Forel, 1886b : 184
Camponotus (Myrmocamelus)
Forel, 1914 : 270
Camponotus (Myrmosaga)
Wheeler, 1922 : 1045
Camponotus (Mayria)
Emery, 1925 : 122
Bolton, 1995 : 92
Taxonomic revision of the Malagasy Camponotus subgenus Mayria (Hymenoptera, Formicidae) using qualitative and quantitative morphology
Rakotonirina, Jean Claude
Fisher, Brian L.
Zootaxa
2018
2018-06-20
4438
1
1
58
QDM4
Forel
Forel
[151,461,1651,1677]
Insecta
Formicidae
Camponotus
GBIF
Animalia
Hymenoptera
18
19
Arthropoda
species
christi
Camponotus christi Forel, 1886b: 184. Lectotype minor worker, present designation, Centre de Madagascar [Ambatomanjaka, Miarinarivo, -18.766947, 46.869107, 1344 m] (Hildebrandt), AntWeb CASENT0101544 (MHNG). Paralectotype: 1 minor worker with same data as lectotype but specimen coded: CASENT0101435 (Camboué) (MNHN) [examined]. [Combination in Camponotus (Myrmocamelus): Forel, 1914: 270; in Camponotus (Myrmosaga): Wheeler, 1922: 1045; in Camponotus (Mayria): Emery, 1925: 122; Bolton, 1995: 92, 131].
Diagnosis.Median portion of clypeus without longitudinal carina; mandible with six teeth; anteromedian margin of clypeus not bordered by a lamella; in lateral view, mesosoma long and low, its dorsal outline not a dome-like structure; propodeal declivity inclined anteriorly; body color entirely yellow to light brown, legs with same color as body.
Description. Minor worker.In full-face view, head elongate (CWb/CL: 0.76±0.03; 0.69–0.83), lateral margins straight and more or less parallel to each other, rounding broadly to the slightly, medially excised posterior margin. Anterior clypeal margin straight, with broadly angulate junction to lateral margin; median longitudinal carina absent. Eyes not breaking lateral outlines of head; their posterior margin located approximately at posterior third portion of head (PoOc/CL: 0.34±0.01; 0.31–0.36). Mandible triangular, apical margin with six sharp teeth. Antennal scape long, more than its distal half extending beyond posterior cephalic border. In lateral view, pronotum and anterior half of mesonotum forming feeble convexity, posterior half of mesonotum and propodeal dorsum straight and long, joining declivity surface with noticeable angle. Propodeal dorsum roughly three times as long as height of declivity; propodeal spiracle circular. Petiolar node as high as or higher than long, its dorsal margin rounding to anterior face and joining posterior face at an angle. Dorsum of head covered with numerous yellowish, elongate, erect hairs; pronotum and mesonotum each with a pair of erect hairs, propodeum with a pair of hairs found on junction to declivity; sparse, short pubescence present on dorsum of body. Body color entirely yellowish-orange to orange-brown and funicular segments becoming dark brown toward the apex. Major worker.Differing from minor worker in the following characters: enlarged head, with markedly concave posterior margin; apical third of antennal scape surpassing posterior cephalic margin; robust mesosoma, metanotum distinctly visible, propodeal dorsum rounding to declivity and twice as long as height of declivity; petiolar node tapering dorsally. More pairs of erect hairs on junction of propodeal dorsum and declivity and on posterodorsal margin of petiolar node.
Discussion.Workers of C. christiare similar to those of C. mainty, C. foersteriand C. maculiventrisin that their clypeus lacks a median longitudinal carina, and in lateral view, their mesosoma is long and low. However, C. christiworkers are easily separable by their entirely yellow to light brown body color and legs. In the members of C. mainty, their body is entirely black to reddish-black. In the two latter species, at least the trochanters and the distal portion of the coxa are yellowish to white. The cluster of C. christishown by the dendrogram of NC-PART clustering is confirmed by the cumulative LDA at 100% identification success, indicating the support of the species hypothesized by qualitative morphologybased taxonomy. The cluster for C. christiis located next to that of C. foersteriand C. mainty, which suggests that the three species are more similar to each other than other species. This finding corroborates their close morphological similarity as detected by the qualitative analysis.
Distribution and biology. Camponotus christiis widely distributed across Madagascar. It occupies sclerophyll forests of the central high plateau, tropical dry forests in the west to coastal scrub and littoral rainforests, as well as montane rainforests in the east. While its workers have been found foraging on the forest floor and the lower portions of vegetation, their nest sites have been located in root mats in the ground layer, rotten logs, rotting tree stumps, rotten pockets above the ground, dead branches and bamboo above the ground, and in canopy moss and leaf litter.
