Forel, 1886b : 184 Camponotus (Myrmocamelus) Forel, 1914 : 270 Camponotus (Myrmosaga) Wheeler, 1922 : 1045 Camponotus (Mayria) Emery, 1925 : 122 Bolton, 1995 : 92 Taxonomic revision of the Malagasy Camponotus subgenus Mayria (Hymenoptera, Formicidae) using qualitative and quantitative morphology Rakotonirina, Jean Claude Fisher, Brian L. Zootaxa 2018 2018-06-20 4438 1 1 58 QDM4 Forel Forel [151,461,1651,1677] Insecta Formicidae Camponotus GBIF Animalia Hymenoptera 18 19 Arthropoda species christi     Camponotus christi  Forel, 1886b: 184. Lectotype minor worker, present designation, Centre de Madagascar [Ambatomanjaka, Miarinarivo, -18.766947, 46.869107, 1344 m] (Hildebrandt), AntWeb CASENT0101544 (MHNG). Paralectotype: 1 minor worker with same data as lectotype but specimen coded: CASENT0101435 (Camboué) (MNHN) [examined]. [Combination in  Camponotus (Myrmocamelus):  Forel, 1914: 270; in  Camponotus (Myrmosaga):  Wheeler, 1922: 1045; in  Camponotus (Mayria):  Emery, 1925: 122;  Bolton, 1995: 92, 131].   Diagnosis.Median portion of clypeus without longitudinal carina; mandible with six teeth; anteromedian margin of clypeus not bordered by a lamella; in lateral view, mesosoma long and low, its dorsal outline not a dome-like structure; propodeal declivity inclined anteriorly; body color entirely yellow to light brown, legs with same color as body.   Description. Minor worker.In full-face view, head elongate (CWb/CL: 0.76±0.03; 0.69–0.83), lateral margins straight and more or less parallel to each other, rounding broadly to the slightly, medially excised posterior margin. Anterior clypeal margin straight, with broadly angulate junction to lateral margin; median longitudinal carina absent. Eyes not breaking lateral outlines of head; their posterior margin located approximately at posterior third portion of head (PoOc/CL: 0.34±0.01; 0.31–0.36). Mandible triangular, apical margin with six sharp teeth. Antennal scape long, more than its distal half extending beyond posterior cephalic border. In lateral view, pronotum and anterior half of mesonotum forming feeble convexity, posterior half of mesonotum and propodeal dorsum straight and long, joining declivity surface with noticeable angle. Propodeal dorsum roughly three times as long as height of declivity; propodeal spiracle circular. Petiolar node as high as or higher than long, its dorsal margin rounding to anterior face and joining posterior face at an angle. Dorsum of head covered with numerous yellowish, elongate, erect hairs; pronotum and mesonotum each with a pair of erect hairs, propodeum with a pair of hairs found on junction to declivity; sparse, short pubescence present on dorsum of body. Body color entirely yellowish-orange to orange-brown and funicular segments becoming dark brown toward the apex.  Major worker.Differing from minor worker in the following characters: enlarged head, with markedly concave posterior margin; apical third of antennal scape surpassing posterior cephalic margin; robust mesosoma, metanotum distinctly visible, propodeal dorsum rounding to declivity and twice as long as height of declivity; petiolar node tapering dorsally. More pairs of erect hairs on junction of propodeal dorsum and declivity and on posterodorsal margin of petiolar node.   Discussion.Workers of  C. christiare similar to those of  C. mainty, C. foersteriand  C. maculiventrisin that their clypeus lacks a median longitudinal carina, and in lateral view, their mesosoma is long and low. However,  C. christiworkers are easily separable by their entirely yellow to light brown body color and legs. In the members of  C. mainty, their body is entirely black to reddish-black. In the two latter species, at least the trochanters and the distal portion of the coxa are yellowish to white. The cluster of  C. christishown by the dendrogram of NC-PART clustering is confirmed by the cumulative LDA at 100% identification success, indicating the support of the species hypothesized by qualitative morphologybased taxonomy. The cluster for  C. christiis located next to that of  C. foersteriand  C. mainty, which suggests that the three species are more similar to each other than other species. This finding corroborates their close morphological similarity as detected by the qualitative analysis.   