guar´a), ( 1F), 15.V.2003, Equipe Probio ( MCNZ 54182); Dom Feliciano( Fazenda Chapada), ( 1F), 18–19.III.2000, Bonaldo A. ( MCNZ 54588); Barra do Quaraí( Espinilho Leste), ( 1F), 11.I.2004, Bunde & Schwertner ( UFRG 824); ibid., Espinilho, ( 1F, 1M), 12.I.2004, Bunde & Schwertner, coletado em Scutiasp., ( UFRG 825, UFRG 826); ibid.( Fazenda Espinilho), ( 1F, 2M), 10.I.2004, Bunde & Schwertner ( UFRG 817, UFRG 818, UFRG 819); ibid., ( 2M), 11.I.2004, Bunde & Schwertner ( UFRG 821, UFRG 822); ibid., ( 1M), 17.IX.2003, Schwertner & Iserhard ( UFRG 816); ibid.( marco geogr´afico, coletado em limoeiro), ( 1M), 11.I.2004, Bunde & Schwertner ( UFRG 820); ibid., ( 1F), 11.I.2004, Bunde & Schwertner ( UFRG 823); ibid., ( 1F), 17.IX.2003, Schwertner & Iserhard ( UFRG 815); ibid., ( 2F, 4M), 24.III.2003, Bunde & Schwertner (coletado em mata ciliar) ( UFRG 809, UFRG 810, UFRG 811, UFRG 812, UFRG 813, UFRG 814); Sant’ ana do Livramento, ( 1M), 5.I.1971, Carvalho S. ( MCNZ 13133); Bage´( Rio Negro), BR-153, ( 1M), 29.III.2006, Schwertner & Cohen ( UFRG); Canguçu( Rincao ˜da Ronda), ( 1F), 3.IV.2004, no collector (campo, coletado em borda de mata) ( UFRG 455); Pelotas, ( 1F), 1. V.1962, Biezanko C.M. ( MCNZ 1989); Santa Vit´oria do Palmar( Estaç˜ao Ecol´ogica Taim) ( 1F, 2M), 27.XI.1985, Marques M.A.L. ( MCNZ 49680, MCNZ 49681, MCNZ 49682); PARAGUAY, Concepcion´: Horqueta( 44 kmeast Paraguay River), ( 1M) 8.XII.1934, Schulze A. ( USNM); San Pedro: Laguna Blanca, 665 ft, ( 1F), 19–23.II.2019, Eger J.E. & Bezark L. ( JEEC); Canindeyú: Reserva Natural del Bosque Mbaracayú, fish carrion-baited butterfly trap, 598 ft, ( 3M) 2–5.XII.2017, Eger J.E. ( JEEC); ibid., ( 6M), 19–22.XI.2016, Eger J.E. ( JEEC#271/2016); Cordillera: ( 1F), 3.III.2005, Rives J.; Caacupe´( Camp. Evangelica Neatestamenta), fish carrion-baited butterfly trap, 988 ft, ( 1M) 15 18. XI.2016, Eger J.E. ( JEEC#271/2016); Naranjo, ( 1M), 3.III.2005, Aguilar C.; ibid., ( 1M), 25.X.2006, Aguilar C. ( JEEC); Central: Capiat´a, ( 1M), 21.II.1994, Arriagada G. ( JEEC); Paraguarí: Cerro Hû, vic. Posada del Cerro, UV and MV lights, 531 ft, ( 1F), 5–6.II.2019, Eger J.E. & Bezark L.G.; Castymi, ( 1M), 5.II.2004, Rivas J. ( JEEC); Guaira´: Ybyturuzú, ( 1M), I.1994, Arriagada G. ( JEEC); Misiones: San Ignacio(vic. Hotel Rural), fish carrion-baited butterfly trap, 595 ft, ( 2M), 5–8.XII.2019, Eger, Tyson & Leavengood ( JEEC); Itapúa: Pro Cosara Nature Reserve, fish carrion-baited butterfly trap, 933 ft, ( 3M), 9.10. XII.2019, Eger, Tyson & Leavengood ( JEEC); ARGENTINA, Salta: 3–8 kmSW Campo Quijano, in Aloysia gratissima, ( 2M), 17–18.III.1995, Gill B.D. ( DBTC); Corrientes: Alvear, ( 3F, 1M) II.1991, no collector ( DARC); ibid., ( 2F, 1M) III.1991, no collector ( DARC); Entre Ríos: Pueblo Liebig, ( 1M), II–III.1997, Caire S.R. ( JEEC); ibid., ( 9F, 9M), XII.1996, Caire S.R. ( JEEC); ibid., ( 3F), VII.1988, G. Arriagada( JEEC); ibid., ( 4F, 7M), II.1991, no collector; ibid., ( 5F, 1M) III.1990, no collector; ibid., ( 2F), II.1990, no collector; ibid., ( 2F), X.1988, no collector ( DARC); URUGUAY, Canelones: ( plain on east sideof Arroyo Pando), ( 1F, 2M), 19.I.2014, Wibmer G.J.( JEEC).