Additional material examined.‘Andrangoloaka], (Sikora) (MHNG); [antananarivo, Museum Paris, Grandidier 1893], (Camboué) (MNHN); [Divers, tous les Divers sont de la Forêt d’Andrangoloaka, 20, Museum Paris, Grandidier 1899] (MNHN); [ Forêt d’Analamanita Province Antananarivo], ( Camboué) ( MHNG); 3 km41° NE Andranomay, 11.5 km147° SSE Anjozorobe, 1300 m, -18.47333, 47.96, montane rainforest, (Fisher, Griswold et al.) (CASC); Analamanga Region, District of Ankazobe, Ambohitantely, 46 kmNE of Ankazobe, 701 m, - 18.198, 47.2815, sclerophyll forest, (Mike, Rinha) ( CASC); Forêtde galerie, Andranorovitra, 24.0 km NNE Ankazobe, 1491 m, -18.11243, 47.19757, disturbed gallery montane forest, ( B.L. Fisher et al.) ( CASC); Mandraka, 1312 m, -18.91813, 47.91717, montane rainforest, ( B.L. Fisher et al.) ( CASC); Mandraka Park, 1360 m, -18.9019, 47.90786, montane shrubland, ( B.L. Fisher et al.) ( CASC); Réserve Spéciale d'Ambohitantely, Forêt d Ambohitantely, 20.9 km72° NE d Ankazobe, 1410 m, -18.22528, 47.28683, montane rainforest, (Fisher, Griswold et al.) ( CASC); Réserve Spéciale d'Ambohitantely, 1580 m, -18.18762, 47.28576, montane forest, ( B.L. Fisher et al.) ( CASC); Réserve Spéciale d'Ambohitantely, 1490 m, -18.22444, 47.2774, montane forest, ( B.L. Fisher et al.) ( CASC); Tananarive, (Waterlot) (MNHN); Province Antsiranana:[ Diego Suarez], ( Ch. Alluaud) ( MHNG); 11.0 km WSW Befingotra, Réserve Anjanaharibe-Sud, 1565 m, -14.75, 49.45, montane rainforest, (B.L. Fisher) (CASC); 6.5 kmSSW Befingotra, Réserve Anjanaharibe-Sud, 875 m, -14.75, 49.5, rainforest, ( B.L. Fisher) ( CASC); 9.2 kmWSW Befingotra, Réserve Anjanaharibe-Sud, 1280 m, -14.75, 49.46667, montane rainforest, ( B.L. Fisher) ( CASC); 9.2 kmWSW Befingotra, Réserve Anjanaharibe-Sud, 1200 m, -14.75, 49.46667, montane rainforest, ( B.L. Fisher) ( CASC); Ampasindava, Andranomatavy Forest, 543 m, -13.66296, 47.97936, disturbed dry forest, ( B.L. Fisher et al.) ( CASC); Ampasindava, Forêt d'Ambilanivy, 3.9 km181° S Ambaliha, 600 m, - 13.79861, 48.16167, rainforest, (Fisher, Griswold et al.) ( CASC); AntsirananaII Pref: Antsahampano S.- Pref: Montagne d'Ambre. SiteMD1, 1049 m, -12.52765, 49.17235, in Commelinaregrowth on path next to degraded primary riparian rainforest, (D. Lees, R. Ranaivosolo & P. Razafindraibe) ( CASC); Betaolana Forest, along Bekona River, 880 m, -14.52996, 49.44039, rainforest, ( B.L. Fisher et al.) ( CASC); Forêt de Binara, 9.1 km233° SW Daraina, 725 m, -13.26333, 49.60333, rainforest, ( B.L. Fisher) ( CASC); Galokochain, Mont Galoko, 520 m, - 13.58487, 48.71818, rainforest, ( B.L. Fisher et al.) ( CASC); Galokochain, Mont Galoko, 980 m, -13.5888, 48.72864, montane forest, ( B.L. Fisher et al.) ( CASC); Galokochain, Mont Galoko, 1100 m, -13.59358, 48.73157, montane forest, ( B.L. Fisher et al.) ( CASC); Makirovana forest, 415 m, -14.17066, 49.95409, rainforest, ( B.L. Fisher et al.) ( CASC); Makirovana forest, 715 m, -14.16666, 49.95, rainforest, ( B.L. Fisher et al.) ( CASC); Makirovana forest, 390 m, -14.10295, 50.01984, rainforest, ( B.L. Fisher et al.) ( CASC); Makirovana forest, 715 m, -14.16666, 49.95, rainforest, ( B.L. Fisher et al.) ( CASC); Montagne d'Ambre National Park, Roussette Camp 7 kmSW Parkentrance, 960 m, -12.51444, 49.18139, rainforest, (Mike, Frank, Rin'ha) ( CASC); Parc National de Marojejy, 25.7 km32° NNE Andapa, 10.3 km314° NW Manantenina, 1575 m, -14.445, 49.74167, montane rainforest, ( B.L. Fisher) ( CASC); Parc National de Marojejy, Antranohofa, 26.6 km31° NNE Andapa, 10.7 km318° NW Manantenina, 1325 m, -14.44333, 49.74333, montane rainforest, ( B.L. Fisher) ( CASC); Parc National de Marojejy, Manantenina River, 27.6 km35° NE Andapa, 9.6 km327° NNW Manantenina, 775 m, -14.435, 49.76, rainforest, ( B.L. Fisher et al.) ( CASC); Parc National Montagne d'Ambre, 1100 m, -12.52306, 49.17901, montane rainforest, ( B.L. Fisher et al.) ( CASC); Parc National Montagne d'Ambre, 3.6 km235° SW Joffreville, 925 m, - 12.53444, 49.1795, montane rainforest, (Fisher, Griswold et al.) ( CASC); Parc National Montagne d'Ambre, Mahasarika, 1135 m, -12.53176, 49.17662, montane rainforest, (B.L. Fisher et