Distribution and biology.  Camponotus christiis widely distributed across Madagascar. It occupies sclerophyll forests of the central high plateau, tropical dry forests in the west to coastal scrub and littoral rainforests, as well as montane rainforests in the east. While its workers have been found foraging on the forest floor and the lower portions of vegetation, their nest sites have been located in root mats in the ground layer, rotten logs, rotting tree stumps, rotten pockets above the ground, dead branches and bamboo above the ground, and in canopy moss and leaf litter.   Additional material examined.‘Andrangoloaka], (Sikora) (MHNG); [antananarivo, Museum Paris, Grandidier 1893], (Camboué) (MNHN); [Divers, tous les Divers sont de la Forêt d’Andrangoloaka, 20, Museum Paris, Grandidier 1899] (MNHN); [ Forêt d’Analamanita Province Antananarivo], ( Camboué) ( MHNG); 3 km41° NE Andranomay, 11.5 km147° SSE Anjozorobe, 1300 m, -18.47333, 47.96, montane rainforest, (Fisher, Griswold et al.) (CASC);  Analamanga Region, District of Ankazobe, Ambohitantely,  46 kmNE of Ankazobe,  701 m, - 18.198, 47.2815, sclerophyll forest, (Mike, Rinha) ( CASC);  Forêtde galerie, Andranorovitra, 24.0 km NNE Ankazobe,  1491 m, -18.11243, 47.19757, disturbed gallery montane forest, ( B.L. Fisher et al.) ( CASC);  Mandraka,  1312 m, -18.91813, 47.91717, montane rainforest, ( B.L. Fisher et al.) ( CASC);  Mandraka Park,  1360 m, -18.9019, 47.90786, montane shrubland, ( B.L. Fisher et al.) ( CASC); Réserve Spéciale d'Ambohitantely, Forêt d Ambohitantely, 20.9 km72° NE d Ankazobe,  1410 m, -18.22528, 47.28683, montane rainforest, (Fisher, Griswold et al.) ( CASC);  Réserve Spéciale d'Ambohitantely,  1580 m, -18.18762, 47.28576, montane forest, ( B.L. Fisher et al.) ( CASC);  Réserve Spéciale d'Ambohitantely,  1490 m, -18.22444, 47.2774, montane forest, ( B.L. Fisher et al.) ( CASC); Tananarive, (Waterlot) (MNHN); Province  Antsiranana:[ Diego Suarez], ( Ch. Alluaud) ( MHNG); 11.0 km WSW Befingotra, Réserve Anjanaharibe-Sud, 1565 m, -14.75, 49.45, montane rainforest, (B.L. Fisher) (CASC);   6.5 kmSSW Befingotra, Réserve Anjanaharibe-Sud,  875 m, -14.75, 49.5, rainforest, ( B.L. Fisher) ( CASC);   9.2 kmWSW Befingotra, Réserve Anjanaharibe-Sud,  1280 m, -14.75, 49.46667, montane rainforest, ( B.L. Fisher) ( CASC);   9.2 kmWSW Befingotra, Réserve Anjanaharibe-Sud,  1200 m, -14.75, 49.46667, montane rainforest, ( B.L. Fisher) ( CASC);  Ampasindava, Andranomatavy Forest,  543 m, -13.66296, 47.97936, disturbed dry forest, ( B.L. Fisher et al.) ( CASC); Ampasindava, Forêt d'Ambilanivy,  3.9 km181° S Ambaliha,  600 m, - 13.79861, 48.16167, rainforest, (Fisher, Griswold et al.) ( CASC);  AntsirananaII Pref: Antsahampano S.- Pref: Montagne d'Ambre. SiteMD1, 1049 m, -12.52765, 49.17235, in  Commelinaregrowth on path next to degraded primary riparian rainforest, (D. Lees, R. Ranaivosolo & P. Razafindraibe) ( CASC);  Betaolana Forest, along Bekona River,  880 m, -14.52996, 49.44039, rainforest, ( B.L. Fisher et al.) ( CASC);  Forêt de Binara,  9.1 km233° SW Daraina,  725 m, -13.26333, 49.60333, rainforest, ( B.L. Fisher) ( CASC);  Galokochain, Mont Galoko,  520 m, - 13.58487, 48.71818, rainforest, ( B.L. Fisher et al.) ( CASC);  Galokochain, Mont Galoko,  980 m, -13.5888, 48.72864, montane forest, ( B.L. Fisher et al.) ( CASC);  Galokochain, Mont Galoko,  1100 m, -13.59358, 48.73157, montane forest, ( B.L. Fisher et al.) ( CASC);  Makirovana forest,  415 m, -14.17066, 49.95409, rainforest, ( B.L. Fisher et al.) ( CASC);  Makirovana forest,  715 m, -14.16666, 49.95, rainforest, ( B.L. Fisher et al.) ( CASC);  Makirovana forest,  390 m, -14.10295, 50.01984, rainforest, ( B.L. Fisher et al.) ( CASC);  Makirovana forest,  715 m, -14.16666, 49.95, rainforest, ( B.L. Fisher et al.) ( CASC); Montagne d'Ambre National Park, Roussette Camp  7 kmSW Parkentrance,  960 m, -12.51444, 49.18139, rainforest, (Mike, Frank, Rin'ha) ( CASC);  Parc National de Marojejy,  25.7 km32° NNE Andapa, 10.3 km314° NW Manantenina,  1575 m, -14.445, 49.74167, montane rainforest, ( B.L. Fisher) ( CASC);  Parc National de Marojejy, Antranohofa, 26.6 km31° NNE Andapa, 10.7 km318° NW Manantenina,  1325 m, -14.44333, 49.74333, montane rainforest, ( B.L. Fisher) ( CASC);  Parc