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        <cito:cites>Hydrochus elsjeae Makhan 1994: 40</cito:cites>
        <cito:cites>Hydrochus obscurus Sharp</cito:cites>
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        <dc:title>Taxonomy of water beetles in the genus Hydrochus Leach, 1817, from Bolivia, Brazil and Paraguay (Coleoptera: Hydrochidae)</dc:title>
        <dc:creator>Perkins, Philip D.</dc:creator>
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        <bibo:journal>Zootaxa</bibo:journal>
        <dc:date>2021</dc:date>
        <bibo:pubDate>2021-06-28</bibo:pubDate>
        <bibo:volume>4994</bibo:volume>
        <bibo:issue>1</bibo:issue>
        <bibo:pageStart>1</bibo:pageStart>
        <bibo:pageEnd>93</bibo:pageEnd>
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        <dwc:ID-CoL>3N67G</dwc:ID-CoL>
        <dwc:authority>Makhan</dwc:authority>
        <dwc:authorityName>Makhan</dwc:authorityName>
        <dwc:authorityYear>2021</dwc:authorityYear>
        <dwc:box>[151,489,1683,1710]</dwc:box>
        <dwc:class>Insecta</dwc:class>
        <dwc:family>Hydrochidae</dwc:family>
        <dwc:genus>Hydrochus</dwc:genus>
        <dwc:kingdom>Animalia</dwc:kingdom>
        <dwc:order>Coleoptera</dwc:order>
        <dwc:pageId>43</dwc:pageId>
        <dwc:pageNumber>44</dwc:pageNumber>
        <dwc:phylum>Arthropoda</dwc:phylum>
        <dwc:rank>species</dwc:rank>
        <dwc:species>elsjeae</dwc:species>
        <dwc:status>stat. nov.</dwc:status>
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        <spm:hasContent>     Hydrochus elsjeae Makhan 1994: 40   Hydrochus obscurusSharp; Oliva (1996), incorrect synonymy by Oliva</spm:hasContent>
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        <spm:hasContent>   Originallydescribed from Bolivia, based on one male. Only the holotypehas been examined in this study.</spm:hasContent>
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        <spm:hasContent>  Remarks.I have examined the holotypeof  H. elsjeaeMakhan, including the male genitalia (ZMHB). Oliva (1996)synonymized  H. elsjeaeMakhanwith  H. obscurusSharp. However, this is undoubtedly incorrect as the male genitalia of the two markedly differ. Among the most obvious differences is that the male genitalia of  H. elsjeaehas large pseudoparameres, whereas  H. obscuruslacks pseudoparameres. Labels on the holotypeof  H. elsjeaeare: “Bo- livien 750m/ Prov. Sara II-06 / J. Steinbach S.V. // Zool. Mus. Berlin // Hydrochus elsjeae/ det. D. Makhan / 1994 // [red] Holotype// Hydrochus obscurus Sharp 1882det. A. Oliva 1996// HABITUS/GENIT. DIGITAL IMAGES captured 2019 P.D. Perkins”.</spm:hasContent>
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        <dwc:box>[151,1335,1852,1879]</dwc:box>
        <dwc:country>Bolivia</dwc:country>
        <dwc:location>Originally</dwc:location>
        <dwc:pageId>43</dwc:pageId>
        <dwc:pageNumber>44</dwc:pageNumber>
        <dwc:specimenCount>1</dwc:specimenCount>
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