al.) (CASC); Parc National Montagne d'Ambre, Roussettes, 1025 m, -12.52574, 49.17238, montane rainforest, (B.L. Fisher et al.) (CASC); Réserve Spéciale Manongarivo, 12.8 km228° SW Antanambao, 780 m, -13.97667, 48.42333, rainforest, ( B.L. Fisher) ( CASC); SAVARegion, District of Sambava, Marojejy National Park, 5 kmW of Manantenina village, 1st Campsite ( Mantella), 487 m, -14.43817, 49.774, Lowaltitude rainforest, (Rin'ha, Mike) ( CASC); 3 kmW Sakalava Beach[white dunes site], 40 m, -12.28617, 49.36667, white dunes in littoral forest, (Harin'Hala, Irwin, Schlinger) ( CASC); Montaigne Français, 150 m, -12.325, 49.33333, along forested limestone ridge, ( R. Harin'Hala) ( CASC); Parc National Montagne d'Ambre[1 st campsite], 960 m, -12.51444, 49.18139, rainforest, ( R. Harin'Hala) ( CASC); Parc National Montagne d'Ambre[ Petit Lacroad], 1125 m, -12.52028, 49.17917, rainforest, ( R. Harin'Hala) ( CASC); Sakalava Beach[vegetated beach dunes], 10 m, -12.26278, 49.3975, across sandy trail in dwarf littoral forest, ( R. Harin'Hala) ( CASC); Province Fianarantsoa:[ Imerina], ( Camboué) ( MHNG); [ Nosibe Imerina], ( Sikora) ( MHNG); 3 kmW Ranomafana, nr. Ifandiana, 950 m, -21.25, 47.41667, rainforest, ( P.S. Ward) ( PSWC); 38 kmS Ambalavao, Réserve Andringitra, 1680 m, -22.2, 46.96667, montane rainforest, ( B.L. Fisher) ( CASC); 40 kmS Ambalavao, Réserve Andringitra, 1275 m, -22.21667, 46.96667, montane rainforest, ( B.L. Fisher) ( CASC); 7.6 km122° Kianjavato, Forêt Classée Vatovavy, 175 m, -21.4, 47.94, rainforest, (B.L. Fisher et al.) (CASC); 8.0 km NE Ivohibe, 1200 m, -22.42167, 46.89833, montane rainforest, (B.L. Fisher & Sylvain) (CASC); 9.0 km NE Ivohibe, 900 m, -22.42667, 46.93833, rainforest, (B.L. Fisher & Sylvain) (CASC); Belle Vuetrail, Ranomafana National Park, FianarantsoaProv., 1020 m, -21.2665, 47.42017, mixed tropical forest, (M.E. Irwin, F.D. Parker, R. Harin'Hala) ( CASC); Fitovavy Fitovinany Region, District of Ifanadiana Bellevue area 1200 mS of Ranomafana National Parkentrance, 1018 m, -21.2665, 47.42017, rainforest, (Rin'ha, Mike) ( CASC); Fitovavy Fitovinany Region, District of Ifanadiana, 12 kmW of Ranomafana, 1127 m, -21.25083, 47.40717, forest edge, open area, (Rin'ha, Mike) ( CASC); Forêt d'Atsirakambiaty, 7.6 km285° WNW Itremo, 1550 m, -20.59333, 46.56333, montane rainforest, (Fisher, Griswold et al.) ( CASC); Forêt de Vevembe, 66.6 km293° Farafangana, 600 m, -22.791, 47.18183, rainforest, transition to montane forest, ( B.L. Fisher et al.) ( CASC); JIRAMA water works near river, Ranomafana National Park, FianarantsoaProv., 690 m, -21.2485, 47.45217, open area near stream, (M.E. Irwin, F.D. Parker, R. Harin'Hala) ( CASC); Réserve Spéciale Manombo, 32 kmSE of Farafangana, 36 m, -23.02183, 47.72, Lowlandrainforest, (Rin'ha, Mike) ( CASC); Miandritsara Forest, 40 kmS of Ambositra, 822 m, -20.79267, 47.17567, Lowaltitude rainforest, (Rin'ha, Mike) ( CASC); Parc National Befotaka-Midongy, Papango 27.7 kmS Midongy-Sud, Mount Papango, 940 m, -23.83517, 46.96367, rainforest, ( B.L. Fisher et al.) ( CASC); Parc National Befotaka-Midongy, Papango 28.5 kmS Midongy-Sud, Mount Papango, 1250 m, - 23.84083, 46.9575, montane rainforest, ( B.L. Fisher et al.) ( CASC); Parc National de Ranomafana, Vatoharanana River, 4.1 km231° SW Ranomafana, 1100 m, -21.29, 47.43333, montane rainforest, (Fisher, Griswold et al.) ( CASC); Réserve Spéciale Ivohibe8.0 km E Ivohibe, 1200 m, -22.48333, 46.96833, montane rainforest, ( B.L. Fisher& Sylvain) ( CASC); Réserve Spéciale Ivohibe, 7.5 kmENE Ivohibe, 900 m, -22.47, 46.96, rainforest, ( B.L. Fisher& Sylvain) ( CASC); Réserve Spéciale Manombo 24.5 km228° Farafangana, 30 m, -23.01583, 47.719, rainforest, ( B.L. Fisher et al.) ( CASC); radio tower, Ranomafana National Park, FianarantsoaProv., 1130 m, - 21.25083, 47.40717, forest edge, mixed tropical forest, open area, ( M. Irwin, R. Harin'Hala) ( CASC); Ranomafana National Park, Talatakely area, 0.4 kmWSW of Park Entrance, 900 m, -21.41667, 47.68333, mixed tropical forest, ( D. H. & K.M. Kavanaugh) ( CASC); stream area, 900 mE of Isalo National Park Interpretive Center, Fianarantsoa Prov., 750 m, -22.62667, 