National de Marojejy, Manantenina River, 27.6 km35° NE Andapa, 9.6 km327° NNW Manantenina,  775 m, -14.435, 49.76, rainforest, ( B.L. Fisher et al.) ( CASC);  Parc National Montagne d'Ambre,  1100 m, -12.52306, 49.17901, montane rainforest, ( B.L. Fisher et al.) ( CASC);  Parc National Montagne d'Ambre,  3.6 km235° SW Joffreville,  925 m, - 12.53444, 49.1795, montane rainforest, (Fisher, Griswold et al.) ( CASC); Parc National Montagne d'Ambre, Mahasarika, 1135 m, -12.53176, 49.17662, montane rainforest, (B.L. Fisher et al.) (CASC); Parc National Montagne d'Ambre, Roussettes, 1025 m, -12.52574, 49.17238, montane rainforest, (B.L. Fisher et al.) (CASC);  Réserve Spéciale Manongarivo,  12.8 km228° SW Antanambao,  780 m, -13.97667, 48.42333, rainforest, ( B.L. Fisher) ( CASC);  SAVARegion, District of Sambava, Marojejy National Park,  5 kmW of Manantenina village, 1st Campsite ( Mantella),  487 m, -14.43817, 49.774, Lowaltitude rainforest, (Rin'ha, Mike) ( CASC);   3 kmW Sakalava Beach[white dunes site],  40 m, -12.28617, 49.36667, white dunes in littoral forest, (Harin'Hala, Irwin, Schlinger) ( CASC);  Montaigne Français,  150 m, -12.325, 49.33333, along forested limestone ridge, ( R. Harin'Hala) ( CASC);  Parc National Montagne d'Ambre[1 st campsite],  960 m, -12.51444, 49.18139, rainforest, ( R. Harin'Hala) ( CASC);  Parc National Montagne d'Ambre[ Petit Lacroad],  1125 m, -12.52028, 49.17917, rainforest, ( R. Harin'Hala) ( CASC);  Sakalava Beach[vegetated beach dunes],  10 m, -12.26278, 49.3975, across sandy trail in dwarf littoral forest, ( R. Harin'Hala) ( CASC); Province  Fianarantsoa:[ Imerina], ( Camboué) ( MHNG); [ Nosibe Imerina], ( Sikora) ( MHNG);  3 kmW Ranomafana, nr.  Ifandiana,  950 m, -21.25, 47.41667, rainforest, ( P.S. Ward) ( PSWC);   38 kmS Ambalavao, Réserve Andringitra,  1680 m, -22.2, 46.96667, montane rainforest, ( B.L. Fisher) ( CASC);   40 kmS Ambalavao, Réserve Andringitra,  1275 m, -22.21667, 46.96667, montane rainforest, ( B.L. Fisher) ( CASC); 7.6 km122° Kianjavato, Forêt Classée Vatovavy, 175 m, -21.4, 47.94, rainforest, (B.L. Fisher et al.) (CASC); 8.0 km NE Ivohibe, 1200 m, -22.42167, 46.89833, montane rainforest, (B.L. Fisher & Sylvain) (CASC); 9.0 km NE Ivohibe, 900 m, -22.42667, 46.93833, rainforest, (B.L. Fisher & Sylvain) (CASC);  Belle Vuetrail, Ranomafana National Park, FianarantsoaProv.,  1020 m, -21.2665, 47.42017, mixed tropical forest, (M.E. Irwin, F.D. Parker, R. Harin'Hala) ( CASC);  Fitovavy Fitovinany Region, District of Ifanadiana Bellevue area  1200 mS of Ranomafana National Parkentrance,  1018 m, -21.2665, 47.42017, rainforest, (Rin'ha, Mike) ( CASC);  Fitovavy Fitovinany Region, District of Ifanadiana,  12 kmW of Ranomafana,  1127 m, -21.25083, 47.40717, forest edge, open area, (Rin'ha, Mike) ( CASC);  Forêt d'Atsirakambiaty,  7.6 km285° WNW Itremo,  1550 m, -20.59333, 46.56333, montane rainforest, (Fisher, Griswold et al.) ( CASC);  Forêt de Vevembe, 66.6 km293° Farafangana,  600 m, -22.791, 47.18183, rainforest, transition to montane forest, ( B.L. Fisher et al.) ( CASC); JIRAMA water works near river, Ranomafana National Park, FianarantsoaProv.,  690 m, -21.2485, 47.45217, open area near stream, (M.E. Irwin, F.D. Parker, R. Harin'Hala) ( CASC);  Réserve Spéciale Manombo,  32 kmSE of Farafangana,  36 m, -23.02183, 47.72, Lowlandrainforest, (Rin'ha, Mike) ( CASC);  Miandritsara Forest,  40 kmS of Ambositra,  822 m, -20.79267, 47.17567, Lowaltitude rainforest, (Rin'ha, Mike) ( CASC); Parc National Befotaka-Midongy, Papango  27.7 kmS Midongy-Sud, Mount Papango,  940 m, -23.83517, 46.96367, rainforest, ( B.L. Fisher et al.) ( CASC); Parc National Befotaka-Midongy, Papango  28.5 kmS Midongy-Sud, Mount Papango,  1250 m, - 23.84083, 46.9575, montane rainforest, ( B.L. Fisher et al.) ( CASC);  Parc National de Ranomafana, Vatoharanana River,  4.1 km231° SW Ranomafana,  1100 m, -21.29, 47.43333, montane rainforest, (Fisher, Griswold et al.) ( CASC);  Réserve Spéciale Ivohibe8.0 km E Ivohibe,  1200 m, -22.48333, 46.96833, montane rainforest, ( B.L. Fisher& Sylvain) ( CASC);  Réserve Spéciale Ivohibe,  7.5 kmENE Ivohibe,  900 m, -22.47, 46.96, rainforest, ( B.L. Fisher& Sylvain) ( CASC);  Réserve Spéciale Manombo 24.5 km228° Farafangana,  30 m, -23.01583, 47.719, rainforest, ( B.L. Fisher