Eumenes stenogaster Isely, 1917
Eumenes tricinctus Isely, 1917
Eumenes verticalis
neoboreus Bequaert, 1944
Eumenes cruciferoides Giordani Soika, 1978
Taxonomic Review of Eumenes Latreille, 1802 (Hymenoptera, Vespidae, Eumeninae) from the New World
Grandinete, Yuri Campanholo
Noll, Fernando Barbosa
Carpenter, James
Zootaxa
2018
2018-08-14
4459
1
1
52
3CDQ8
Say, 1824
Say
1824
[151,504,620,646]
Insecta
Vespidae
Eumenes
GBIF
Animalia
Hymenoptera
29
30
Arthropoda
species
verticalis
Typedata: Typedestroyed. Typelocality:“ Pennsylvania” de Saussure, 1852, 1: 41. Cresson, 1887: 287 (cat.). Dalla Torre, 1894, 9: 33 (cat.). Dalla Torre, 1904, 19: 25 (cat.). Bequaert, 1928, 101: 1004. Bequaert, 1938, 33: 61 (key), 64, 70 ( typelost). Bequaert & Taylor, 1942, 42: 68 (list). Bequaert, 1944, 71: 78 (key), 86. MacLachlan, 1980, 53: 618, 620 (key). Buck et al.2008, 5: 44 (key), 155. Eumenes coloradensisCresson, 1875, 5: 717. REVISED STATUS. Type data:Lectotype female at ANSP (examined). Typelocality: Colorado( USA). Dalla Torre, 1894, 9: 21 (cat.). Dalla Torre, 1904, 19: 21 (cat.). Tucker, 1909, 22: 286. Cresson, 1916, 1: 104 (designation of Lectotype). Isely, 1917, 10: 348 (key), 359. Bequaert, 1938, 33: 65, 69 (note on types). Bequaert, 1944, 71: 78 (key), 81. Eumenes( Alpha) cruciferorumViereck, 1908, 33: 388. REVISED STATUS. Typedata: Lectotypefemale (by present designation) and Paralectotypemale at KUNHM (examined). Typelocality:“ Thomas’ Ranch, Oak CreekCanon, 6000 ft., 20 milessouthwest of Flagstaff, Coconino Co., Arizona” (USA). Eumenes cruciferorum; Bequaert, 1938, 33: 69 (note on types). Bohart, 1951: 885. Eumenes( Alpha) enigmatusViereck, 1908, 33: 389. NEW STATUS. Typedata: Lectotypemale (by present designation) and Paralectotypemale at KUNHM (examined). Type locality:“Thomas’ Ranch, Oak Creek Canon, 6000 ft., 20 milessouthwest of Flagstaff, Coconino Co., Arizona”. Eumenes enigmatus; Bequaert, 1938, 33: 69 ( enigmaticus[!]; note on type). Bohart, 1951: 885.
Eumenes stenogaster Isely, 1917, 10: 353. NEW STATUS.
Type data:Holotype female at USNM (n° 21379) (examined). Typelocality:“ White Mts, Novo Mexico; Rio Ruidoso; about 6500 ft.” ( USA). Bequaert, 1938, 33: 70 (note on types). Bohart, 1951: 885.
Eumenes tricinctus Isely, 1917, 10: 361. REVISED STATUS.
Type data:Holotype female at USNM (n° 21383) (examined). Typelocality:“ Oregon”. Bequaert, 1938, 33: 70 (note on type).
Eumenes verticalisvar. (or subsp.) neoboreus Bequaert, 1944, 71: 76, 87, NEW STATUS
Typedata: Holotypemale at MCZ(n° 33185) (examined). Three paratypesmale at ROM (examined). Typelocality:“ Canada... Quebec: La Trappe” ( Holotype); “ Canada: Ontario( paratypes). Eumenes verticalis tricinctus; (4) 24 (9): 315. Bohart, 1951: 885 (cat.) Krombein, 1979: 1508 (cat.). MacLachlan, 1980, 53: 618. Ruiz C. et al., 1993, 88: 87 (list). Rodríguez-Palafox, 1996: 479 (list). Eumenes verticalis verticalis; Bohart, 1951: 885 (cat.). Krombein, 1979: 1508 (cat.). MacLachlan, 1980, 53: 618. Ahlstrom, 1995, 1: 102 (list). Eumenes verticalis neoboreus; Bohart, 1951: 885 (cat.). Krombein, 1979: 1508 (cat.). MacLachlan, 1980, 53: 618. Eumenes verticalis coloradensis; Bohart, 1951: 885. Krombein, 1958: 163; Krombein, 1979: 1508 (cat.). MacLachlan, 1980, 53: 618. Ruiz C. et al., 1993, 88: 83, 87 (list). Rodríguez-Palafox, 1996: 479 (list).