45.35817, open area near stream, (R. Harin'Hala) (CASC); Vohiparara, -21.23333, 47.36667, (A. Pauly) (CASC); Vohipararabroken bridge, FianarantsoaProv., 1110 m, -21.22617, 47.36983, high altitude rainforest, ( R. Harin'Hala) ( CASC); Province Mahajanga: Forêt de Tsimembo, 8.7 km336° NNW Soatana, 20 m, -19.02139, 44.44067, tropical dry forest, ( Fisher-Griswold Arthropod Team) ( CASC); Réserve Spéciale Marotandrano, Marotandrano 48.3 kmS Mandritsara, 865 m, -16.28322, 48.81443, transition humid forest, ( B.L. Fisher et al.) ( CASC); Region Sofia, Bemanevika, 1606 m, -14.337, 48.58874, montane rainforest, (B.L. Fisher et al.) (CASC); Region Sofia, Bemanevika, 1657 m, -14.32826, 48.58406, montane rainforest, (B.L. Fisher et al.) (CASC); Province Toamasina: Baie d’Antongil( Mocquerys) ( MSNG); 19 kmESE Maroantsetra, 250 m, -15.48333, 49.9, rainforest, (P.S. Ward) ( PSWC); 1 kmSSW Andasibe(= Perinet), 920 m, -18.93333, 48.41667, rainforest, ( P.S. Ward) ( PSWC); 5.3 kmSSE Ambanizana, Andranobe, 600 m, -15.66667, 49.96667, rainforest, ( B.L. Fisher) ( CASC); 5.3 kmSSE Ambanizana, Andranobe, 425 m, -15.67133, 49.97395, rainforest, ( B.L. Fisher) ( CASC); 6.3 kmS Ambanizana, Andranobe, 25 m, -15.6813, 49.958, rainforest, ( B.L. Fisher) ( CASC); 6.9 kmNE Ambanizana, Ambohitsitondroina, 825 m, -15.58506, 50.00952, rainforest, ( B.L. Fisher) ( CASC); 6.9 kmNE Ambanizana, Ambohitsitondroina, 1000 m, -15.56667, 50, montane rainforest, ( B.L. Fisher) ( CASC); 6.9 kmNE Ambanizana, Ambohitsitondroina, 1080 m, -15.56667, 50, montane rainforest, ( B.L. Fisher) ( CASC); 6 kmESE Andasibe(= Perinet), 900 m, -18.95, 48.46667, rainforest, ( P.S. Ward) ( PSWC); 7 SE Andasibe National Park Headquarters, 1050 m, -18.96278, 48.45267, tropical forest, ( M.E. Irwin, R. Harin'Hala) ( CASC); Ambalahasina, 62.4 km19° Toamasina, 15 m, -17.59452, 49.46785, coastal scrub, ( B. Blaimer, F.N. Raharimalala) ( CASC); Ambatovy, 12.4 kmNE Moramanga, 1010 m, -18.84963, 48.2947, montane rainforest, ( B.L. Fisher et al.) ( CASC); Ambatovy, 12.4 kmNE Moramanga, 1000 m, -18.84773, 48.29568, montane rainforest, ( B.L. Fisher et al.) ( CASC); Ambatovy, 12.4 kmNE Moramanga, 1080 m, -18.83937, 48.30842, montane rainforest, ( B.L. Fisher et al.) ( CASC); Analamay, 1068 m, -18.80623, 48.33707, montane rainforest, ( Malagasyant team) ( CASC); Andasibe National Park, botanic garden near entrance, west of ANGAPoffice, 1025 m, -18.92639, 48.40783, tropical forest, ( M.E. Irwin, R. Harin'Hala) ( CASC); Ankerana, 865 m, -18.40062, 48.81311, rainforest, ( B.L. Fisher et al.) ( CASC); Ankerana, 750 m, -18.40829, 48.82107, rainforest, ( B.L. Fisher et al.) ( CASC); Ankerana, 1035 m, -18.4017, 48.80605, montane forest, ( B.L. Fisher et al.) ( CASC); Bevolota 17.1 kmN Andasibe, 995 m, - 18.77071, 48.43164, montane rainforest, ( B.L. Fisher et al.) ( CASC); Corridor Forestier Analamay-Mantadia, Ambatoharanana, 1016 m, -18.79944, 48.40375, rainforest, (B.L. Fisher et al.) (CASC); Corridor Forestier Analamay-Mantadia, Ambatoharanana, 968 m, -18.80424, 48.40081, rainforest, (B.L. Fisher et al.) (CASC); Corridor Forestier Analamay-Mantadia, Ambatoharanana, 1013 m, -18.80388, 48.40506, rainforest, (B.L. Fisher et al.) (CASC); Corridor Forestier Analamay-Mantadia, Ambatoharanana, 968 m, -18.80424, 48.40081, rainforest, (B.L. Fisher et al.) (CASC); Corridor Forestier Analamay-Mantadia, Ambatoharanana, 1064 m, -18.80398, 48.40358, rainforest, (B.L. Fisher et al.) (CASC); Corridor Forestier Analamay-Mantadia, Ambatoharanana, 968 m, -18.80424, 48.40081, rainforest, (B.L. Fisher et al.) (CASC); Corridor Forestier Analamay-Mantadia, Ambatoharanana, 960 m, -18.80438, 48.40735, rainforest, (B.L. Fisher et al.) (CASC); Corridor Forestier Analamay-Mantadia, Ambohibolakely, 1044 m, -18.76087, 48.37128, rainforest, (B.L. Fisher et al.) (CASC); Corridor Forestier Analamay-Mantadia, Ambohibolakely, 983 m, -18.76131, 48.36437, rainforest, (B.L. Fisher et al.) (CASC); Corridor Forestier Analamay-Mantadia, Ambohibolakely, 918 m, -18.77898, 48.36375, rainforest, (B.L. Fisher et al.) (CASC); Corridor Forestier Analamay-Mantadia, Ambohibolakely, 983 m, -18.76131, 