Color pattern. Highly polychromatic, brownish to ferrugineus, usually with orangish intermittent stripes dorsally ( Fig. 4D–F). Head. Dorsal surface with posterior longitudinal bands of black punctures between the eyes. Clypeus with, or without posterior and median 1 + 1 spots ( Fig. 7B). Antennomeres I bicolored (first half yellow and distal half dark). Rostrum with segment III mostly yellow with a median dark blotch, segment IV mainly dark. Thorax. Pronotum with at least 1 + 1 distinct inner maculate rows with anterior and posterior portions of these rows with indistinct patterns of macules. Scutellum with darker sparse punctures distributed sparsely in longitudinal stripes. Femora with proximal small dark blotch and larger distal blotch; tibiae with medial and distal dark blotches; first and last tarsomeres dark, remaining tarsomere yellow to ferrugineus. Abdomen. Connexiva with dark punctations evenly distributed. Ventral pattern. Lateral rows usually absent, sometimes few dark spots on each sternite. Median rows with more than one solid blotch on each sternite, inner row with a few aggregations of dark spots on each sternite, usually forming two thin sub rows. Stridulatory areas usually without blotches. Area adjacent to spiracles concolorous ( Fig. 4F). Head. Wider than long. Clypeus slightly surpassing mandibular plates ( Fig. 7B). Maxillar plates subequal to mandibular plates in length. Rostrum reaching the middle of sternite III. Thorax. Pronotum with anterolateral margins sub-rectilinear, humeral angles round and slightly produced, by less than half the width of the eye ( Fig. 4D–F). Peritreme spout-like reduced, about as wide as long ( Fig. 8B). Abdomen. Posterior margin of sternite VII sinuous medially ( Fig. 9B). Genital plates. Valvifers VIII subtriangular, anterior margin convex, sutural margin posteriorly truncated, posterior margin strongly sinuous. Laterotergites IX subtriangular, anterior margin convex (partially hidden by valvifers VIII), lateral angles strongly produced and constricted by valvifer VIII ( Fig. 9B). Gynatrium. Ring sclerites with almost the entire inner margin with thick "leaf-like" sclerotized process. Posteromedian gynatrial pouch with a median trapezoidal sclerite and 1 + 1 lateral oval sclerites about the same size as the median one ( Fig. 10B). Genital capsule. Posterior margin sub-rectilinear ( Fig. 12B). Ventral rim is also sub-rectilinear in posterior view. Parameres. Crown not surpassing the width of the stem, apex obtuse ( Fig. 12K). Phallus. Aedeagus sensu strictodistal half abruptly tapered, slightly sinuous in lateral view with apex curved dorsally. Conjunctival processes II with a thin process with a distinct base. Conjunctival processes III distally sclerotized, apex acute and curved ( Fig. 13D and E). Measures.Body length (10.1–17.4) and width (7.0–11.4). Head length (2.0–3.0) and width (3.0–4.2).
Distribution:Brazil (Bahia to Rio Grande do Sul), Argentina (south till Neuqu´en) ( Coscar´on, 2017), Paraguay, and Uruguay.