et al.) ( CASC); radio tower, Ranomafana National Park, FianarantsoaProv.,  1130 m, - 21.25083, 47.40717, forest edge, mixed tropical forest, open area, ( M. Irwin, R. Harin'Hala) ( CASC);  Ranomafana National Park, Talatakely area,  0.4 kmWSW of Park Entrance,  900 m, -21.41667, 47.68333, mixed tropical forest, ( D. H. & K.M. Kavanaugh) ( CASC); stream area,  900 mE of Isalo National Park Interpretive Center, Fianarantsoa Prov., 750 m, -22.62667, 45.35817, open area near stream, (R. Harin'Hala) (CASC); Vohiparara, -21.23333, 47.36667, (A. Pauly) (CASC);  Vohipararabroken bridge, FianarantsoaProv.,  1110 m, -21.22617, 47.36983, high altitude rainforest, ( R. Harin'Hala) ( CASC); Province  Mahajanga: Forêt de Tsimembo,  8.7 km336° NNW Soatana,  20 m, -19.02139, 44.44067, tropical dry forest, ( Fisher-Griswold Arthropod Team) ( CASC); Réserve Spéciale Marotandrano, Marotandrano  48.3 kmS Mandritsara,  865 m, -16.28322, 48.81443, transition humid forest, ( B.L. Fisher et al.) ( CASC); Region Sofia, Bemanevika, 1606 m, -14.337, 48.58874, montane rainforest, (B.L. Fisher et al.) (CASC); Region Sofia, Bemanevika, 1657 m, -14.32826, 48.58406, montane rainforest, (B.L. Fisher et al.) (CASC); Province  Toamasina: Baie d’Antongil( Mocquerys) ( MSNG);   19 kmESE Maroantsetra,  250 m, -15.48333, 49.9, rainforest, (P.S. Ward) ( PSWC);   1 kmSSW Andasibe(= Perinet),  920 m, -18.93333, 48.41667, rainforest, ( P.S. Ward) ( PSWC);   5.3 kmSSE Ambanizana, Andranobe,  600 m, -15.66667, 49.96667, rainforest, ( B.L. Fisher) ( CASC);   5.3 kmSSE Ambanizana, Andranobe,  425 m, -15.67133, 49.97395, rainforest, ( B.L. Fisher) ( CASC);   6.3 kmS Ambanizana, Andranobe,  25 m, -15.6813, 49.958, rainforest, ( B.L. Fisher) ( CASC);   6.9 kmNE Ambanizana, Ambohitsitondroina,  825 m, -15.58506, 50.00952, rainforest, ( B.L. Fisher) ( CASC);   6.9 kmNE Ambanizana, Ambohitsitondroina,  1000 m, -15.56667, 50, montane rainforest, ( B.L. Fisher) ( CASC);   6.9 kmNE Ambanizana, Ambohitsitondroina,  1080 m, -15.56667, 50, montane rainforest, ( B.L. Fisher) ( CASC);   6 kmESE Andasibe(= Perinet),  900 m, -18.95, 48.46667, rainforest, ( P.S. Ward) ( PSWC); 7 SE Andasibe National Park Headquarters,  1050 m, -18.96278, 48.45267, tropical forest, ( M.E. Irwin, R. Harin'Hala) ( CASC);  Ambalahasina, 62.4 km19° Toamasina,  15 m, -17.59452, 49.46785, coastal scrub, ( B. Blaimer, F.N. Raharimalala) ( CASC);  Ambatovy,  12.4 kmNE Moramanga,  1010 m, -18.84963, 48.2947, montane rainforest, ( B.L. Fisher et al.) ( CASC);  Ambatovy,  12.4 kmNE Moramanga,  1000 m, -18.84773, 48.29568, montane rainforest, ( B.L. Fisher et al.) ( CASC);  Ambatovy,  12.4 kmNE Moramanga,  1080 m, -18.83937, 48.30842, montane rainforest, ( B.L. Fisher et al.) ( CASC);  Analamay,  1068 m, -18.80623, 48.33707, montane rainforest, ( Malagasyant team) ( CASC);  Andasibe National Park, botanic garden near entrance, west of ANGAPoffice,  1025 m, -18.92639, 48.40783, tropical forest, ( M.E. Irwin, R. Harin'Hala) ( CASC);  Ankerana,  865 m, -18.40062, 48.81311, rainforest, ( B.L. Fisher et al.) ( CASC);  Ankerana,  750 m, -18.40829, 48.82107, rainforest, ( B.L. Fisher et al.) ( CASC);  Ankerana,  1035 m, -18.4017, 48.80605, montane forest, ( B.L. Fisher et al.) ( CASC);  Bevolota  17.1 kmN Andasibe,  995 m, - 18.77071, 48.43164, montane rainforest, ( B.L. Fisher et al.) ( CASC); Corridor Forestier Analamay-Mantadia, Ambatoharanana, 1016 m, -18.79944, 48.40375, rainforest, (B.L. Fisher et al.) (CASC); Corridor Forestier Analamay-Mantadia, Ambatoharanana, 968 m, -18.80424, 48.40081, rainforest, (B.L. Fisher et al.) (CASC); Corridor Forestier Analamay-Mantadia, Ambatoharanana, 1013 m, -18.80388, 48.40506, rainforest, (B.L. Fisher et al.) (CASC); Corridor Forestier Analamay-Mantadia, Ambatoharanana, 968 m, -18.80424, 48.40081, rainforest, (B.L. Fisher et al.) (CASC); Corridor Forestier Analamay-Mantadia, Ambatoharanana, 1064 m, -18.80398, 48.40358, rainforest, (B.L. Fisher et al.) (CASC); Corridor Forestier Analamay-Mantadia, Ambatoharanana, 968 m, -18.80424, 48.40081, rainforest, (B.L. Fisher et al.) (CASC); Corridor Forestier Analamay-Mantadia, Ambatoharanana, 960 m, -18.80438, 48.40735, rainforest, (B.L. Fisher et al.) (CASC); Corridor Forestier Analamay-Mantadia, Ambohibolakely, 