48.36437, rainforest, (B.L. Fisher et al.) (CASC); Corridor Forestier Analamay-Mantadia, Tsaravoniana, 984 m, - 18.76369, 48.4203, rainforest, (B.L. Fisher et al.) (CASC); Corridor Forestier Analamay-Mantadia, Tsaravoniana, 1039 m, -18.76465, 48.41938, rainforest, (B.L. Fisher et al.) (CASC); Corridor Forestier Analamay-Mantadia, Tsaravoniana, 1018 m, -18.75737, 48.42302, rainforest, (B.L. Fisher et al.) (CASC); F.C. Andriantantely, 530 m, - 18.695, 48.81333, rainforest, (H.J. Ratsirarson) (CASC); F.C. Didy, 960 m, -18.19833, 48.57833, rainforest, (H.J. Ratsirarson) (CASC); F.C. Sandranantitra, 450 m, -18.04833, 49.09167, rainforest, (H.J. Ratsirarson) (CASC); Forêt Ambatovy, 14.3 km57° Moramanga, 1075 m, -18.85083, 48.32, montane rainforest, ( Malagasyant team) ( CASC); Manakambahiny Atsinanana, -17.75, 48.71667, Primary forest, ( A. Pauly) ( CASC); Montagne d'Akirindro 7.6 km341° NNW Ambinanitelo, 600 m, -15.28833, 49.54833, rainforest, (Fisher, Griswold et al.) ( CASC); Montagne d'Anjanaharibe, 18.0 km 21° NNE Ambinanitelo, 470 m, -15.18833, 49.615, rainforest, (Fisher, Griswold et al.) ( CASC); Montagne d'Anjanaharibe, 19.5 km27° NNE Ambinanitelo, 1100 m, -15.17833, 49.635, montane rainforest, (Fisher, Griswold et al.) ( CASC); Nosy Mangabe, 300 m, -15.5, 49.76667, rainforest, ( P.S. Ward) ( PSWC); Nosy Mangabe, 5 m, -15.5, 49.76667, littoral vegetation, ( P.S. Ward) ( PSWC); Nosy Mangabe, 7.43 kmS Maroantsetra, 3 m, -15.4973, 49.76223, littoral rainforest, ( B.L. Fisher et al.) ( CASC); Parc National de Masoala, 39.4 km150° SSE Maroantsetra, 200 m, -15.71, 49.97, rainforest, ( B.L. Fisher, H.J. Ratsirarson) ( CASC); Parc National Mananara-Nord, 7.1 km261° Antanambe, 225 m, -16.455, 49.7875, rainforest, ( B.L. Fisher et al.) ( CASC); Parcelle K7 Tampolo, 10 m, -17.28333, 49.41667, littoral forest, ( Malagasyant team) ( CASC); Réserve Nationale Integrale Betampona, Betampona 35.1 kmNW Toamasina, 500 m, - 17.91801, 49.20074, rainforest, ( B.L. Fisher et al.) ( CASC); Réserve Spéciale Ambatovaky, Sandrangato river, 450 m, -16.77274, 49.26551, rainforest, ( B.L. Fisher et al.) ( CASC); Réserve Spéciale Ambatovaky, Sandrangato river, 475 m, -16.76912, 49.26704, rainforest, ( B.L. Fisher et al.) ( CASC); Réserve Spéciale Ambatovaky, Sandrangato river, 520 m, -16.7633, 49.26692, rainforest, ( B.L. Fisher et al.) ( CASC); Réserve Spéciale Ambatovaky, Sandrangato river, 470 m, -16.7702, 49.26638, rainforest, ( B.L. Fisher et al.) ( CASC); Réserve Spéciale Ambatovaky, Sandrangato river, 360 m, -16.81753, 49.29498, rainforest, ( B.L. Fisher et al.) ( CASC); Réserve Spéciale Ambatovaky, Sandrangato river, 425 m, -16.8162, 49.29202, rainforest, ( B.L. Fisher et al.) ( CASC); Réserve Spéciale Ambatovaky, Sandrangato river, 460 m, -16.81209, 49.29216, rainforest, ( B.L. Fisher et al.) ( CASC); Réserve Spéciale Ambatovaky, Sandrangato river, 480 m, -16.80561, 49.29507, rainforest, ( B.L. Fisher et al.) ( CASC); Réserve Ambodiriana, 4.8 km306° Manompana, along Manompana river, 125 m, -16.67233, 49.70117, rainforest, ( B.L. Fisher et al.) ( CASC); Réserve Perinet-Analamazaotra, 950 m, -18.93333, 48.43333, rainforest, (D. M. O lson) ( PSWC); Réserve Betampona, Camp Rendrirendry 34.1 km332° Toamasina, 390 m, - 17.924, 49.19967, rainforest, ( B.L. Fisher et al.) ( CASC); S.F. Tampolo, 10 kmNNE Fenoarivo Atn., 10 m, - 17.2825, 49.43, littoral rainforest, (B.L. Fisher) ( CASC); Tampolo, Masoala Peninsula, 40.4 km154° SSE Maroantsetra, 30 m, -15.73, 49.96, rainforest, ( B.L. Fisher, H.J. Ratsirarson) ( CASC); Torotorofotsy, 1005 m, - 18.77048, 48.43043, montane rainforest, ( B.L. Fisher et al.) ( CASC); Torotorofotsy, 1070 m, -18.87082, 48.34737, montane rainforest, marsh edge, ( Malagasyant team) ( CASC); Province Toliara: 10 kmNW Enakara, Res Andohahela, 420 m, -24.56667, 46.81667, rainforest, ( B.L. Fisher) ( CASC); 13 kmNW Enakara, Réserve Andohahela, 1250 m, -24.55, 46.8, montane rainforest, ( B.L. Fisher) ( CASC); 13 kmNW Enkara, Res Andohahela, 1140 m, -24.55, 46.8, montane rainforest, ( B.L. Fisher) ( CASC); 6 kmESE Imonty, Réserve Andohahela, 1200 m, -24.85, 46.75, rainforest, ( P.S. Ward) ( PSWC); Anosy