Biological notes:Collected manually, with net and entomological umbrella. Attracted to ultraviolet light, mercury vapor light and fish carrion-baited butterfly traps ( Eger et al., 2015b). Found on A. gratissima(Gillies and Hook)( Verbenaceae), S. terebinthifoliaRaddi( Anacardiaceae), Scutiasp.Brongn ( Rhamnaceae), Citrussp.L. ( Rutaceae) and in a nest of Anumbius annumbi(Vieillot)( Aves: Furnariidae), was also found in diapause (via label). Recorded by Berg (1879)on Vachellia caven(Mol.), V. farnesiana(L.)( Mimosaceae), Cestrum euanthesSchltdl., Solanum bonarienseL.( Solanaceae) and Gleditsia triacanthosL. ( Fabaceae).
Comments:The only species with nymphal stages described ( Berg, 1879). The species is highly polychromatic, and as described by Berg: “It’ s so variable in color, in dots and black spots, that no two specimens are completely identical”. Differs from its congeners by the abdominal inner row of blotches that are separated by the stridulatory area forming two distinct rows, peritreme smaller and aedeagus sensu strictolong and slender.
1908-12-06
CMNH
Brazil
Bahia
16
113
CMNH 3702
1
1
Bahia
1980-01-25
DBTC
Brazil
Prata
Prata
16
113
1
1
Minas Gerais
1997-02-09
UFRG
Brazil
Fazenda Leiteiro
Dourados
16
113
2
2
Mato Grosso do Sul
1945-01
JEEC
Brazil
Guatapar´a
16
113
MCNZ 2253, IBSP 7167, IBSP 7168
5
2
3
Stao Paulo
1953-01
MCNZ
Brazil
Stao Bento do Sapucai
16
113
MCNZ 2240
1
1
Stao Paulo
1935-11
MCNZ
Brazil
Stao Jose´dos Campos
16
113
MCNZ 2244, MCNZ 2245
2
2
Stao Paulo
1941-11
MCNZ
Brazil
Stao Paulo
Stao Paulo
16
113
MCNZ 2264
1
1
Stao Paulo
1926-05
IBSP
Brazil
16
113
IBSP 7173
1
1
Stao Paulo
[1009,1484,1891,1910]
1935-10
IBSP
Brazil
Sorocaba
16
113
IBSP 6810
1
1
Stao Paulo
[818,1484,1918,1938]
1988-02-03
Brazil
Curitiba
Curitiba
16
113
2
2
Parana´
[818,1484,1947,1966]
1973-10-10
JEEC
Brazil
Campo do Tenente
16
113
DZUP 217364
1
1
Parana ´
[100,721,147,167]
1954-11
DZUP
Brazil
Corup´a
17
114
DZUP 217376
1
1
Santa Catarina
2011-10-18
Brazil
Jaguaruna
Jaguaruna
17
114
2
2
Santa Catarina
UFRG
Brazil
Ararangu´a
17
114
1
1
Santa Catarina
1982-12-22
UFRG
Brazil
Parque Estadual do Turvo
Derrubadas
17
114
1
Rio Grande do Sul
1947-07-21
MCNZ
Brazil
Vila Oliva
Caxias do Sul
17
114
MCNZ 2265
1
1
Rio Grande Do Sul
1976-03-02
MCNZ
Brazil
Canela
Canela
17
114
MCNZ 45165
1
1
Rio Grande Do Sul
UFRG
Brazil
Stao Francisco de Paula (FLONA)
17
114
1
1
