Revision of the Mexican and Central American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae)
Blahnik, Roger J.
Holzenthal, Ralph W.
Zootaxa
2008
2008-02-22
1711
1
1
72
Blahnik & Holzenthal, 2008
Blahnik & Holzenthal
2008
[151,403,311,337]
Insecta
Glossosomatidae
Mortoniella
Animalia
Trichoptera
21
22
Arthropoda
species
carinula
sp. nov.
Fig. 8 Mortoniella carinulais similar to a group of species, including M. opinionis, n. sp., M. papillata, n. sp., M. redunca, n. sp., M. sicula, n. sp, and M. umbonata, n. sp., all distinguished by the character combination of having an inferior appendage invaginated mesally and with protruding, often apically acute, apicolateral projections, and also a phallicata with well developed dorsolateral processes. Among these species, M. carinulais differentiated by the form of the dorsal phallic spine, which is more or less linearly extended apically, rather than being distinctly upturned, and keeled apicomesally, making it appear slightly widened at the apex in lateral view. Some minor variation was encountered in the form of the apicolateral projections of the inferior appendage, as well as in the form of tergum X, as shown in Fig. 8. Adult. Length of forewing: male 3.5–4.1 mm, female 3.8–4.7 mm. Forewing with forks I, II, and III, hind wing with forks II and III. Overall color medium brown, palps darker brown, apices of tibiae, tarsal and basal antennal segments light brown. Wing bar at anastamosis indistinctly marked with pale, light brown setae. Ventral process of abdominal segment VI (male) short, ventrally oriented, subtriangular, subacute apically, not constricted basally. Male genitalia. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin forming rounded projection in dorsal half, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, dorsal excision narrow, much less than half width of segment. Tergum X with short, acute apicomesal projection and projecting lateral lobes, lateral lobes with apices narrowed, subacute, mesally curved. Inferior appendage without apicomesal projection, apicolateral projections very narrow, elongate, apices acuminate, usually slightly ventrally recurved ( Fig. 8A), sometimes nearly straight or slightly dorsally curved ( Fig. 8E), extreme apex scabrous; basolaterally, with short, rounded projections on each side; mesal pockets of inferior appendage with apical processes short, dorsally curved. Paramere appendages elongate, narrow, apex widened and scabrous. Dorsal phallic spine slightly enlarged ventrally, dorsal contour sinuous, apex only slightly upturned, appearing somewhat widened due to dorsomesal carina; as viewed dorsally, with apex acute. Phallicata relatively short, dorsally with paired, upright, subquadrate processes, laterally with broadly rounded, sclerotized explanate projection on either side. Endophallic membrane with single prominent, stout, curved spine.
Holotypemale: COSTA RICA: Cartago: Reserva Tapantí, Quebrada Segunda@ administration building, 09°45'40"N, 083°47'13"W, 1250 m, 9–10.v.1990, Holzenthal& Blahnik( UMSP000001358) (pinned) ( UMSP). Paratypes: COSTA RICA: Cartago: Reserva[Parque Nacional] Tapantí, Río Grande de Orosí, 09°41'10"N, 083°45'22"W, 1650 m, 18–21.iii.1987, Holzenthal, Hamilton& Heyn— 1 male(pinned) ( UMSP); Quebrada Palmitos& falls, ca. 9 km(road) NW tunnel, 09°43'12"N, 083°46'48"W, 1400 m, 2– 3.vi.1990, Holzenthal, Blahnik& Muñoz— 2 females(pinned) ( UMSP); same locality, 1–2.viii.1990, Holzenthal, Blahnik& Muñoz— 