Region, Anosyenne Mts, 29.9 kmNW Manantenina, 750 m, -24.13894, 47.06804, montane rainforest, ( B.L. Fisher, F.A. Esteves et al.) ( CASC); Anosy Region, Anosyenne Mts, 31.2 kmNW Manantenina, 1125 m, -24.13894, 47.06804, rainforest, ( B.L. Fisher, F.A. Esteves et al.) ( CASC); Anosy Region, Anosyenne Mts, 31.2 kmNW Manantenina, 1315 m, -24.13632, 47.05485, rainforest, ( B.L. Fisher, F.A. Esteves et al.) ( CASC); Anosy Region, Parc National Andohahela, Col de Tanatana, 400 m, -24.74969, 46.84949, rainforest, ( B.L. Fisher, F.A. Esteves et al.) ( CASC); Forêt Ivohibe55.0 km N Tolagnaro, 200 m, -24.569, 47.204, rainforest, ( B.L. Fisher et al.) ( CASC); Forêt Ivohibe 55.6 kmN Tolagnaro, 650 m, -24.56167, 47.20017, rainforest, ( B.L. Fisher et al.) ( CASC); Parc National Andohahela, Col de Tanatana, 33.3 kmNW Tolagnaro, 275 m, -24.7585, 46.85367, rainforest, ( B.L. Fisher et al.) ( CASC); Parc National Andohahela, Manangotry, 33.8 kmNW Tolagnaro, 575 m, -24.75117, 46.85783, rainforest, ( B.L. Fisher et al.) ( CASC); Parc National d'Andohahela, Col du Sedro, 3.8 km113° ESE Mahamavo, 37.6 km341° NNW Tolagnaro, 900 m, -24.76389, 46.75167, montane rainforest, ( Fisher-Griswold Arthropod Team) ( CASC); Réserve Privée Berenty, Forêt de Bealoka, Mandrare River, 14.6 km329° NNW Amboasary, 35 m, -24.95694, 46.2715, gallery forest, ( Fisher-Griswold Arthropod Team) ( CASC); Réserve Spéciale Kalambatritra, 1365 m, -23.4185, 46.4583, grassland, ( B.L. Fisher et al.) ( CASC); Réserve Spéciale Kalambatritra, Ambinanitelo, 1325 m, -23.4502, 46.45658, montane rainforest, (B.L. Fisher et al.) (CASC); Réserve Spéciale Kalambatritra, Ampanihy, 1270 m, - 23.4635, 46.4631, montane rainforest, (B.L. Fisher et al.) (CASC); Réserve Spéciale Kalambatritra, Ampanihy, 1269 m, -23.463, 46.47057, montane rainforest, (B.L. Fisher et al.) (CASC); [ Central Madagascar], ( MHNG); [ Central Madagascar], ( MHNG); [de Diversa, Museum Paris, Grandidier 1899], (MNHN); [Madagasc.], (MHNG); 11 kmSE Ampasimanolotra(= Brickaville), 5 m, -18.9, 49.13333, littoral rainforest, ( P.S. Ward) ( PSWC); massif du Tsaratanana(versant Sud), Andohanambatoafo, 2030 m, ( P. Soga) ( MNHN).
1851012870
[544,1319,1411,1436]
MHNG
Camboue
Foret d'Analamanita Province
20
21
1
Antananarivo
1851013005
CASC
Analamanga Region
701
Ambohitantely
District of Ankazobe
20
21
1
Antananarivo
1851012916
CASC
B. L. Fisher
Foret
1491
Ankazobe
Andranorovitra
20
21
1
Antananarivo
1851012798
CASC
B. L. Fisher
1312
Mandraka
20
21
1
Antananarivo
1851013007
CASC
B. L. Fisher
1360
Mandraka Park
20
21
1
Antananarivo
1851013006
CASC
1410
Ankazobe
Ambohitantely
20
21
1
Antananarivo
1851012869
CASC
B. L. Fisher
1580
Reserve Speciale d'Ambohitantely
20
21
1
Antananarivo
1851012788
CASC
B. L. Fisher
1490
Reserve Speciale d'Ambohitantely
20
21
1
Antananarivo
1851012917
[641,1375,1768,1793]
MHNG
Diego Suarez & Ch. Alluaud
Antsiranana
20
21
1
Antsiranana
1851012915
CASC
B. L. Fisher
875
Reserve Anjanaharibe-Sud
20
21
1
Antsiranana
1851012912
CASC
B. L. Fisher
1280
Reserve Anjanaharibe-Sud
20
21
1
Antsiranana
1851012871
CASC
B. L. Fisher
1200
Reserve Anjanaharibe-Sud
20
21
1
Antsiranana
1851012872
CASC
B. L. Fisher
543
Andranomatavy Forest
Ampasindava
20
21
1
Antsiranana
1851012800
CASC
600
21
22
3.9 km 181 ° S Ambaliha
20
21
1
Antsiranana
1851012852
CASC
Pref & Antsahampano S. & Montagne d'Ambre. Site
Antsiranana
21
22
1
Antsiranana
1851012855
CASC
B. L. Fisher
880
Bekona River
Betaolana Forest
21
22
1
Antsiranana
1851012911
CASC
B. L. Fisher
725
Foret de Binara
21
22
1
Antsiranana
1851012695
CASC
B. L. Fisher
520
Mont Galoko
Galoko
21
22
1
Antsiranana
1851012866
CASC
B. L. Fisher
980
Mont Galoko
Galoko
21
22
1
Antsiranana
1851012796
CASC
B. L. Fisher
1100
Mont Galoko
Galoko
21
22
1
Antsiranana
1851012862
CASC
B. L. Fisher
415
Makirovana forest
21
22
1
Antsiranana
1851012909
[429,1431,437,462]
CASC
B. L. Fisher
715
Makirovana forest
21
22
1
Antsiranana
1851012868
[151,1129,473,498]
CASC
B. L. Fisher
390
Makirovana forest
21
22
1
Antsiranana
1851012856
CASC
B. L. Fisher
715
Makirovana forest
21
22
1
Antsiranana
1851013002
CASC
960
7 km SW Park
21
22
1
Antsiranana
1851013004
CASC
B. L. Fisher