Rio Grande Do Sul
1946-11
MCNZ
Brazil
Pareci Novo
17
114
MCNZ 2235, MCNZ 2241
2
2
Rio Grande Do Sul
1991-02-28
MCNZ
Brazil
Torres
Torres
17
114
MCNZ 50372
1
1
Rio Grande Do Sul
[258,713,370,390]
1991-02-22
MCNZ
Brazil
17
114
MCNZ 50373
1
1
Rio Grande Do Sul
1978-01
MCNZ
Brazil
17
114
MCNZ 15100
1
1
Rio Grande Do Sul
2010-03-10
MCNZ
Brazil
Sanga da Areia
Stao Francisco de Assis
17
114
MCNZ 181640
1
1
Rio Grande Do Sul
[193,766,454,474]
2009-05-27
MCNZ
Brazil
17
114
MCNZ 181706
1
1
Rio Grande Do Sul
2009-08-20
MCNZ
Brazil
17
114
MCNZ 181737, MCNZ 181738
2
2
Rio Grande Do Sul
[198,707,510,530]
1962-07
MCNZ
Brazil
Canoas
17
114
MCNZ 1993
1
1
Rio Grande Do Sul
1973-02-24
MCNZ
Brazil
Santa Terezinha
Imbe
17
114
MCNZ 8493
1
1
Rio Grande Do Sul
1955-09
MCNZ
Brazil
Tramandai
Tramandai
17
114
MCNZ 2260
1
1
Rio Grande Do Sul
1981-05-12
MCNZ
Brazil
Triunfo
Triunfo
17
114
MCNZ 16031
1
1
Rio Grande Do Sul
1979-01-23
MCNZ
Brazil
17
114
MCNZ 13004
1
1
Rio Grande Do Sul
[251,723,622,641]
1999-06-22
MCNZ
Brazil
17
114
MCNZ 53169
1
1
Rio Grande Do Sul
2008-02-14
MCNZ
Brazil
Parque COPESUL
17
114
MCNZ 180430
1
1
Rio Grande Do Sul
[100,590,678,697]
1966-12-19
CIBM
Brazil
Gravatai
17
114
1
1
Rio Grande Do Sul
1999-08-01
MCNZ
Brazil
Fazenda Kramm
Eldorado do Sul
17
114
MCNZ 53175
1
1
Rio Grande Do Sul
[183,577,733,753]
1999-01-19
MCNZ
Brazil
17
114
1
1
Rio Grande Do Sul
2000-01-05
2000-01-07
2000-01-05
MCNZ
Brazil
Fazenda Sao t Jos´e
17
114
MCNZ 53190
1
1
Rio Grande Do Sul
1953-05-20
MCNZ
Brazil
Porto Alegre
17
114
MCNZ 15081
1
1
Rio Grande Do Sul
MCNZ
Brazil
17
114
MCNZ 2263
1
1
Rio Grande Do Sul
1964-02-25
MCNZ
Brazil
17
114
MCNZ 15070
1
1
Rio Grande Do Sul
[181,661,845,864]
1951-05-23
MCNZ
Brazil
17
114
MCNZ 15080
1
1
Rio Grande Do Sul
1955-02
MCNZ
Brazil
17
114
MCNZ 2259
1
1
Rio Grande Do Sul
1959-01
MCNZ
Brazil
17
114
MCNZ 15064
1
1
Rio Grande Do Sul
[183,626,900,920]
1956-09-15
CIBM
Brazil
17
114
CIBM 6246
1
1
Rio Grande Do Sul
MCNZ
Brazil
17
114
MCNZ 2940
1
1
Rio Grande Do Sul
1999-04-15
MCNZ
Brazil
Ilha das Flores
17
114
MCNZ 53171
1
1
Rio Grande Do Sul
[346,766,956,976]
1956-11
MCNZ
Brazil
17
114
MCNZ 2256
1
1
Rio Grande Do Sul
[100,641,984,1004]
1979-01-25
MCNZ
Brazil
Lami
17
114
MCNZ 13005
1
1
Rio Grande Do Sul
1964-01-14
MCNZ
Brazil
Morro do Sabi´a
17
114
MCNZ 15065
1
1
Rio Grande Do Sul
1982-08-13
UFRG
Brazil
Morro Santana
17
114
UFRG 970
1
1
Rio Grande Do Sul
