Deeleman-Reinhold, 1986b : 217
Huber 2011 : 55
C. anai
C. phyllicola
Revision and cladistic analysis of the Southeast Asian leaf-dwelling spider genus Calapnita Simon (Araneae, Pholcidae)
Bernhard A. Huber
Zootaxa
2017
4219
1
1
63
92X73
Deeleman-Reinhold, 1986
Deeleman-Reinhold
1986
[151,747,1534,1560]
Arachnida
Pholcidae
Calapnita
Animalia
Araneae
25
26
Arthropoda
species
phasmoides
Calapnita phasmoides Deeleman-Reinhold, 1986b: 217, figs 46–51, 59e ( ♂ ♀). Huber 2011: 55(except specimens from Sumatra and Java; see C. anai), figs 194–195 (not figs 70–71, 196–197, see C. phyllicola).
Diagnosis. Males are easily distinguished from most congeners (except C. anaiand C. deelemanae) by simple appendix with small subdistal side-branch ( Fig. 101); from C. anaiand C. deelemanaeby tip of procursus (not strongly bent; with distinct retrolateral process; dorsal flat process not gradually but abruptly widening; Figs 98– 100); females of these three species and C. subphyllicolamay be indistinguishable externally, but the slightly elongated and anteriorly converging pore plates might be distinctive of C. phasmoides( Fig. 102).
Notes. For the present study, almost all material listed in Huber (2011)was restudied (except specimens from Sumatrabecause fresh material from the same locality in Sumatrawas available; see C. anaibelow); in addition, the holotypefrom East Kalimantan, Borneo, deposited in RMNH( ARA9574), was also examined. As mentioned earlier ( Huber 2011), males from outside East and Central Kalimantan(at that time only Sumatraand Java, now also Sabahand Malay Peninsula) differ slightly from males from Borneo. All specimens from outside Borneo are here separated from C. phasmoidesand formally described as C. anai(see below). FIGURES 89–97. Calapnita subphyllicolaDeeleman-Reinhold, SEM micrographs (ZFMK, Ar 16004–05). 89. Male cheliceral apophyses. 90–91. Right procursus tip, retrolatero-dorsal and retrolateral views. 92. Left procursus tip, prolaterodorsal view. 93. Left genital bulb, prolateral view (arrow points at membranous process). 94–95. Male and female ALS (arrows point at PMS spigots). 96. Male gonopore. 97. Epigynum, ventral view. a, appendix; b, genital bulb; e, embolus; p, procursus. Scale lines: 10 µm (94–96), 50 µm (89–92), 100 µm (93, 97). FIGURES 98–106. Calapnita phasmoidesDeeleman-Reinhold(98–102; ♂ from Tumbang Tahai, RMNH, ARA 17451, ♀ from Sepaku, RMNH, ARA 17379) and C. sp. from Ulu Senegang (103–106; RMNH, ARA 17810, 17812). 98–100, 103–105. Left procursi, prolateral, dorsal, and retrolateral views. 101, 106. Left bulbal appendices (a), prolateral views (e, embolus; arrow points at membranous process). 102. Cleared female genitalia, dorsal view (arrow points at membranous sac). Scale lines: 0.5 mm (98–101, 103–106; all male structures at same scale), 0.3 mm (102). Anadditional very similar species seems to occur in Sabah, Ulu Senegang( 5.36°N, 116.03°E, 3♂ 6♀, RMNH, ARA17810–12). In males of this putative species, the procursus looks intermediate (in lateral view almost identical to C. anai; in dorsal view with distinct retrolateral process as in C. phasmoides; Figs 103–105). The appendix is very similar to C. phasmoidesand C. anai( Fig. 106). These specimens are all in very bad condition and the species is thus not formally described. For the present study, the genitalia of one of the four females available from East and Central Kalimantanwere cleared ( Fig. 102). It became obvious that the previously cleared female ( Huber 2011: figs 70–71, 196–197) from Tumbang Tahai, Central Kalimantan( RMNH, ARA17813), which at that time accompanied an unambiguous C. phasmoidesmale ( RMNH, ARA17451), was a misidentified C. phyllicola. In contrast to C. phyllicola, the female internal genitalia of C. phasmoidesare characterized by oval rather than drop-shaped pore plates and an unpaired rather than paired internal ‘sac’ ( Fig. 102).
Distribution. Known from East and Central Kalimantanonly ( Fig. 281). Specimens from Sumatraand Javapreviously assigned tentatively to this species are here assigned to the newly described C. anai(see below).
1424726748
Sumatra and Java
25
26
2
1
1
Sumatra and Java
1424726738
RMNH, ARA
28
29
5.36
Ulu Senegang
783
116.03
An
27
28
ARA 17810-12
9
6
3
Sabah