Kulczyński, 1911: 13 Zaitunia schmitzi Lehtinen 1967: 275 Zonstein 2009b: 127 A revision of the spider genus Zaitunia (Araneae, Filistatidae) Sergei Zonstein Yuri M. Marusik European Journal of Taxonomy 2016 2016-07-20 214 1 97 7GBLC Kulczynski, 1911 Kulczynski 1911 [572,1015,1578,1605] Arachnida Filistatidae Zaitunia Animalia Araneae 14 15 Arthropoda species schmitzi   Figs 6–7, 43A, 50A     Filistata schmitzi  Kulczyński, 1911: 13, pl. 1, fig. 3 ( ♂ ♀).    Zaitunia schmitzi–  Lehtinen 1967: 275, fig. 21 (♂♀). —  Zonstein 2009b: 127, figs 1–12 (♂♀).    Diagnosis Males differ from those of all other congeners by a long and large cigar-shaped tegulum, a very thick spermophore occupying the whole bulb, a strongly slanting tip of the cymbium and by the extremely short and sideward-directed embolus (vs smaller coniform and longer forward-directed ones in other species). Females are easily distinguished by their intense dark coloration (darker than that in any other species) as well as their low and small globular receptacles, with outer lobe only slightly exceeding the median one in size ( Fig. 7E–K).    Type material  Lectotype(designated by Zonstein 2009b: 127) ISRAEL: ♂, Jerusalem (the current depository unknown; not examined).   Paralectotype ISRAEL: 1 ♀, Galilee (the current depository unknown; not examined).    Note  Lehtinen (1967)indicated the typeseries as being kept in IZW, but according to Wioletta Wawer, the curator, these typeswere not found there (pers. comm.).   Material examined(47 ♂♂, 28 ♀♀, 8 juvs)   EGYPT: 1 ♀, South Sinai, Mt. Catarina, c. 28°33' N, 33°57' E,  16 Jun. 1968, unknown collector ( HUJ).   ISRAEL: 1 ♀, Upper Galilee, Mt. Meron, 33°01' N, 35°23' E,  900 m,  4–5 Feb. 2007, T. Levanony( TAU);  1 ♂, same data, but  5–12 Apr. 2007( TAU);  1 ♂, same data, but  12–16 May 2007( TAU);  2 ♂♂, WesternGalilee, ‘ EnhaMifraz, 32°54' N, 35°05' E,  19 Mar. 1944, A. Shulov( HUJ);  1 ♂, same region, ‘ En Ya’aqov, 33°00' N, 35°14' E,  26–31 Jul. 2007, I. Shtirberg( TAU);  1 ♂, Carmel Ridge, Har Horshan, 32°35' N, 35°00' E, near Bat-Shlomo,  28 May 1993, Y. Lubin( HUJ);  2 ♀♀, same ridge, Elyaqim, 32°38' N, 35°04' E,  15 Mar. 1963, M. Warburg( TAU);  1 ♀, JordanValley, Ein Duyuk,  11 Apr. 1968, M. Pener( HUJ);  1 ♀, same locality,  10 Jun. 1970, M. Pener( HUJ);  1 juv., Yizre’el Valley, Bet HaShita, 32°33' N, 35°26' E,  6 Feb. 1937, A. Shulov( TAU); 2 juvs, same region, ‘ En Harod, 32°34' N, 35°23' E,  11 Mar. 1945, A. Shulov( HUJ);  1 ♀, Jerusalem,  16 Jul. 1941, A. Shulov( HUJ);  1 juv., same locality,  12 Dec. 1949, A. Shulov( HUJ);  1 ♀, same locality,  10 Mar. 1961, A. Shulov( HUJ);  1 ♀, same area, Ramat Rahel, 31°44' N, 35°13' E,  23 Apr. 1973, unknown collector ( HUJ);  4 ♂♂, 1 juv., Judean Hills, Sansan Reserve, 31°42' N, 35°05' E,  500–700 m,  22 Mar. 2002, Y. Mandelik( HUJ);  6 ♂♂, same data, but  31 Mar. 2001( HUJ);  1 ♂, same data, but  14 May 2002( HUJ);  7 ♂♂, same area, Ramat Avishur, 31°39' N, 34°55' E,  300–400 m,  22 Feb. 2001, Y. Mandelik( HUJ);  3 ♂♂, same area, Giv’at Zekharya, 31°42' N, 34°57' E,  250–300 m,  22 Mar. 2002, Y. Mandelik( HUJ);  1 ♂, same locality, but  14 May 2002( HUJ);  8 ♂♂, 2 ♀♀, 1 juv., same region, National Park Adullam,  1–30 Apr. 2003, U. Columbus& T. Levanony( TAU);  1 ♀, same region, Bet Shemesh, 31°45' N, 34°59' E,  3 Mar. 1954, A. Weissman( HUJ);  1 ♀, same area, 6 kmSW of Bet Shemesh, 31°40' N, 34°57' E,  350 m,  10 May 2009, S. Zonstein( TAU);  1 ♀, same region, 3.5 kmE of Bet Guvrin, 31°37' N, 34°56' E,  350 m,  8 Nov. 2011, S. Zonstein( TAU);  2 ♂♂, same area, near Giv’at Yeshayahu, 31°40' N, 34°57' E,  300–350 m,  20 May 2007, O. Skutelsky( TAU);  1 ♂, same data, but  20 May 2008( TAU);  1 ♂, same area, Matta, 31°43' N, 35°04' E,  620 m,  13–18 Apr. 2007, I. Shtirberg( TAU);  1 ♂, same region, near Lahav, 31°33' N, 34°52' E,  500 m,  25–30 Jul. 2006, I. Shtirberg( TAU);  1 ♂, 2 ♀♀, same data, but  13–18 Jan. 2007( TAU);  2 ♀♀, NorthernNegev, Dorot – Ruhama, c. 31°30' N, 34°41' E,  31 Jan. 1972, I. Vaisberg( HUJ);  1 ♂, same region, HaBesor, 31°18' N, 34°29' E,  70 m,  2 Apr. 2010, C. Drees& L. Friedman( TAU);  1 ♀, Dead Sea Area, ‘ Enot Zuqim(En Fashkha),  24 Feb. 1942, A. Shulov( HUJ);  1 ♂, same region, Nahal Perat(Wadi Kelt),  13 Jan. 1945, A. Shulov( HUJ);  1 ♂, 3 ♀♀, same region, surroundings of ‘ En Gedi, 31°27' N, 35°23' E, 0 m,  19 Jan. 2013, S. Zonstein( TAU);  2 ♂♂, 1 juv., Judean Desert, ‘ Arad, 31°16' N, 35°13' E,  20 Dec. 1962, P. Amitai( HUJ);  4 ♀♀, 1 juv., same locality,  13 Feb. 1969, M. Warburg( TAU);  1 ♀, Ma’ale Adummim,  30 Apr. 1972, unknown collector ( HUJ);  1 ♀, CentralNegev, Yeroham, 30°59' N, 34°56' E,  30 Nov. 1964, Blond& Watz( HUJ);  1 juv., same region, crater Makhtesh Gadol, c. 30°57' N, 35°00' E,  1 Apr. 1962, A. Shulov( HUJ).    Description  Male(from Jerusalem area) HABITUS. See Fig. 6B. BODY LENGTH. 2.83. COLOUR. Carapace dark yellowish-brown with margins, postocular area and eye tubercle dark brown to brownish-black; chelicerae, labium, sternum and maxillae brown; legs brown with diffuse and poorly developed dark-brown fasciae; abdomen uniformly dark brown. CARAPACE ( Fig. 6E). 1.27 long, 1.03 wide. EYES. AME 0.06, ALE 0.11, PLE 0.08, PME 0.06, AME–AME 0.06. PALP ( Figs 7A-D, 43A). Femur equal in length to tibia; tibia thick, uniformly swollen. Cymbium moderately long, with or without small dorsodistal projection. Broad, blunt-tipped tegulum with thick spermophore forming 4 complete coils closely adjoining each other. Embolus short, lacking a neck, flattened and bent retrolaterally. LEG MEASUREMENTS. ♂(♀)    Femur Patella Tibia Metatarsus Tarsus Total  Palp 0.61 (0.80) 0.25 (0.31) 0.59 (0.43) ― 0.24 (0.79) 1.69 (2.33)  I 1.31 (1.52) 0.49 (0.49) 1.28 (1.41) 1.10 (1.20) 0.73 (0.91) 4.91 (5.53)  II 1.04 (1.15) 0.47 (0.46) 0.98 (0.95) 0.93 (0.85) 0.65 (0.74) 4.07 (4.15)  III 0.98 (0.97) 0.46 (0.46) 0.78 (0.59) 0.95 (0.74) 0.64 (0.65) 3.81 (3.41)  IV 1.25 (1.36) 0.52 (0.48) 1.16 (1.05) 1.26 (1.06) 0.74 (0.73) 4.93 (4.68)  Female(from Jerusalem area) HABITUS. See Figs 6A, 50A. BODY LENGTH. 