1044 m, -18.76087, 48.37128, rainforest, (B.L. Fisher et al.) (CASC); Corridor Forestier Analamay-Mantadia, Ambohibolakely, 983 m, -18.76131, 48.36437, rainforest, (B.L. Fisher et al.) (CASC); Corridor Forestier Analamay-Mantadia, Ambohibolakely, 918 m, -18.77898, 48.36375, rainforest, (B.L. Fisher et al.) (CASC); Corridor Forestier Analamay-Mantadia, Ambohibolakely, 983 m, -18.76131, 48.36437, rainforest, (B.L. Fisher et al.) (CASC); Corridor Forestier Analamay-Mantadia, Tsaravoniana, 984 m, - 18.76369, 48.4203, rainforest, (B.L. Fisher et al.) (CASC); Corridor Forestier Analamay-Mantadia, Tsaravoniana, 1039 m, -18.76465, 48.41938, rainforest, (B.L. Fisher et al.) (CASC); Corridor Forestier Analamay-Mantadia, Tsaravoniana, 1018 m, -18.75737, 48.42302, rainforest, (B.L. Fisher et al.) (CASC); F.C. Andriantantely, 530 m, - 18.695, 48.81333, rainforest, (H.J. Ratsirarson) (CASC); F.C. Didy, 960 m, -18.19833, 48.57833, rainforest, (H.J. Ratsirarson) (CASC); F.C. Sandranantitra, 450 m, -18.04833, 49.09167, rainforest, (H.J. Ratsirarson) (CASC);  Forêt Ambatovy, 14.3 km57° Moramanga,  1075 m, -18.85083, 48.32, montane rainforest, ( Malagasyant team) ( CASC);  Manakambahiny Atsinanana, -17.75, 48.71667, Primary forest, ( A. Pauly) ( CASC);  Montagne d'Akirindro  7.6 km341° NNW Ambinanitelo,  600 m, -15.28833, 49.54833, rainforest, (Fisher, Griswold et al.) ( CASC);  Montagne d'Anjanaharibe, 18.0 km 21° NNE Ambinanitelo,  470 m, -15.18833, 49.615, rainforest, (Fisher, Griswold et al.) ( CASC);  Montagne d'Anjanaharibe,  19.5 km27° NNE Ambinanitelo,  1100 m, -15.17833, 49.635, montane rainforest, (Fisher, Griswold et al.) ( CASC);  Nosy Mangabe,  300 m, -15.5, 49.76667, rainforest, ( P.S. Ward) ( PSWC);  Nosy Mangabe,  5 m, -15.5, 49.76667, littoral vegetation, ( P.S. Ward) ( PSWC);  Nosy Mangabe,  7.43 kmS Maroantsetra,  3 m, -15.4973, 49.76223, littoral rainforest, ( B.L. Fisher et al.) ( CASC);  Parc National de Masoala,  39.4 km150° SSE Maroantsetra,  200 m, -15.71, 49.97, rainforest, ( B.L. Fisher, H.J. Ratsirarson) ( CASC);  Parc National Mananara-Nord, 7.1 km261° Antanambe,  225 m, -16.455, 49.7875, rainforest, ( B.L. Fisher et al.) ( CASC);  Parcelle K7 Tampolo,  10 m, -17.28333, 49.41667, littoral forest, ( Malagasyant team) ( CASC); Réserve Nationale Integrale Betampona, Betampona  35.1 kmNW Toamasina,  500 m, - 17.91801, 49.20074, rainforest, ( B.L. Fisher et al.) ( CASC);  Réserve Spéciale Ambatovaky, Sandrangato river,  450 m, -16.77274, 49.26551, rainforest, ( B.L. Fisher et al.) ( CASC);  Réserve Spéciale Ambatovaky, Sandrangato river,  475 m, -16.76912, 49.26704, rainforest, ( B.L. Fisher et al.) ( CASC);  Réserve Spéciale Ambatovaky, Sandrangato river,  520 m, -16.7633, 49.26692, rainforest, ( B.L. Fisher et al.) ( CASC);  Réserve Spéciale Ambatovaky, Sandrangato river,  470 m, -16.7702, 49.26638, rainforest, ( B.L. Fisher et al.) ( CASC);  Réserve Spéciale Ambatovaky, Sandrangato river,  360 m, -16.81753, 49.29498, rainforest, ( B.L. Fisher et al.) ( CASC);  Réserve Spéciale Ambatovaky, Sandrangato river,  425 m, -16.8162, 49.29202, rainforest, ( B.L. Fisher et al.) ( CASC);  Réserve Spéciale Ambatovaky, Sandrangato river,  460 m, -16.81209, 49.29216, rainforest, ( B.L. Fisher et al.) ( CASC);  Réserve Spéciale Ambatovaky, Sandrangato river,  480 m, -16.80561, 49.29507, rainforest, ( B.L. Fisher et al.) ( CASC);  Réserve Ambodiriana, 4.8 km306° Manompana, along Manompana river,  125 m, -16.67233, 49.70117, rainforest, ( B.L. Fisher et al.) ( CASC);  Réserve Perinet-Analamazaotra,  950 m, -18.93333, 48.43333, rainforest, (D. M. O lson) ( PSWC); Réserve Betampona, Camp Rendrirendry 34.1 km332° Toamasina,  390 m, - 17.924, 49.19967, rainforest, ( B.L. Fisher et al.) ( CASC);  S.F. Tampolo,  10 kmNNE Fenoarivo Atn.,  10 m, - 17.2825, 49.43, littoral rainforest, (B.L. Fisher) ( CASC); Tampolo, Masoala Peninsula,  40.4 km154° SSE Maroantsetra,  30 m, -15.73, 49.96, rainforest, ( B.L. Fisher, H.J. Ratsirarson) ( CASC);  Torotorofotsy,  1005 m, - 18.77048, 48.43043, montane rainforest, ( B.L. Fisher et al.) ( CASC);  Torotorofotsy,  1070 m, -18.87082, 48.34737, montane rainforest, marsh edge, ( Malagasyant team) ( CASC); Province  Toliara:  10 kmNW Enakara, Res Andohahela,  420 m, -24.56667, 