Revision of the Mexican and Central American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) Blahnik, Roger J. Holzenthal, Ralph W. Zootaxa 2008 2008-02-22 1711 1 1 72 Blahnik & Holzenthal, 2008 Blahnik & Holzenthal 2008 [151,403,311,337] Insecta Glossosomatidae Mortoniella Animalia Trichoptera 21 22 Arthropoda species carinula sp. nov.   Fig. 8   Mortoniella carinulais similar to a group of species, including  M. opinionis, n. sp.,  M. papillata, n. sp.,  M. redunca, n. sp.,  M. sicula, n. sp, and  M. umbonata, n. sp., all distinguished by the character combination of having an inferior appendage invaginated mesally and with protruding, often apically acute, apicolateral projections, and also a phallicata with well developed dorsolateral processes. Among these species,  M. carinulais differentiated by the form of the dorsal phallic spine, which is more or less linearly extended apically, rather than being distinctly upturned, and keeled apicomesally, making it appear slightly widened at the apex in lateral view. Some minor variation was encountered in the form of the apicolateral projections of the inferior appendage, as well as in the form of tergum X, as shown in Fig. 8. Adult. Length of forewing: male 3.5–4.1 mm, female 3.8–4.7 mm. Forewing with forks I, II, and III, hind wing with forks II and III. Overall color medium brown, palps darker brown, apices of tibiae, tarsal and basal antennal segments light brown. Wing bar at anastamosis indistinctly marked with pale, light brown setae. Ventral process of abdominal segment VI (male) short, ventrally oriented, subtriangular, subacute apically, not constricted basally. Male genitalia. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin forming rounded projection in dorsal half, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, dorsal excision narrow, much less than half width of segment. Tergum X with short, acute apicomesal projection and projecting lateral lobes, lateral lobes with apices narrowed, subacute, mesally curved. Inferior appendage without apicomesal projection, apicolateral projections very narrow, elongate, apices acuminate, usually slightly ventrally recurved ( Fig. 8A), sometimes nearly straight or slightly dorsally curved ( Fig. 8E), extreme apex scabrous; basolaterally, with short, rounded projections on each side; mesal pockets of inferior appendage with apical processes short, dorsally curved. Paramere appendages elongate, narrow, apex widened and scabrous. Dorsal phallic spine slightly enlarged ventrally, dorsal contour sinuous, apex only slightly upturned, appearing somewhat widened due to dorsomesal carina; as viewed dorsally, with apex acute. Phallicata relatively short, dorsally with paired, upright, subquadrate processes, laterally with broadly rounded, sclerotized explanate projection on either side. Endophallic membrane with single prominent, stout, curved spine.     Holotypemale: COSTA RICA: Cartago: Reserva Tapantí, Quebrada Segunda@ administration building, 09°45'40"N, 083°47'13"W,  1250 m,  9–10.v.1990, Holzenthal& Blahnik( UMSP000001358) (pinned) ( UMSP).    Paratypes: COSTA RICA: Cartago: Reserva[Parque Nacional] Tapantí, Río Grande de Orosí, 09°41'10"N, 083°45'22"W,  1650 m,  18–21.iii.1987, Holzenthal, Hamilton& Heyn— 1 male(pinned) ( UMSP);  Quebrada Palmitos& falls, ca. 9 km(road) NW tunnel, 09°43'12"N, 083°46'48"W,  1400 m,  2– 3.vi.1990, Holzenthal, Blahnik& Muñoz— 2 females(pinned) ( UMSP); same locality,  1–2.viii.1990, Holzenthal, Blahnik& Muñoz— 1 male, 3 females(pinned) ( NMNH); same locality,  24–25.iii.1991, Holzenthal, Muñoz& Huisman— 2 males(pinned), 1 male(alcohol) ( UMSP); same locality,  21.ii.1992, Holzenthal, Muñoz& Kjer— 1 male(pinned) ( UMSP);  Río Dos Amigos& falls, ca. 6 km(road) NW tunnel, 09°42'14"N, 083°46'59"W,  1500 m,  4–5.viii.1990, Holzenthal, Blahnik& Muñoz— 1 male(pinned) ( UMSP); waterfall, ca. 1 km(road) NW tunnel, 09°41'24"N, 083°45'36"W,  1600 m,  2–3.viii.1990, Holzenthal, Blahnik& Muñoz— 3 males, 1 female(pinned) ( NMNH); same locality and date as holotype— 2 females(pinned), 1 male(alcohol) ( UMSP); same locality,  23.viii.1990, Holzenthal& Huisman— 1 female(pinned) ( UMSP);   Puntarenas: Río Bellavista, ca.  1.5 kmNW Las Alturas, 08°57'04"N, 082°50'46"W,  1400 m,  2–3.viii.1987, Holzenthal Morse& Clausen— 1 male, 6 females(pinned) ( UMSP);   San José: Quebrada   FIGURE 8.  Mortoniella carinula, new species. Male genitalia: A—lateral; B—segment IX and tergum X, dorsal; C— phallic ensemble, ventral; D—dorsal phallic spine, dorsal; E—inferior appendage variation, lateral; F—tergum X variation, dorsal   Caraiges, 2.5 km(road) SW La Legua, 09°44'02"N, 084°07'12"W,  1400 m,  22.i.1992, Holzenthal, Kjer& Quesada— 3 males, 4 females(pinned) ( UMSP); El Salvaje, Río Tabarcia, 8 km(road) E Palmichal, 09°50'49"N, 084°07'12"W,  1650 m,  19–20.i.1992, Holzenthal, Kjer& Quesada— 6 males, 6 females(pinned) ( INBIO). Etymology. This species is named  M. carinula, diminutive of the Latin word carina, meaning keel, and referring to the projecting keeled apex of the dorsal phallic spine that is a distinctive characteristic of this species. 1990-05-09 1990-05-10 1990-05-09 UMSP Holzenthal & Blahnik Costa Rica 1250 9.761111 Quebrada Segunda 21 -83.78694 Reserva Tapanti 21 22 UMSP000001358 1 Cartago holotype 1987-03-18 1987-03-21 1987-03-18 UMSP Holzenthal & Hamilton & Heyn Costa Rica Reserva 1650 9.686111 Rio Grande de Orosi 21 -83.75611 Tapanti 21 22 1 1 Cartago paratype 1990-06-02 1990-06-03 1990-06-02 UMSP Holzenthal & Blahnik & Munoz Costa Rica 1400 9.719999 Quebrada Palmitos 21 -83.78001 21 22 2 2 Cartago paratype 1990-08-01 1990-08-02 1990-08-01 NMNH Holzenthal & Blahnik & Munoz Costa Rica 1400 9.719999 Quebrada Palmitos 21 -83.78001 21 22 4 3 1 Cartago paratype 1991-03-24 1991-03-25 1991-03-24 UMSP Holzenthal & Munoz & Huisman Costa Rica 1400 9.719999 Quebrada Palmitos 21 -83.78001 21 22 3 3 Cartago paratype 1992-02-21 1991-03-25 1991-03-24 UMSP Holzenthal & Munoz & Kjer Costa Rica 1400 9.719999 Quebrada Palmitos 21 -83.78001 21 22 1 1 Cartago paratype 1990-08-04 1990-08-05 1990-08-04 UMSP Holzenthal & Blahnik & Munoz Costa Rica 1500 9.703889 Rio Dos Amigos 21 -83.78306 21 22 1 1 Cartago paratype 1990-08-02 1990-08-03 1990-08-02 NMNH Holzenthal & Blahnik & Munoz Costa Rica 1600 9.690001 21 -83.76 21 22 4 1 3 Cartago paratype 1990-08-02 1990-08-03 1990-08-02 UMSP Holzenthal & Blahnik & Munoz Costa Rica 1600 9.690001 21 -83.76 21 22 3 2 1 Cartago holotype 1990-08-23 1990-08-03 1990-08-02 UMSP Holzenthal & Huisman Costa Rica 1600 22 23 9.690001 21 -83.76 21 22 1 1 Cartago paratype 1987-08-02 1987-08-03 1987-08-02 UMSP Holzenthal Morse & Clausen Costa Rica 1400 8.951111 Rio Bellavista 21 -82.846115 22 23 7 6 1 Puntarenas paratype [1195,1436,1998,2024] Quebrada Costa Rica San Jose 22 23 1 San Jose paratype 1992-01-19 1992-01-22 1992-01-19 UMSP, INBIO Caraiges & SW La Legua & Holzenthal & Kjer & Quesada & El Salvaje & E Palmichal 1400 9.846944 Rio Tabarcia 21 -84.12 23 24 19 10 9