Revision and cladistic analysis of the Southeast Asian leaf-dwelling spider genus Calapnita Simon (Araneae, Pholcidae) Bernhard A. Huber Zootaxa 2017 4219 1 1 63 93W53 [151,354,151,177] Arachnida Pholcidae Calapnita Animalia Araneae 14 15 Arthropoda species kubah sp. nov.   Diagnosis. Males are easily distinguished from known congeners by morphology of male palps (heavily sclerotized appendix with three terminal tines; subdistally widened and sclerotized embolus; Figs 33, 39); females are difficult to separate from similar species (  C. deelemanae,  C. phasmoides,  C. subphyllicola), but seem to differ in their internal genitalia (pore plates not round; membranous ‘sac’ absent; Fig. 37).   Etymology. The species name is derived from the typelocality; noun in apposition.    Material examined. Holotype. MALAYSIA-BORNEO: ♂, ZFMK( Ar15969), Sarawak, Kubah National Park, along Main Trail( 1.611°N, 110.191– 110.195°E),  160–200 ma.s.l., undersides of palm leaves,  13.vii.2014( B.A. Huber, S.B. Huber).   Other material. MALAYSIA-BORNEO: 10♂ 9♀ 1 juv., ZFMK(Ar 15970–71), and 1♂ 1♀, SMK, same data as holotype;  5♀in absolute ethanol, ZFMK( Bor191), same data.  2♂ 1♀, ZFMK( Ar15972), Sarawak, Kubah National Park, near entrance to Waterfall Trail( 1.606°N, 110.187°E),  300 ma.s.l., undersides of large palm leaves,  12.vii.2014( B.A. Huber, S.B. Huber);  5♀3 juvs in absolute ethanol, ZFMK( Bor222), same data.   Description. Male( holotype) MEASUREMENTS. Total body length 6.9, carapace width 1.1. Leg 1: 42.9 (10.1 + 0.5 + 10.3 + 20.0 + 2.0), tibia 2: 7.6, tibia 3: 4.5, tibia 4: 7.2; tibia 1 L/d: 98. Distance PME-PME 245 µm, diameter PME 115 µm, distance PME- ALE ~25 µm; no trace of AME. COLOR. Prosoma mostly pale whitish, sternum with indistinct darker marks; legs pale ochre-yellow with dark brown patellae and tibia-metatarsus joints; abdomen monochromous pale ochre-gray. BODY. Habitus as in Fig. 7; ocular area barely elevated ( Fig. 42), each triad on very low hump; carapace without median furrow; clypeus unmodified; sternum wider than long (0.70/0.64), unmodified. CHELICERAE. As in Fig. 35, with pair of simple scaly apophyses ( Fig. 38) near lamellae and pair of indistinct lateral humps proximally; without modified hairs; without stridulatory ridges. PALPS. As in Figs 33–34; coxa unmodified; trochanter with long retrolatero-ventral apophysis; femur without processes; procursus rather straight, long, with strong prolatero-ventral ‘knee’, with prolateral process at halflength continuing towards tip of procursus as transparent lamina; bulb with small membranous (but not wormshaped) proximal process; large appendix with three heavily sclerotized distal tines ( Fig. 39); long embolus arising from base of appendix, with distinctive sclerotized subdistal widening ( Fig. 39), with semitransparent tip. LEGS. Without spines and curved hairs; few vertical hairs; retrolateral trichobothrium on tibia 1 at 2%; prolateral trichobothrium absent on tibia 1, present on other tibiae; tarsus 1 pseudosegments not visible in dissecting microscope.  Male(variation). Tibia 1 in13 other males: 9.5–10.4 (mean 9.9).  Female. In general similar to male; eye triads slightly closer together (distance PME-PME 220 µm). Tibia 1 in 11 females: 7.6–9.2 (mean 8.4). Epigynum very simple, weakly sclerotized and folded ( Fig. 43), internal anterior arch visible through cuticle ( Fig. 36); with simple short posterior ‘knob’; internal genitalia as in Fig. 37.  Natural history. The spiders were only found on the undersides of palm leaves. At one site (“along Main Trail”), palms were common and spiders abundant; at the other site (“near entrance to Waterfall Trail”) palms were rare and so were spiders.   Distribution. Known from typelocality in western  Sarawakonly ( Fig. 281). 1424726758 2014-07-13 ZFMK B. A. Huber & S. B. Huber Malaysia Main Trail 180 1.611 Ar 78 110.195 National Park 15 16 1 1 holotype 1424726760 2014-07-13 ZFMK, SMK B. A. Huber & S. B. Huber Malaysia Main Trail 180 1.611 Ar 78 110.195 National Park 15 16 22 10 11 1424726759 [299,909,1860,1885] 2014-07-13 ZFMK Bor Malaysia Main Trail 180 1.611 Ar 78 110.195 National Park 15 16 5 5 1424726755 2014-07-12 ZFMK B. A. Huber & S. B. Huber Malaysia Waterfall Trail 300 1.606 Ar 78 110.187 National Park 15 16 3 1 2 1424726756 [587,1252,1930,1955] 2014-07-12 ZFMK Bor Malaysia Waterfall Trail 300 1.606 Ar 78 110.187 National Park 15 16 5 5