46.81667, rainforest, ( B.L. Fisher) ( CASC);   13 kmNW Enakara, Réserve Andohahela,  1250 m, -24.55, 46.8, montane rainforest, ( B.L. Fisher) ( CASC);   13 kmNW Enkara, Res Andohahela,  1140 m, -24.55, 46.8, montane rainforest, ( B.L. Fisher) ( CASC);   6 kmESE Imonty, Réserve Andohahela,  1200 m, -24.85, 46.75, rainforest, ( P.S. Ward) ( PSWC);  Anosy Region, Anosyenne Mts,  29.9 kmNW Manantenina,  750 m, -24.13894, 47.06804, montane rainforest, ( B.L. Fisher, F.A. Esteves et al.) ( CASC);  Anosy Region, Anosyenne Mts,  31.2 kmNW Manantenina,  1125 m, -24.13894, 47.06804, rainforest, ( B.L. Fisher, F.A. Esteves et al.) ( CASC);  Anosy Region, Anosyenne Mts,  31.2 kmNW Manantenina,  1315 m, -24.13632, 47.05485, rainforest, ( B.L. Fisher, F.A. Esteves et al.) ( CASC);  Anosy Region, Parc National Andohahela, Col de Tanatana,  400 m, -24.74969, 46.84949, rainforest, ( B.L. Fisher, F.A. Esteves et al.) ( CASC);  Forêt Ivohibe55.0 km N Tolagnaro,  200 m, -24.569, 47.204, rainforest, ( B.L. Fisher et al.) ( CASC);  Forêt Ivohibe  55.6 kmN Tolagnaro,  650 m, -24.56167, 47.20017, rainforest, ( B.L. Fisher et al.) ( CASC);  Parc National Andohahela, Col de Tanatana,  33.3 kmNW Tolagnaro,  275 m, -24.7585, 46.85367, rainforest, ( B.L. Fisher et al.) ( CASC); Parc National Andohahela, Manangotry,  33.8 kmNW Tolagnaro,  575 m, -24.75117, 46.85783, rainforest, ( B.L. Fisher et al.) ( CASC);  Parc National d'Andohahela, Col du Sedro,  3.8 km113° ESE Mahamavo, 37.6 km341° NNW Tolagnaro,  900 m, -24.76389, 46.75167, montane rainforest, ( Fisher-Griswold Arthropod Team) ( CASC);  Réserve Privée Berenty, Forêt de Bealoka, Mandrare River,  14.6 km329° NNW Amboasary,  35 m, -24.95694, 46.2715, gallery forest, ( Fisher-Griswold Arthropod Team) ( CASC);  Réserve Spéciale Kalambatritra,  1365 m, -23.4185, 46.4583, grassland, ( B.L. Fisher et al.) ( CASC); Réserve Spéciale Kalambatritra, Ambinanitelo, 1325 m, -23.4502, 46.45658, montane rainforest, (B.L. Fisher et al.) (CASC); Réserve Spéciale Kalambatritra, Ampanihy, 1270 m, - 23.4635, 46.4631, montane rainforest, (B.L. Fisher et al.) (CASC); Réserve Spéciale Kalambatritra, Ampanihy, 1269 m, -23.463, 46.47057, montane rainforest, (B.L. Fisher et al.) (CASC); [ Central Madagascar], ( MHNG); [ Central Madagascar], ( MHNG); [de Diversa, Museum Paris, Grandidier 1899], (MNHN); [Madagasc.], (MHNG);   11 kmSE Ampasimanolotra(= Brickaville),  5 m, -18.9, 49.13333, littoral rainforest, ( P.S. Ward) ( PSWC); massif du Tsaratanana(versant Sud), Andohanambatoafo,  2030 m, ( P. Soga) ( MNHN). 1851012870 [544,1319,1411,1436] MHNG Camboue Foret d'Analamanita Province 20 21 1 Antananarivo 1851013005 CASC Analamanga Region 701 Ambohitantely District of Ankazobe 20 21 1 Antananarivo 1851012916 CASC B. L. Fisher Foret 1491 Ankazobe Andranorovitra 20 21 1 Antananarivo 1851012798 CASC B. L. Fisher 1312 Mandraka 20 21 1 Antananarivo 1851013007 CASC B. L. Fisher 1360 Mandraka Park 20 21 1 Antananarivo 1851013006 CASC 1410 Ankazobe Ambohitantely 20 21 1 Antananarivo 1851012869 CASC B. L. Fisher 1580 Reserve Speciale d'Ambohitantely 20 21 1 Antananarivo 1851012788 CASC B. L. Fisher 1490 Reserve Speciale d'Ambohitantely 20 21 1 Antananarivo 1851012917 [641,1375,1768,1793] MHNG Diego Suarez & Ch. Alluaud Antsiranana 20 21 1 Antsiranana 1851012915 CASC B. L. Fisher 875 Reserve Anjanaharibe-Sud 20 21 1 Antsiranana 1851012912 CASC B. L. Fisher 1280 Reserve Anjanaharibe-Sud 20 21 1 Antsiranana 1851012871 CASC B. L. Fisher 1200 Reserve Anjanaharibe-Sud 20 21 1 Antsiranana 1851012872 CASC B. L. Fisher 543 Andranomatavy Forest Ampasindava 20 21 1 Antsiranana 1851012800 CASC 600 21 22 3.9 km 181 ° S Ambaliha 20 21 1 Antsiranana 1851012852 CASC Pref & Antsahampano S. & Montagne d'Ambre. Site Antsiranana 21 22 1 Antsiranana 1851012855 CASC B. L. Fisher 880 Bekona River Betaolana Forest 21 22 1 Antsiranana 1851012911 CASC B. L. Fisher 725 Foret de Binara 21 22 1 Antsiranana 1851012695 CASC B. L. Fisher 520 Mont Galoko Galoko 21 22 1 Antsiranana 1851012866 CASC B. L. Fisher 980 Mont Galoko Galoko 21 22 1 Antsiranana 1851012796 CASC B. L. Fisher 1100 Mont Galoko Galoko 21 22 1 Antsiranana 1851012862 CASC B. L. Fisher 415 Makirovana forest 21 22 1 Antsiranana 1851012909 [429,1431,437,462] CASC B. L. Fisher 715 Makirovana forest 21 22 1 