1 male, 3 females(pinned) ( NMNH); same locality, 24–25.iii.1991, Holzenthal, Muñoz& Huisman— 2 males(pinned), 1 male(alcohol) ( UMSP); same locality, 21.ii.1992, Holzenthal, Muñoz& Kjer— 1 male(pinned) ( UMSP); Río Dos Amigos& falls, ca. 6 km(road) NW tunnel, 09°42'14"N, 083°46'59"W, 1500 m, 4–5.viii.1990, Holzenthal, Blahnik& Muñoz— 1 male(pinned) ( UMSP); waterfall, ca. 1 km(road) NW tunnel, 09°41'24"N, 083°45'36"W, 1600 m, 2–3.viii.1990, Holzenthal, Blahnik& Muñoz— 3 males, 1 female(pinned) ( NMNH); same locality and date as holotype— 2 females(pinned), 1 male(alcohol) ( UMSP); same locality, 23.viii.1990, Holzenthal& Huisman— 1 female(pinned) ( UMSP); Puntarenas: Río Bellavista, ca. 1.5 kmNW Las Alturas, 08°57'04"N, 082°50'46"W, 1400 m, 2–3.viii.1987, Holzenthal Morse& Clausen— 1 male, 6 females(pinned) ( UMSP); San José: Quebrada FIGURE 8. Mortoniella carinula, new species. Male genitalia: A—lateral; B—segment IX and tergum X, dorsal; C— phallic ensemble, ventral; D—dorsal phallic spine, dorsal; E—inferior appendage variation, lateral; F—tergum X variation, dorsal Caraiges, 2.5 km(road) SW La Legua, 09°44'02"N, 084°07'12"W, 1400 m, 22.i.1992, Holzenthal, Kjer& Quesada— 3 males, 4 females(pinned) ( UMSP); El Salvaje, Río Tabarcia, 8 km(road) E Palmichal, 09°50'49"N, 084°07'12"W, 1650 m, 19–20.i.1992, Holzenthal, Kjer& Quesada— 6 males, 6 females(pinned) ( INBIO). Etymology. This species is named M. carinula, diminutive of the Latin word carina, meaning keel, and referring to the projecting keeled apex of the dorsal phallic spine that is a distinctive characteristic of this species.
1990-05-09
1990-05-10
1990-05-09
UMSP
Holzenthal & Blahnik
Costa Rica
1250
9.761111
Quebrada Segunda
21
-83.78694
Reserva Tapanti
21
22
UMSP000001358
1
Cartago
holotype
1987-03-18
1987-03-21
1987-03-18
UMSP
Holzenthal & Hamilton & Heyn
Costa Rica
Reserva
1650
9.686111
Rio Grande de Orosi
21
-83.75611
Tapanti
21
22
1
1
Cartago
paratype
1990-06-02
1990-06-03
1990-06-02
UMSP
Holzenthal & Blahnik & Munoz
Costa Rica
1400
9.719999
Quebrada Palmitos
21
-83.78001
21
22
2
2
Cartago
paratype
1990-08-01
1990-08-02
1990-08-01
NMNH
Holzenthal & Blahnik & Munoz
Costa Rica
1400
9.719999
Quebrada Palmitos
21
-83.78001
21
22
4
3
1
Cartago
paratype
1991-03-24
1991-03-25
1991-03-24
UMSP
Holzenthal & Munoz & Huisman
Costa Rica
1400
9.719999
Quebrada Palmitos
21
-83.78001
21
22
3
3
Cartago
paratype
1992-02-21
1991-03-25
1991-03-24
UMSP
Holzenthal & Munoz & Kjer
Costa Rica
1400
9.719999
Quebrada Palmitos
21
-83.78001
21
22
1
1
Cartago
paratype
1990-08-04
1990-08-05
1990-08-04
UMSP
Holzenthal & Blahnik & Munoz
Costa Rica
1500
9.703889
Rio Dos Amigos
21
-83.78306
21
22
1
1
Cartago
paratype
1990-08-02
1990-08-03
1990-08-02
NMNH
Holzenthal & Blahnik & Munoz
Costa Rica
1600
9.690001
21
-83.76
21
22
4
1
3
Cartago
paratype
1990-08-02
1990-08-03
1990-08-02
UMSP
Holzenthal & Blahnik & Munoz
Costa Rica
1600
9.690001
21
-83.76
21
22
3
2
1
Cartago
holotype
1990-08-23
1990-08-03
1990-08-02
UMSP
Holzenthal & Huisman
Costa Rica
1600
22
23
9.690001
21
-83.76
21
22
1
1
Cartago
paratype
1987-08-02
1987-08-03
1987-08-02
UMSP
Holzenthal Morse & Clausen
Costa Rica
1400
8.951111
Rio Bellavista
21
-82.846115
22
23
7
6
1
Puntarenas
paratype
[1195,1436,1998,2024]
Quebrada
Costa Rica
San Jose
22
23
1
San Jose
paratype
1992-01-19
1992-01-22
1992-01-19
UMSP, INBIO
Caraiges & SW La Legua & Holzenthal & Kjer & Quesada & El Salvaje & E Palmichal
1400
9.846944
Rio Tabarcia
21
-84.12
23
24
19
10
9