Argia jocosa Hagen in Selys 1865: 408 Calvert 1909: 120 Heckman 2008: 361 Pérez-Gutiérrez & Palacino-Rodriguez 2011: 212 De Marmels 2012: 253 A. cupraurea A. orichalcea A. jocosa Hag. A. cuprea Damselflies of the genus Argia (Odonata: Coenagrionidae) from Ecuador with descriptions of five new species Garrison, Rosser W. Ellenrieder, Natalia Von Zootaxa 2018 2018-09-03 4470 1 1 69 GGPG Hagen in Selys, 1865 Hagen in Selys 1865 [151,568,332,358] Insecta Coenagrionidae Argia Animalia Odonata 17 18 Arthropoda species jocosa   Figs. 3(rear of head ♀), 7 (labrum ♀); 17 (head, thorax, S1– 4 ♂); 28 (thorax ♀); 50–52 (S1– 10 ♂), 62 (S1– 5 ♀), 77–79 (mesostigmal plates ♀), 95 (genital ligula ♂), 111 (appendages ♂), 128 ( typelabels), 139 (map).      Argia jocosaHagen in Selys 1865: 408(36 separate), (description of ♂, "Santa-Fe de Bogota, par Lindig. (Coll. Hagen.)"); Calvert 1901; 71 (key); Calvert inCalvert & Hagen 1902: 111 (catalog);  Calvert 1909: 120, 143 (key, Colombia);  Heckman 2008: 361(key, listed erroneously from Ecuador);  Pérez-Gutiérrez & Palacino-Rodriguez 2011: 212( Colombia):  De Marmels 2012: 253(distribution in Venezuela, illustrations and comparison with  A. cupraureaand  A. orichalcea).  Types.  LECTOTYPEby present designation, ♂. " St. Fede Bogota/ Lindig 1863", " A. jocosa Hag./ an A. cupreavar?", "Penis/ drawn [written by C.H. Kennedy]", "Hagen [printed]". MCZ 12194 ( Fig. 128).  Paralectotype ♂, same locality label. [ MCZ].   Specimens examined (only those from Ecuadorlisted).  1 ♂, Sucumbios Prov., Bellavista River, 32 kmsoutheast of Lago Agrio["A wide river, 3–4 meters across with good flow, bottom stony with some deeper holes, bordered by secondary forest (W-721)"] ( 0°0'21" S, 76°38'37" W,  285 m),  11 January 2014, David Wagner, William Haber, Fred Morrisonleg. [ RWG].  De Marmels (2012)provided a thorough review of this species within Venezuelaand cleared up its misidentification with  A. cupraurea. Its male is the most variable of all of the metallic red species. The lectotype(Fig. 50) is characterized, as Calvert (1901)noted by "Segments 3–7 black on dorsum, basal half of 3–6 blue, 8–10 blue on dorsum." Most males of this species (including all those known from Venezuela) differ in having S3–5 blue for two-thirds or more of their length (as in Fig. 52). The male from Bellavista River, Ecuador(Fig. 52) has the inferior black on S3–6 thicker than on males from Colombiaand Venezuela. Another male from Lloraudo River, Colombiahas S3–7 predominantly black (Fig. 51) thus approaching the color pattern of  cuprea-like species  A. divesand  A. philipi. The thoracic dorsum is normally entirely metallic (as in Fig. 20) but the male from Bellavista River, Ecuador( Fig. 17) has a narrow blue antehumeral stripe and another male from Reserva Natural El Rey Zamuro, Colombiahas a similar pale antehumeral stripe that is about as wide as the middorsal black stripe. The middorsal and humeral stripe, the latter of which is divided above at its upper third, show no metallic coloration although metallic reflections are present on the prothorax and head. All of these males conform to what we believe to represent  A. jocosaas the male appendages and genital ligula (unique for this species) match those of the lectotype( Fig. 95). The Bellavista River pair represents a new record for Ecuador. We also examined an additional 296 ♂♂and 75♀♀from Colombiaand Venezuelafor this account. See under  A. philipiaccount below. 2608963152 C. H. Kennedy St. Fe 17 18 1 1 Bogota lectotype 2608963367 MCZ C. H. Kennedy St. Fe 17 18 1 1 Bogota paralectotype 2608963051 2014-01-11 RWG David Wagner & William Haber & Fred Morrison Ecuador 285 -0.0058333334 Lago Agrio 21 -76.64361 Bellavista River 17 18 1 1 Sucumbios