1980-01-28
1980-01-29
1980-01-28
MCNZ
Brazil
Guaiba
Guaiba
17
114
MCNZ 14080
1
1
Rio Grande Do Sul
MCNZ
Brazil
Arroio Petim
17
114
MCNZ 16996, MCNZ 45002, MCNZ 45016, MCNZ 45022, MCNZ 45032
5
5
Rio Grande Do Sul
1955-03-15
MCNZ
Brazil
Florida
17
114
MCNZ 2257
1
1
Rio Grande Do Sul
[241,625,1152,1171]
CIBM
Brazil
Cidreira
17
114
1
1
Rio Grande Do Sul
1952-09
MCNZ
Brazil
Viamtao
17
114
MCNZ 2237
1
1
Rio Grande Do Sul
1956-04
MCNZ
Brazil
17
114
MCNZ 2266
1
1
Rio Grande Do Sul
1957-05
MCNZ
Brazil
17
114
MCNZ 2254, MCNZ 2255, MCNZ 2258, MCNZ 2262, MCNZ 2267
5
3
2
Rio Grande Do Sul
MCNZ
Brazil
17
114
MCNZ 15095
1
1
Rio Grande Do Sul
1983-02-13
MCNZ
Brazil
17
114
MCNZ 47962
1
1
Rio Grande Do Sul
MCNZ
Brazil
17
114
MCNZ 47916, MCNZ 47920
2
1
1
Rio Grande Do Sul
1983-04-12
MCNZ
Brazil
17
114
MCNZ 47905, MCNZ 47907
2
2
Rio Grande Do Sul
[245,716,1347,1367]
1983-04-13
MCNZ
Brazil
17
114
MCNZ 47964
1
1
Rio Grande Do Sul
1983-05-10
MCNZ
Brazil
17
114
MCNZ 47975, MCNZ 47977
2
1
1
Rio Grande Do Sul
2018-10-20
MCNZ
Schmitz L. R.
Brazil
Morro do Coco
17
114
1
1
Rio Grande Do Sul
2003-04-10
MCNZ
Brazil
Ilha Grande
Palmares do Sul
17
114
MCNZ 53717, MCNZ 53718, MCNZ 53877
3
1
2
Rio Grande Do Sul
2003-05-13
MCNZ
Brazil
Fazenda Boa Vista
Barra do Ribeiro
17
114
MCNZ 53969, MCNZ 53998
2
1
1
Rio Grande Do Sul
2003-05-14
MCNZ
Brazil
17
114
MCNZ 54261
1
1
Rio Grande Do Sul
2003-05-16
MCNZ
Brazil
17
114
MCNZ 54079, MCNZ 54086, MCNZ 176143, MCNZ 176235
4
2
2
Rio Grande Do Sul
1963-11
MCNZ
Brazil
Tapes
Tapes
17
114
MCNZ 1945, MCNZ 1946, MCNZ 1988, MCNZ 1991, MCNZ 1992, MCNZ 1996, MCNZ 1998
7
3
4
Rio Grande Do Sul
[167,633,1626,1645]
1983-04-01
MCNZ
Brazil
17
114
MCNZ 47886
1
1
Rio Grande Do Sul
2003-05-15
MCNZ
Brazil
Fazenda Guar´a
17
114
MCNZ 54182
1
1
Rio Grande Do Sul
MCNZ
Brazil
Fazenda Chapada
Dom Feliciano
17
114
MCNZ 54588
1
1
Rio Grande Do Sul
2004-01-11
UFRG
Brazil
Espinilho Leste
Barra do Quarai
17
114
UFRG 824
1
1
Rio Grande Do Sul
2004-01-12
UFRG
Brazil
Espinilho
17
114
UFRG 825, UFRG 826
2
1
1
Rio Grande Do Sul
2004-01-10
UFRG
Brazil
Fazenda Espinilho
17
114
UFRG 817, UFRG 818, UFRG 819
3
1
2
Rio Grande Do Sul
2004-01-11
UFRG
Brazil
17
114
UFRG 821, UFRG 822
2
2
Rio Grande Do Sul
[215,766,1849,1869]
2003-09-17
UFRG
Brazil
17
114
UFRG 816
1
1
Rio Grande Do Sul
2004-01-11
UFRG
Brazil
marco geogr´afico
17
114
UFRG 820
1
1
Rio Grande Do Sul
2004-01-11
