Alexander, 1958 : 49 Alexander, Alexander 1965 : 98 Alexander, Alexander 1967 : 171 A revision of Bittacomorphinae with a review of the monophyly of extant subfamilies of Ptychopteridae (Diptera) Fasbender, Andrew Courtney, Gregory W. Zootaxa 2017 4309 1 1 69 5WKFT Alexander 1958 Alexander 1958 [151,668,1819,1845] Insecta Ptychopteridae Bittacomorphella Animalia Diptera 39 40 Arthropoda species pacifica     Bittacomorphella pacifica  Alexander, 1958: 49–50 (original description)  Bittacomorphella pacifica  Alexander, Alexander 1965: 98(catalog)    Bittacomorphella pacifica  Alexander, Alexander 1967: 171(illustration) Diagnosis. Adult( Figs. 7c, 12). Legs with white confined to distal tip of 1st, 2nd & 3rd tarsomeres., paramere straight, subaedeagal sclerite of aedeagus with dorsal tooth.   Description.Larvae and pupae unknown. Adult.Measurements (n=5). Overall length 10.45mm( 9.68– 11.16mm); head width (n=3): 0.85mm( 0.82–0.89mm), height, 0.81mm( 0.74–0.87mm), length 0.45mm( 0.38– 0.53mm); antenna flagellum length 6.78mm, flagellomere proportions: 1.00, 0.70, 0.70, 0.70, 0.70, 0.70, 0.67, 0.67, 0.56, 0.56, 0.51, 0.48, 0.43, 0.43, 0.43, 0.40, 0.40, 0.35, 0.35, 0.08; maxillary palpus length 1.95mm; palpus segment proportions: 0.28, 1, 1.52, 0.96, 3.68; thorax length 1.49mm( 1.31–1.67mm), height: 1.42mm( 1.34– 1.55mm); wing length 7.38mm( 6.70–8.30mm), width (n=4): 1.59mm( 1.41–1.85mm); R3+4+5 stem 0.54mmstraight, R4+5 stem 1.55mm, R4 after fork 1.16mm, R5 after fork 1.13mm, R3+4+5:R4+5-stem:R4 1.00, 3.03, 2.3, r-m 0.25mm, M distal to r-m 2.50mm, Mdistal:R3+4+5 5.13:1.00, R4:Mdistal 0.06, 1.00, CuA2 1.08mm; leg proportions, femur 1.0, 1.01, 1.04; tibia 0.96, 0.94, 1.04; tarsomeres, foreleg 0.53, 0.18, 0.13, 0.03, 0.03; midleg 0.56, 0.20, 0.14, 0.03, 0.03; hind leg 0.68, 0.23, 0.15, 0.03, 0.03; abdomen length: 8.81mm(8.00– 9.96mm), Male tergite proportions 1.00, 3.37, 1.98, 1.87, 1.53, 0.78, 0.14; male genitalia width: 0.76mm, height 0.71mm. Frons medial suture indistinct, most prominent between antennal bases. Antennal scape with three setiform sensilla midway along anterior margin; 20 flagellomeres. Face narrow, incised by clypeus nearly to dorsal margin. Hypopharynx subconical triangular, directed anterior-ventrally. Labium extending beyond clypeus. Labella partially fused medially. Antepronotum dorsal margin rounded. Scutum glabrous, pollinose, fuscous. Scutellum muddy yellow, prescutellum absent. Mediotergite ruddy-brown. Paratergite poorly developed, narrow, scythe-like. Sub-paratergal membrane subquadrate, lightly sclerotized, anepisternal suture absent. Pleurites ruddy brown. Wing membrane with scattered macrotrichia on distal 1/10 of wing, in cells R1+2, R3, R4, R5, M; veins C to wing apex, Sc, R s, R1, R3, R4 & R5 after forking and distal portion of M with macrotrichia; alula undeveloped. Posthalter poorly developed protrusion, free of webbing. Coxae yellow, overall leg coloration brown. White banding confined to distal tip of 1st, 2nd & 3rd tarsomeres. Empodium notched proximally. Abdominal tergites and sternites ruddy brown. Male.Epandrium about 1.5x as wide as long; weakly bowed dorsally, rarely with weak internal suture present, posterior corner subobtuse, internal fold arcing weakly anteriorly, ending near medial margin of posteriomedial style; posterior margin emarginate across 1/2 of medial margin, posteromedial style blunt triangular lobule. Epandrial claspers evenly tapering along length to sub-acute point, curved sub-circularly dorsoanteriorly. Parameres glabrous, dorsoventrally flattened basally; distal portion dorsoposteriorly directed acute straight spine tapering to fine point. Gonocoxite conical, expanding to gonostylus. Gonostylus 1.5x length of gonocoxite, moderately expanded at base, inner apex sensilla single row of 10–11 along distal-medial surface, arising from dorsal margin anteriorly and shifting ventrally posteriorly; three additional sensilla on medial surface near gonocoxite. Basal division of hypandrium with medial suture present to lacrimiform membranous region; terminal division wellarticulated, proximal portion dorsally cupped