1575
Manantenina
Parc National de Marojejy
21
22
1
Antsiranana
1851012907
CASC
B. L. Fisher
1325
Antranohofa
Parc National de Marojejy
21
22
1
Antsiranana
1851012794
CASC
B. L. Fisher
775
Manantenina River
Parc National de Marojejy
21
22
1
Antsiranana
1851012902
CASC
B. L. Fisher
1100
Parc National Montagne d'Ambre
21
22
1
Antsiranana
1851012697
CASC
925
Parc National Montagne d'Ambre
21
22
1
Antsiranana
1851012861
CASC
B. L. Fisher
780
Reserve Speciale Manongarivo
21
22
1
Antsiranana
1851012786
SAVA, CASC
Low
Marojejy National Park
487
Mantella
Camp
21
22
1
Antsiranana
1851012910
CASC
40
3 km W Sakalava Beach
21
22
1
Antsiranana
1851012691
CASC
R. Harin'Hala
150
Montaigne Francais
21
22
1
Antsiranana
1851012696
CASC
R. Harin'Hala
960
Parc National Montagne d'Ambre
21
22
1
Antsiranana
1851012853
CASC
R. Harin'Hala
1125
Petit Lac
Parc National Montagne d'Ambre
21
22
1
Antsiranana
1851012783
CASC
R. Harin'Hala
10
Sakalava Beach
21
22
1
Antsiranana
1851012903
[678,1326,1188,1213]
MHNG
Imerina & Camboue
Fianarantsoa
21
22
1
Fianarantsoa
1851012858
MHNG
Nosi & Imerina & Sikora
3 km W Ranomafana
21
22
1
Fianarantsoa
1851012849
PSWC
P. S. Ward
950
Ifandiana
21
22
1
Fianarantsoa
1851012859
CASC
B. L. Fisher
1680
Reserve Andringitra
21
22
1
Fianarantsoa
1851012865
CASC
B. L. Fisher
1275
Reserve Andringitra
21
22
1
Fianarantsoa
1851012860
CASC
R. Harin'Hala
1020
Ranomafana National Park
Belle Vue
21
22
1
Fianarantsoa
1851012904
CASC
1018
District of Ifanadiana Belle
Fitovavy Fitovinany Region
21
22
1
Fianarantsoa
1851012845
CASC
1127
District of Ifanadiana
Fitovavy Fitovinany Region
21
22
1
Fianarantsoa
1851012905
CASC
1550
Foret d'Atsirakambiaty
21
22
1
Fianarantsoa
1851012693
CASC
B. L. Fisher
600
Farafangana
Foret de Vevembe
21
22
1
Fianarantsoa
1851012793
CASC
R. Harin'Hala
690
Ranomafana National Park
21
22
1
Fianarantsoa
1851012847
CASC
Lowland
36
Reserve Speciale Manombo
21
22
1
Fianarantsoa
1851012779
CASC
Low
822
Miandritsara Forest
21
22
1
Fianarantsoa
1851012787
CASC
B. L. Fisher
940
Mount Papango
21
22
1
Fianarantsoa
1851012854
CASC
B. L. Fisher
1250
Mount Papango
21
22
1
Fianarantsoa
1851012770
CASC
1100
Vatoharanana River
Parc National de Ranomafana
21
22
1
Fianarantsoa
1851012851
CASC
E Ivohibe & B. L. Fisher & Sylvain
1200
Reserve Speciale Ivohibe
21
22
1
Fianarantsoa
1851012781
CASC
B. L. Fisher & Sylvain
900
22
23
Reserve Speciale Ivohibe
21
22
1
Fianarantsoa
1851012908
CASC
B. L. Fisher
30
Farafangana
Reserve Speciale Manombo
22
23
1
Fianarantsoa
1851012700
CASC
M. Irwin & R. Harin'Hala
1130
Ranomafana National Park
22
23
1
Fianarantsoa
1851012846
CASC
D. H. & K. M. Kavanaugh
900
Talatakely area
Ranomafana National Park
22
23
1
Fianarantsoa
1851012914
900
Fianarantsoa Prov
Isalo National Park Interpretive Center
22
23
1
Fianarantsoa
1851012789
CASC
R. Harin'Hala
1110
Vohiparara
22
23
1
Fianarantsoa
1851013001
CASC
Fisher-Griswold Arthropod Team
20
Foret de Tsimembo
22
23
1
Mahajanga
1851012790
CASC
B. L. Fisher
865
48.3 km S Mandritsara
22
23
1
Mahajanga
1851013003
[417,1125,580,605]
MSNG
Baie d'Antongil & Mocquerys
Toamasina
22
23
1
Toamasina
1851012797
PSWC
Ward
250
19 km ESE Maroantsetra
22
23
1
Toamasina
1851012692
PSWC
P. S. Ward
920
Perinet
22
23
1
Toamasina
1851012792
CASC
B. L. Fisher
600
Andranobe
22
23
1
Toamasina
1851012901
CASC
B. L. Fisher
425
Andranobe
22
23
1
Toamasina
1851012784
CASC
B. L. Fisher
25
Andranobe
22
23
1
Toamasina
1851012864
CASC
B. L. Fisher
825
Ambohitsitondroina
22
23
1
Toamasina
1851012799
CASC
B. L. Fisher
1000
Ambohitsitondroina
22
23
1
Toamasina
1851012698
CASC
B. L. Fisher
1080
Ambohitsitondroina
22
23
1
Toamasina
1851012848
[257,1260,866,891]
PSWC
P. S. Ward
900
Perinet
22
23
1
Toamasina
1851012780
CASC
M. E. Irwin & R. Harin'Hala
1050
Andasibe National Park Headquarters
22
23
1
Toamasina
1851012850
CASC
B. Blaimer & F. N. Raharimalala
15
Ambalahasina
22
23
1
Toamasina
1851012867
CASC
B. L. Fisher
1010
Ambatovy
22
23
1
Toamasina
1851012857
CASC
B. L. Fisher
1000
Ambatovy
22
23
1
Toamasina
1851012791
CASC
B. L. Fisher
1080
Ambatovy
22
23
1
Toamasina
1851012778
[346,1432,1081,1106]
CASC
Malagasy
1068
Analamay
22
23
1
Toamasina
1851012863
ANGAP, CASC