Antsiranana 1851012868 [151,1129,473,498] CASC B. L. Fisher 390 Makirovana forest 21 22 1 Antsiranana 1851012856 CASC B. L. Fisher 715 Makirovana forest 21 22 1 Antsiranana 1851013002 CASC 960 7 km SW Park 21 22 1 Antsiranana 1851013004 CASC B. L. Fisher 1575 Manantenina Parc National de Marojejy 21 22 1 Antsiranana 1851012907 CASC B. L. Fisher 1325 Antranohofa Parc National de Marojejy 21 22 1 Antsiranana 1851012794 CASC B. L. Fisher 775 Manantenina River Parc National de Marojejy 21 22 1 Antsiranana 1851012902 CASC B. L. Fisher 1100 Parc National Montagne d'Ambre 21 22 1 Antsiranana 1851012697 CASC 925 Parc National Montagne d'Ambre 21 22 1 Antsiranana 1851012861 CASC B. L. Fisher 780 Reserve Speciale Manongarivo 21 22 1 Antsiranana 1851012786 SAVA, CASC Low Marojejy National Park 487 Mantella Camp 21 22 1 Antsiranana 1851012910 CASC 40 3 km W Sakalava Beach 21 22 1 Antsiranana 1851012691 CASC R. Harin'Hala 150 Montaigne Francais 21 22 1 Antsiranana 1851012696 CASC R. Harin'Hala 960 Parc National Montagne d'Ambre 21 22 1 Antsiranana 1851012853 CASC R. Harin'Hala 1125 Petit Lac Parc National Montagne d'Ambre 21 22 1 Antsiranana 1851012783 CASC R. Harin'Hala 10 Sakalava Beach 21 22 1 Antsiranana 1851012903 [678,1326,1188,1213] MHNG Imerina & Camboue Fianarantsoa 21 22 1 Fianarantsoa 1851012858 MHNG Nosi & Imerina & Sikora 3 km W Ranomafana 21 22 1 Fianarantsoa 1851012849 PSWC P. S. Ward 950 Ifandiana 21 22 1 Fianarantsoa 1851012859 CASC B. L. Fisher 1680 Reserve Andringitra 21 22 1 Fianarantsoa 1851012865 CASC B. L. Fisher 1275 Reserve Andringitra 21 22 1 Fianarantsoa 1851012860 CASC R. Harin'Hala 1020 Ranomafana National Park Belle Vue 21 22 1 Fianarantsoa 1851012904 CASC 1018 District of Ifanadiana Belle Fitovavy Fitovinany Region 21 22 1 Fianarantsoa 1851012845 CASC 1127 District of Ifanadiana Fitovavy Fitovinany Region 21 22 1 Fianarantsoa 1851012905 CASC 1550 Foret d'Atsirakambiaty 21 22 1 Fianarantsoa 1851012693 CASC B. L. Fisher 600 Farafangana Foret de Vevembe 21 22 1 Fianarantsoa 1851012793 CASC R. Harin'Hala 690 Ranomafana National Park 21 22 1 Fianarantsoa 1851012847 CASC Lowland 36 Reserve Speciale Manombo 21 22 1 Fianarantsoa 1851012779 CASC Low 822 Miandritsara Forest 21 22 1 Fianarantsoa 1851012787 CASC B. L. Fisher 940 Mount Papango 21 22 1 Fianarantsoa 1851012854 CASC B. L. Fisher 1250 Mount Papango 21 22 1 Fianarantsoa 1851012770 CASC 1100 Vatoharanana River Parc National de Ranomafana 21 22 1 Fianarantsoa 1851012851 CASC E Ivohibe & B. L. Fisher & Sylvain 1200 Reserve Speciale Ivohibe 21 22 1 Fianarantsoa 1851012781 CASC B. L. Fisher & Sylvain 900 22 23 Reserve Speciale Ivohibe 21 22 1 Fianarantsoa 1851012908 CASC B. L. Fisher 30 Farafangana Reserve Speciale Manombo 22 23 1 Fianarantsoa 1851012700 CASC M. Irwin & R. Harin'Hala 1130 Ranomafana National Park 22 23 1 Fianarantsoa 1851012846 CASC D. H. & K. M. Kavanaugh 900 Talatakely area Ranomafana National Park 22 23 1 Fianarantsoa 1851012914 900 Fianarantsoa Prov Isalo National Park Interpretive Center 22 23 1 Fianarantsoa 1851012789 CASC R. Harin'Hala 1110 Vohiparara 22 23 1 Fianarantsoa 1851013001 CASC Fisher-Griswold Arthropod Team 20 Foret de Tsimembo 22 23 1 Mahajanga 1851012790 CASC B. L. Fisher 865 48.3 km S Mandritsara 22 23 1 Mahajanga 1851013003 [417,1125,580,605] MSNG Baie d'Antongil & Mocquerys Toamasina 22 23 1 Toamasina 1851012797 PSWC Ward 250 19 km ESE Maroantsetra 22 23 1 Toamasina 1851012692 PSWC P. S. Ward 920 Perinet 22 23 1 Toamasina 1851012792 CASC B. L. Fisher 600 Andranobe 22 23 1 Toamasina 1851012901 CASC B. L. Fisher 425 Andranobe 22 23 1 Toamasina 1851012784 CASC B. L. Fisher 25 Andranobe 22 23 1 Toamasina 1851012864 CASC B. L. Fisher 825 Ambohitsitondroina 22 23 1 Toamasina 1851012799 CASC B. L. Fisher 1000 Ambohitsitondroina 22 23 1 Toamasina 1851012698 CASC B. L. Fisher 1080 Ambohitsitondroina 22 23 1 Toamasina 1851012848 [257,1260,866,891] PSWC P. S. Ward 900 Perinet 22 23 1 Toamasina 1851012780 CASC M. E. Irwin & R. Harin'Hala 1050 Andasibe National Park Headquarters 22 23 1 Toamasina 1851012850 CASC B. Blaimer & F. N. Raharimalala 15 Ambalahasina 22 23 1 Toamasina 1851012867 CASC B. L. Fisher 1010 Ambatovy 22 23 1 Toamasina 1851012857 CASC B. L. Fisher 1000 Ambatovy 22 23 1 Toamasina 1851012791 CASC B. L. Fisher 