pouch longer than wide; apical portion lateral margins fused to narrow tube, width narrower than proximal portion, end closed. Aedeagus with sperm sac as described for subfamily, except lateral membranous windows highly reduced. Lateral ejaculatory process broad and triangularly winglike, swept-anterior, apex rounded, dorsolaterally compressed at base, rotating to completely laterally compressed apex, low longitudinal ridge present on posterior surface emerging from ventral section; ventral lobe small, rounded, not incising sperm sac. Ejaculatory apodeme teardrop shaped, expanding to rounded apex. Aedeagal sclerites not tapered. Apical apodemes simple, dorsoanterior extension of subaedeagal sclerite articulating only on posterior margin. Subaedeagal sclerite ventral arm triangular, apex directed anterior; external portion narrow tapering, curved dorsally towards apex, dorsal surface with broadly triangular denticle immediately prior to apex. Female.Epigynium trapezoidal, flat dorsally, posterior margin straight, lateral margins straight, posterior margin indistinct, medial suture indistinct. Cerci articulation indistinct, dorsal margin weakly convex, apex broadly rounded, ventral margin slightly convex, with an even arc; apical cylindrical nodule with single elongate trichoid sensilla medially; cercus epigynium ratio 0.5, 1. Hypoproct trapezoidal, narrowing posteriorly, base with heavily sclerotized subtriangular lobe directed posteriorly. Genital fork T-shaped, lateral apices weakly curved anteriorly. Hypogynium quadrate, posterior corners triangular lobes, hypogynial valves not distinguishable, medial notch absent.     Typematerial examined. Holotype. UNITED STATES. WASHINGTON.  Pierce Co. Mt.Rainier, Wash.  4700 ft.,  Jul. 21 1940, H.&M. Townes, Lowerspec.  ♂= Holotype[pinned 1 ♂, head and genitalia on slide; USNM].  Paratypes.Same locality as holotype[pinned 1 ♀, same pin as holotype; USNM)]. CALIFORNIA.    Siskiyou Co.Castle Crags,  6.vii.1953, coll. C.P. Alexander[pinned 2 ♂on same pin; slide 1 ♂; USNM].  OREGON.  Yamhill Co.Dayton,  19.ix.1946, coll. K.M. Fender[pinned 1 ♂; USNM].    Othermaterial examined. CANADA. BRITISH COLUMBIA. Robson,  5.vi.1938, coll. H.H. Foxlee[pinned 1 ♂, 1 ♀; UMMZ].   UNITED STATES. OREGON.  Benton Co.Mid Parker Creek,  8.viii.1981, coll. G.W. Courtney[alcohol 1 ♂; ISIC];   Clackamas Co.Mt.Hood,  29.vii.1921, coll. A.L. Melander[pinned 1 ♂; USNM];   Douglas Co. 3 mi. E. Reedsport, Schofield Slough,  23.iv.1964, coll. J.D. Vertrees, J. Schuh[pinned 1 ♂; WSU].  WASHINGTON.  Snohomish Co.Everett,  4.vii.1924, coll. A.L. Melander[pinned 2 ♂; USNM];   Thurston Co.Olympia,  10.vi.1891, no collector [pinned, 1 ♀; WSU].  Taxonomic notes.  Bittacomorphella pacificacan be distinguished by the combination of a pollinose scutum, straight, needle-like parameres and a dorsal tooth-like projection on the subaedeagal sclerite of the aedeagus. Many specimens determined as  Bl. sackeniiare actually this species.   Distribution. Fig. 19a. 1586248089 1940-07-21 Co. Mt. Rainier & H. & M. Townes & Lower United States of America 1433 Mt. Rainier Pierce Co. Mt. 41 42 2 WASHINGTON holotype 1586248092 USNM United States of America CALIFORNIA 41 42 3 1 2 CALIFORNIA holotype 1586248094 [151,1433,151,176] 1953-07-06 USNM Co. Castle Crags & C. P. Alexander Siskiyou Co. Siskiyou Co. 42 43 3 3 1586248087 [151,1100,188,213] 1946-09-19 USNM Co. Dayton & K. M. Fender Yamhill Co. Yamhill Co. 42 43 1 1 OREGON 1586248083 1938-06-05 UMMZ H. H. Foxlee Canada Robson Other 42 43 2 1 1 BRITISH COLUMBIA 1586248088 1981-08-08 ISIC G. W. Courtney United States of America Benton Co. Mid Parker Creek 42 43 1 1 OREGON 1586248105 [391,1328,295,320] 1921-07-29 USNM A. L. Melander United States of America Clackamas Co. Mt. 42 43 1 1 OREGON 1586248097 1964-04-23 WSU J. D. Vertrees & J. Schuh United States of America Douglas Co. Reedsport 42 43 1 1 Slough 1586248103 [151,1269,367,392] 1924-07-04 USNM A. L. Melander United States of America Snohomish Co. Snohomish Co. 42 43 2 2 WASHINGTON 1586248095 1891-06-10 WSU United States of America Thurston Co. Thurston Co. 42 43 1 1 WASHINGTON