3.85. COLOUR. As in male, but carapace, palps and legs slightly lighter; in contrast, fasciae on palps and legs somewhat darker than in male. CARAPACE ( Fig. 6D). 1.52 long, 1.18 wide. EYES. AME 0.07, ALE 0.12, PLE 0.10, PME 0.07, AME–AME 0.06. ENDOGYNE ( Fig. 7E–K). With 2 pairs of globular receptacles. Median receptacles two times thinner that lateral; median receptacles separated by 3 diameters; lateral receptacles evenly covered by pores, and median receptacles with pores on head only.    Variation Carapace length in males varies from 1.03 to 1.30, in females from 1.37 to 2.00; ground colouration varies from medium yellowish-brown with a darker brown abdomen to an almost uniform dark brown.    Ecology The species inhabits many typesof habitats, from deserts through steppes and maquis to low oak evergreen forest; in all these habitats, it occurs mainly under rocks (see Fig. 50A).    Distribution  Egypt( Sinai) and Israel( Fig. 46). 1291545687 [189,1399,460,487] 1968-06-16 HUJ Egypt 28.55 Mt. Catarina 1232 33.95 17 18 1 1 South Sinai 1291545704 2007-02-04 2007-02-05 2007-02-04 TAU T. Levanony Israel 900 33.016666 Mt. Meron 1207 35.383335 Upper Galilee 17 18 1 1 1291545677 [286,822,561,588] 2007-04-05 2007-04-12 2007-04-05 TAU T. Levanony Israel 900 33.016666 Mt. Meron 1207 35.383335 Upper Galilee 17 18 1 1 1291545709 [838,1394,561,588] 2007-05-12 2007-05-16 2007-05-12 TAU T. Levanony Israel 900 33.016666 Mt. Meron 1207 35.383335 Upper Galilee 17 18 1 1 1291545690 [189,1267,596,623] 1944-03-19 HUJ A. Shulov Israel 32.9 En 1208 35.083332 17 18 2 2 Western 1291545692 2007-07-26 2007-07-31 2007-07-26 TAU I. Shtirberg Israel 33.0 En Ya'aqov 1207 35.233334 17 18 1 1 Western 1291545679 1993-05-28 HUJ Y. Lubin Israel Carmel Ridge 32.583332 Bat-Shlomo 1209 35.0 Har Horshan 17 18 1 1 Western 1291545697 1963-03-15 TAU M. Warburg Israel 32.633335 Elyaqim 1209 35.066666 17 18 2 2 Western 1291545695 1968-04-11 HUJ M. Pener Jordan Ein Duyuk 17 18 1 1 1291545717 [472,1071,737,764] 1970-06-10 HUJ M. Pener Jordan Ein Duyuk 17 18 1 1 1291545743 1937-02-06 TAU A. Shulov Jordan 32.55 Bet HaShita 1210 35.433334 Yizre'el Valley 17 18 1 1291545703 1945-03-11 HUJ A. Shulov Jordan 32.566666 En Harod 1210 35.383335 17 18 1 1291545670 [692,1246,808,834] 1941-07-16 HUJ A. Shulov Jordan Jerusalem 17 18 1 1 Jerusalem 1291545693 1949-12-12 HUJ A. Shulov Jordan Jerusalem 17 18 1 Jerusalem 1291545723 [698,1329,843,869] 1961-03-10 HUJ A. Shulov Jordan Jerusalem 17 18 1 1 Jerusalem 1291545710 1973-04-23 HUJ Jordan 31.733334 Ramat Rahel 1214 35.216667 17 18 1 1 Jerusalem 1291545685 2002-03-22 HUJ Y. Mandelik Jordan 600 31.7 Sansan Reserve 1214 35.083332 Judean Hills 17 18 5 4 Jerusalem 1291545689 [189,690,948,975] 2001-03-31 HUJ Y. Mandelik Jordan 600 31.7 Sansan Reserve 1214 35.083332 Judean Hills 17 18 6 6 Jerusalem 1291545678 [702,1181,948,975] 2002-05-14 HUJ Y. Mandelik Jordan 600 31.7 Sansan Reserve 1214 35.083332 Judean Hills 17 18 1 1 Jerusalem 1291545707 2001-02-22 HUJ Y. Mandelik Jordan 350 31.65 Ramat Avishur 1214 34.916668 17 18 7 7 Jerusalem 1291545719 2002-03-22 HUJ Y. Mandelik Jordan 275 31.7 Giv'at Zekharya 1214 34.95 17 18 3 3 Jerusalem 1291545715 2002-05-14 HUJ Y. Mandelik Jordan 275 31.7 Giv'at Zekharya 1214 34.95 17 18 1 1 Jerusalem 1291545713 2003-04-01 2003-04-30 2003-04-01 TAU U. Columbus & T. Levanony Jordan National Park Adullam 17 18 11 2 8 Jerusalem 1291545734 1954-03-03 HUJ A. Weissman Jordan 31.75 Bet Shemesh 1214 34.983334 17 18 1 1 Jerusalem 1291545726 2009-05-10 TAU S. Zonstein Jordan 350 31.666666 Bet Shemesh 1214 34.95 17 18 1 1 Jerusalem 1291545701 2011-11-08 TAU S. Zonstein Jordan 350 31.616667 Bet Guvrin 1215 34.933334 17 18 1 1 Jerusalem 1291545732 2007-05-20 TAU O. Skutelsky Jordan 325 31.666666 Giv'at Yeshayahu 1214 34.95 17 18 2 2 Jerusalem 1291545682 [571,1043,1230,1257] 2008-05-20 TAU O. Skutelsky Jordan 325 31.666666 Giv'at Yeshayahu 1214 34.95 17 18 1 1 Jerusalem 1291545700 2007-04-13 2007-04-18 2007-04-13 TAU I. Shtirberg Jordan 620 31.716667 Matta 1214 35.066666 17 18 1 1 Jerusalem 1291545702 2006-07-25 2006-07-30 2006-07-25 TAU I. Shtirberg Jordan 500 31.55 Lahav 1215 34.866665 17 18 1 1 Jerusalem 1291545749 2007-01-13 2007-01-18 2007-01-13 TAU I. Shtirberg Jordan 500 31.55 Lahav 1215 34.866665 17 18 3 2 1 Jerusalem 1291545686 [281,1394,1336,1362] 1972-01-31 HUJ I. Vaisberg Jordan 31.5 Dorot - Ruhama 1216 34.683334 17 18 2 2 Northern 1291545718 [189,1371,1371,1397] 2010-04-02 TAU C. Drees & L. Friedman Jordan 70 31.3 HaBesor 1216 34.483334 17 18 1 1 Northern 1291545727 1942-02-24 HUJ A. Shulov Jordan Enot Zuqim Dead Sea Area 17 18 1 1 Northern 1291545714 1945-01-13 HUJ A. Shulov Jordan Nahal Perat 17 18 1 1 Northern 1291545741 2013-01-19 TAU S. Zonstein Jordan 31.45 En Gedi 1216 35.383335 17 18 4 3 1 Northern 1291545733 1962-12-20 HUJ P. Amitai Jordan 31.266666 Arad 1217 35.216667 Judean Desert 17 18 3 2 Northern 1291545711 1969-02-13 TAU M. Warburg Jordan 31.266666 Arad 1217 35.216667 Judean Desert 17 18 5 4 Northern 1291545712 [281,1036,1547,1573] 1972-04-30 HUJ Jordan Ma'ale Adummim 17 18 1 1 Northern 1291545725 1964-11-30 HUJ Blond & Watz Jordan 30.983334 Yeroham 1218 34.933334 17 18 1 1 Central 1291545735 1962-04-01 HUJ A. Shulov Jordan 30.95 Makhtesh Gadol 1218 35.0 17 18 1 Central