UFRG
Brazil
17
114
UFRG 823
1
1
Rio Grande Do Sul
[223,766,1933,1952]
2003-09-17
UFRG
Brazil
17
114
UFRG 815
1
1
Rio Grande Do Sul
2003-03-24
UFRG
Brazil
17
114
UFRG 809, UFRG 810, UFRG 811, UFRG 812, UFRG 813, UFRG 814
6
2
4
Rio Grande Do Sul
1971-01-05
MCNZ
Brazil
Sant' ana do Livramento
Sant' ana do Livramento
17
114
MCNZ 13133
1
1
Rio Grande Do Sul
2006-03-29
UFRG
Brazil
Rio Negro
Bage´
17
114
1
1
Rio Grande Do Sul
2004-04-03
UFRG
Brazil
Rincao tda Ronda
Cangucu
17
114
UFRG 455
1
1
Rio Grande Do Sul
MCNZ
Brazil
Pelotas
Pelotas
17
114
MCNZ 1989
1
1
Rio Grande Do Sul
1985-11-27
MCNZ
Brazil
Estactao Ecol´ogica Taim
Santa Vit´oria do Palmar
17
114
MCNZ 49680, MCNZ 49681, MCNZ 49682
3
1
2
Rio Grande Do Sul
1934-12-08
USNM
Paraguay
Horqueta
17
114
1
1
Concepcion
JEEC
Paraguay
203
Laguna Blanca
Laguna Blanca
17
114
1
1
San Pedro
2017-12-02
2017-12-05
2017-12-02
JEEC
Paraguay
182
Reserva Natural del Bosque Mbaracayu
Reserva Natural del Bosque Mbaracayu
17
114
3
3
Canindeyu
[902,1483,482,502]
JEEC
Paraguay
17
114
6
6
Canindeyu
[818,1163,510,530]
2005-03-03
Paraguay
Cordillera
17
114
1
1
Cordillera
JEEC
Paraguay
301
Camp. Evangelica Neatestamenta
Caacupe´
17
114
1
1
Cordillera
2005-03-03
Paraguay
Naranjo
Naranjo
17
114
1
1
Cordillera
[925,1313,594,613]
2006-10-25
JEEC
Paraguay
17
114
1
1
Cordillera
1994-02-21
JEEC
Paraguay
Capiat´a
17
114
1
1
Central
2019-02-05
2019-02-06
2019-02-05
Paraguay
162
vic. Posada del Cerro
Cerro Hu
17
114
1
1
Paraguari
[872,1284,678,697]
2004-02-05
JEEC
Paraguay
Castymi
Castymi
17
114
1
1
Paraguari
1994-01
JEEC
Paraguay
Ybyturuzu
Ybyturuzu
17
114
1
1
Guaira´
JEEC
Paraguay
181
Hotel Rural
San Ignacio
17
114
2
2
Misiones
JEEC
Paraguay
284
Pro Cosara Nature Reserve
Pro Cosara Nature Reserve
17
114
3
3
Itapua
DBTC
Argentina
Campo Quijano
17
114
2
2
Salta
[818,1385,873,892]
1991-02
DARC
Argentina
Alvear
17
114
4
3
1
Corrientes
1991-03
DARC
Argentina
17
114
3
2
1
Corrientes
JEEC
Argentina
Pueblo Liebig
17
114
1
1
Entre Rios
1996-12
JEEC
Argentina
17
114
18
9
9
Entre Rios
[900,1320,956,976]
1988-07
JEEC
G. Arriagada
Argentina
17
114
3
3
Entre Rios
1991-02
Argentina
17
114
11
4
7
Entre Rios
[1027,1377,984,1004]
1990-03
Argentina
17
114
6
5
1
Entre Rios
1990-02
Argentina
17
114
2
2
Entre Rios
[1054,1484,1012,1032]
1988-10
DARC
Argentina
17
114
2
2
Entre Rios
2014-01-19
JEEC
Wibmer G. J.
Uruguay
Arroyo Pando
17
114
3
1
2
Canelones