The beetle family Carabidae of Costa Rica: The genera of the Cryptobatida group of subtribe Agrina, tribe Lebiini, with new species and notes on their way of life (Insecta: Coleoptera) Erwin, Terry L. Zootaxa 2004 662 1 54 8KTVP [1223,1316,1518,1544] Insecta Carabidae Animalia Coleoptera 5 6 Arthropoda tribe Lebiini    1 Elytron at basal third depressed, surface uneven, tuberculate or not, and/or margin of pronotum angulate or subangulate at mid­lateral setiferous pore ............................... .......................................................................................... Cryptobatida Group... 2  1' Elytron neither depressed nor tuberculate, surface smooth with normal interneurs and intervals (or interneurs effaced); side of pronotum evenly rounded .................... .......................................... Plochionida Group, Calleidida Group, Agrida Group   2 Elytron markedly tuberculate overall or with a series of small setiferous tubercles in intervals 3 and 5 ....................................................................................................... 3 2' Elytron without trace of discal tubercles; lateral marginal intervals with or without calli ................................................................................................................................... 5   3(2) Prothorax somewhat tubular, much narrower than head ...  Otoglossa Chaudoir 1872  3' Prothorax wider than head ...................................................................................... 4   4(3') Sides of pronotum narrowly reflexed except at mid­lateral seta, there wide; neck well­defined, narrow; elytra metallic blue, forebody rufous....1  Thoasia Liebke 1939  4' Sides of pronotum broadly reflexed throughout; neck broad ...................................... ...........................................................................................  Hyboptera Chaudoir 1872   5(2') Antennomere 4 glabrous except for apical ring setae .............................................. 6  5' Antennomere 4 multisetiferous from basal third to apex, in addition to apical ring setae ................................................................................................................................... 7   6(5) Elytron laterally with callus at apical third; male endophallus without flagellum ..... ........................................................................................  ValeriaascheroErwin 2004  6' Elytron laterally without callus at apical third; male endophallus with flagellum ...... ...........................................................................................  Aspasiola Chaudoir 1877   7(5') Head and pronotum densely and evenly punctate .......  Cylindronotum Putzeys 1846  7' Head and pronotum smooth ...................................................................................... 8   8 (7') Pronotum with lateral margin narrowly reflexed from base to apex .......................... ......................................................................................  Pseudotoglossa Mateu 1961  8' Pronotum with lateral margin moderately or markedly reflexed from base to apex 9   9(8') Elytron laterally at apical third with large callus .......  Cryptobatis Eschscholtz 1829  9' Elytron laterally at apical third without callus ........................  Onota Chaudoir 1872