1080 Ambatovy 22 23 1 Toamasina 1851012778 [346,1432,1081,1106] CASC Malagasy 1068 Analamay 22 23 1 Toamasina 1851012863 ANGAP, CASC M. E. Irwin & R. Harin'Hala 1025 Andasibe National Park 22 23 1 Toamasina 1851012785 CASC B. L. Fisher 865 Ankerana 22 23 1 Toamasina 1851012795 [411,1308,1188,1213] CASC B. L. Fisher 750 Ankerana 22 23 1 Toamasina 1851012844 CASC B. L. Fisher 1035 Ankerana 22 23 1 Toamasina 1851012694 CASC B. L. Fisher 995 Bevolota 22 23 1 Toamasina 1851012688 CASC Malagasy 1075 Moramanga Foret Ambatovy 22 23 1 Toamasina 1851012913 [269,1299,1974,1999] CASC Manakambahiny Atsinanana & A. Pauly Primary forest 22 23 1 Toamasina 1851012906 CASC 600 23 24 Montagne d'Akirindro 22 23 1 Toamasina 1851012819 CASC 470 Ambinanitelo Montagne d'Anjanaharibe 23 24 1 Toamasina 1851012681 CASC 1100 Montagne d'Anjanaharibe 23 24 1 Toamasina 1851012830 PSWC P. S. Ward 300 Nosy Mangabe 23 24 1 Toamasina 1851012682 [415,1359,259,284] PSWC P. S. Ward 5 Nosy Mangabe 23 24 1 Toamasina 1851012678 CASC B. L. Fisher 3 Nosy Mangabe 23 24 1 Toamasina 1851012679 CASC B. L. Fisher & H. J. Ratsirarson 200 Parc National de Masoala 23 24 1 Toamasina 1851012759 CASC B. L. Fisher 225 Antanambe Parc National Mananara-Nord 23 24 1 Toamasina 1851012683 CASC Parcelle K & Malagasy 10 Tampolo 23 24 1 Toamasina 1851012670 CASC B. L. Fisher 500 35.1 km NW Toamasina 23 24 1 Toamasina 1851012838 CASC B. L. Fisher 450 Sandrangato river Reserve Speciale Ambatovaky 23 24 1 Toamasina 1851012823 CASC B. L. Fisher 475 Sandrangato river Reserve Speciale Ambatovaky 23 24 1 Toamasina 1851012832 CASC B. L. Fisher 520 Sandrangato river Reserve Speciale Ambatovaky 23 24 1 Toamasina 1851012763 CASC B. L. Fisher 470 Sandrangato river Reserve Speciale Ambatovaky 23 24 1 Toamasina 1851012699 CASC B. L. Fisher 360 Sandrangato river Reserve Speciale Ambatovaky 23 24 1 Toamasina 1851012831 CASC B. L. Fisher 425 Sandrangato river Reserve Speciale Ambatovaky 23 24 1 Toamasina 1851012669 CASC B. L. Fisher 460 Sandrangato river Reserve Speciale Ambatovaky 23 24 1 Toamasina 1851012834 CASC B. L. Fisher 480 Sandrangato river Reserve Speciale Ambatovaky 23 24 1 Toamasina 1851012840 CASC B. L. Fisher Reserve Ambodiriana 125 Manompana river Manompana 23 24 1 Toamasina 1851012841 PSWC 950 Reserve Perinet-Analamazaotra 23 24 1 Toamasina 1851012672 CASC B. L. Fisher 390 Toamasina 23 24 1 Toamasina 1851012771 CASC S. F. Tampolo & Fisher 10 10 km NNE Fenoarivo Atn. 23 24 1 Toamasina 1851012777 CASC B. L. Fisher & H. J. Ratsirarson 30 40.4 km 154 ° SSE Maroantsetra 23 24 1 Toamasina 1851012772 CASC B. L. Fisher 1005 Torotorofotsy 23 24 1 Toamasina 1851012824 CASC Malagasy 1070 Torotorofotsy 23 24 1 Toamasina 1851012755 CASC B. L. Fisher 420 Res Andohahela 23 24 1 Toliara 1851012835 CASC B. L. Fisher 1250 Reserve Andohahela 23 24 1 Toliara 1851012765 CASC B. L. Fisher 1140 Res Andohahela 23 24 1 Toliara 1851012775 PSWC P. S. Ward 1200 Reserve Andohahela 23 24 1 Toliara 1851012753 CASC B. L. Fisher & F. A. Esteves 750 Anosyenne Mts Anosy Region 23 24 1 Toliara 1851012686 CASC B. L. Fisher & F. A. Esteves 1125 Anosyenne Mts Anosy Region 23 24 1 Toliara 1851012760 CASC B. L. Fisher & F. A. Esteves 1315 Anosyenne Mts Anosy Region 23 24 1 Toliara 1851012676 CASC B. L. Fisher & F. A. Esteves Anosy Region 400 Col de Tanatana Parc National Andohahela 23 24 1 Toliara 1851012828 CASC N Tolagnaro & B. L. Fisher 200 Foret Ivohibe 23 24 1 Toliara 1851012671 CASC B. L. Fisher 650 Foret Ivohibe 23 24 1 Toliara 1851012843 CASC B. L. Fisher 275 Col de Tanatana Parc National Andohahela 23 24 1 Toliara 1851012782 CASC B. L. Fisher 575 33.8 km NW Tolagnaro 23 24 1 Toliara 1851012839 CASC Col du Sedro & Fisher-Griswold Arthropod Team 900 Tolagnaro Parc National d'Andohahela 23 24 1 Toliara 1851012685 CASC Fisher-Griswold Arthropod Team Reserve Privee Berenty 35 Mandrare River Foret de Bealoka 23 24 1 Toliara 1851012675 CASC B. L. Fisher 1365 Reserve Speciale Kalambatritra 23 24 1 Toliara 1851012690 [1059,1432,1795,1820] MHNG Madagascar Central Madagascar 23 24 1 1851012820 [151,516,1831,1856] MHNG Madagascar Central Madagascar 23 24 1 1851012842 [151,1350,1867,1892] PSWC P. S. Ward Madagascar 5 Brickaville 23 24 1 1851012836 MNHN du Tsaratanana & P. Soga Madagascar 2030 Andohanambatoafo 23 24 1 Sud