M. E. Irwin & R. Harin'Hala
1025
Andasibe National Park
22
23
1
Toamasina
1851012785
CASC
B. L. Fisher
865
Ankerana
22
23
1
Toamasina
1851012795
[411,1308,1188,1213]
CASC
B. L. Fisher
750
Ankerana
22
23
1
Toamasina
1851012844
CASC
B. L. Fisher
1035
Ankerana
22
23
1
Toamasina
1851012694
CASC
B. L. Fisher
995
Bevolota
22
23
1
Toamasina
1851012688
CASC
Malagasy
1075
Moramanga
Foret Ambatovy
22
23
1
Toamasina
1851012913
[269,1299,1974,1999]
CASC
Manakambahiny Atsinanana & A. Pauly
Primary forest
22
23
1
Toamasina
1851012906
CASC
600
23
24
Montagne d'Akirindro
22
23
1
Toamasina
1851012819
CASC
470
Ambinanitelo
Montagne d'Anjanaharibe
23
24
1
Toamasina
1851012681
CASC
1100
Montagne d'Anjanaharibe
23
24
1
Toamasina
1851012830
PSWC
P. S. Ward
300
Nosy Mangabe
23
24
1
Toamasina
1851012682
[415,1359,259,284]
PSWC
P. S. Ward
5
Nosy Mangabe
23
24
1
Toamasina
1851012678
CASC
B. L. Fisher
3
Nosy Mangabe
23
24
1
Toamasina
1851012679
CASC
B. L. Fisher & H. J. Ratsirarson
200
Parc National de Masoala
23
24
1
Toamasina
1851012759
CASC
B. L. Fisher
225
Antanambe
Parc National Mananara-Nord
23
24
1
Toamasina
1851012683
CASC
Parcelle K & Malagasy
10
Tampolo
23
24
1
Toamasina
1851012670
CASC
B. L. Fisher
500
35.1 km NW Toamasina
23
24
1
Toamasina
1851012838
CASC
B. L. Fisher
450
Sandrangato river
Reserve Speciale Ambatovaky
23
24
1
Toamasina
1851012823
CASC
B. L. Fisher
475
Sandrangato river
Reserve Speciale Ambatovaky
23
24
1
Toamasina
1851012832
CASC
B. L. Fisher
520
Sandrangato river
Reserve Speciale Ambatovaky
23
24
1
Toamasina
1851012763
CASC
B. L. Fisher
470
Sandrangato river
Reserve Speciale Ambatovaky
23
24
1
Toamasina
1851012699
CASC
B. L. Fisher
360
Sandrangato river
Reserve Speciale Ambatovaky
23
24
1
Toamasina
1851012831
CASC
B. L. Fisher
425
Sandrangato river
Reserve Speciale Ambatovaky
23
24
1
Toamasina
1851012669
CASC
B. L. Fisher
460
Sandrangato river
Reserve Speciale Ambatovaky
23
24
1
Toamasina
1851012834
CASC
B. L. Fisher
480
Sandrangato river
Reserve Speciale Ambatovaky
23
24
1
Toamasina
1851012840
CASC
B. L. Fisher
Reserve Ambodiriana
125
Manompana river
Manompana
23
24
1
Toamasina
1851012841
PSWC
950
Reserve Perinet-Analamazaotra
23
24
1
Toamasina
1851012672
CASC
B. L. Fisher
390
Toamasina
23
24
1
Toamasina
1851012771
CASC
S. F. Tampolo & Fisher
10
10 km NNE Fenoarivo Atn.
23
24
1
Toamasina
1851012777
CASC
B. L. Fisher & H. J. Ratsirarson
30
40.4 km 154 ° SSE Maroantsetra
23
24
1
Toamasina
1851012772
CASC
B. L. Fisher
1005
Torotorofotsy
23
24
1
Toamasina
1851012824
CASC
Malagasy
1070
Torotorofotsy
23
24
1
Toamasina
1851012755
CASC
B. L. Fisher
420
Res Andohahela
23
24
1
Toliara
1851012835
CASC
B. L. Fisher
1250
Reserve Andohahela
23
24
1
Toliara
1851012765
CASC
B. L. Fisher
1140
Res Andohahela
23
24
1
Toliara
1851012775
PSWC
P. S. Ward
1200
Reserve Andohahela
23
24
1
Toliara
1851012753
CASC
B. L. Fisher & F. A. Esteves
750
Anosyenne Mts
Anosy Region
23
24
1
Toliara
1851012686
CASC
B. L. Fisher & F. A. Esteves
1125
Anosyenne Mts
Anosy Region
23
24
1
Toliara
1851012760
CASC
B. L. Fisher & F. A. Esteves
1315
Anosyenne Mts
Anosy Region
23
24
1
Toliara
1851012676
CASC
B. L. Fisher & F. A. Esteves
Anosy Region
400
Col de Tanatana
Parc National Andohahela
23
24
1
Toliara
1851012828
CASC
N Tolagnaro & B. L. Fisher
200
Foret Ivohibe
23
24
1
Toliara
1851012671
CASC
B. L. Fisher
650
Foret Ivohibe
23
24
1
Toliara
1851012843
CASC
B. L. Fisher
275
Col de Tanatana
Parc National Andohahela
23
24
1
Toliara
1851012782
CASC
B. L. Fisher
575
33.8 km NW Tolagnaro
23
24
1
Toliara
1851012839
CASC
Col du Sedro & Fisher-Griswold Arthropod Team
900
Tolagnaro
Parc National d'Andohahela
23
24
1
Toliara
1851012685
CASC
Fisher-Griswold Arthropod Team
Reserve Privee Berenty
35
Mandrare River
Foret de Bealoka
23
24
1
Toliara
1851012675
CASC
B. L. Fisher
1365
Reserve Speciale Kalambatritra
23
24
1
Toliara
1851012690
[1059,1432,1795,1820]
MHNG
Madagascar
Central Madagascar
23
24
1
1851012820
[151,516,1831,1856]
MHNG
Madagascar
Central Madagascar
23
24
1
1851012842
[151,1350,1867,1892]
PSWC
P. S. Ward
Madagascar
5
Brickaville
23
24
1
1851012836
MNHN
du Tsaratanana & P. Soga
Madagascar
2030
Andohanambatoafo
23
24
1
Sud