Eumenes cruciferoides Giordani Soika, 1978, 29: 18 (key), 28. NEW SYNONYMY.
Type data:Holotype female at MSNV (examined). Typelocality:“ Messico: Nogares, dintorni di Orizaba, 1540 m.” (Mexico). Rodríguez-Palafox, 1996: 479 (list).
Distribution: Canada(Alta, B.C., Queb, Man*, Ont., Sask*); Mexico(Lower California*, Nuevo León*, Tamaulipas*, Veracruz); U.S.A.(AZ, CA, CT, CO, DC*, DE*, ID, IL, IN*, MA, MD*, MN*, MO, NC, ND, NH, NJ, NM, NV*, NY, OH*, OR, PA*, SD*, UT, VA*, WI*, WV*, WY). * Records from the literature.
Comments and diagnosis. Eumenes verticaliswas originally described by Say (1824), from Pennsylvania (USA) (the type was destroyed), and currently it has four recognized subspecies: E. verticalis coloradensisCresson, 1875; E. verticalis tricinctus Isely, 1917, E. verticalis neoboreus Bequaert, 1944, and E. verticalis verticalis(Say), 1824. We had examined all the typespecimens from all the subspecies of E. verticalisplus 137 specimens( 57 femalesand 80 males, all with male genitalia extracted and examined) and as we stated for E. consobrinus, E. verticalishas some intraspecific variation along its distribution, mainly in color and punctation of T2 throughout the east coast of the USAand southeast Canada, which leads us to propose the abandonment of these subspecies. Because the primary typeof the species was destroyed, we compared specimens of the typical E. verticaliswith the original description of Say (1824) and we selected one of these for the redescription of the species. This specimen is established as the lectotype. Giordani Soika (1978)described E. cruciferoides( Fig. 12) and noted that it resembles E. crucifera bolliformisbut also noted that they differ in the “punctation of the clypeus, punctation and color (p. 29)”. We compared the typeof E. cruciferoidesand to the typeof E. bolliformisand, although the former is known only from the type, we maintain it as a different species based on the following features: the punctures on clypeus are coarser and extend to the apical region; the punctation on the mesepisternum is denser, with the interspaces forming carinae, particularly in the center-basal region; the pubescence on the head and mesosoma is slightly longer, most evident on the frons and scape. Although E. cruciferoidesis known only from the typespecimen, we compared it with the typical E. verticalisand no morphological differences were found besides having the ferrugineous marks contiguous with the yellow marks along body. As we observed for E. fraternusand E. cruciferavar. bolliformis, specimens found the the southern USAand/or Mexicopresent ferrugineous marks along the body. As with E. verticalis, E. cruciferoidesalso has the clypeus with coarse punctation and long pubescence along the body, including on the scape (longer than the pedicel). For these reasons, we treat E. cruciferoidesas a new synonym of E. verticalis.
1917355187
[199,509,752,774]
Pennsylvania
29
30
1
Pennsylvania
holotype
1917355144
[199,527,1150,1172]
United States of America
Colorado
29
30
1
Colorado
holotype
1917355160
[199,1436,1457,1479]
Coconino Co.
1829
Oak Creek Canon
Thomas' Ranch
29
30
1
Arizona
holotype
1917355205
[199,627,1794,1816]
White Mts
29
30
1
Novo Mexico
holotype
1917355150
[636,1006,1794,1816]
United States of America
1981
Rio Ruidoso
29
30
1
holotype
1917355154
[199,457,1948,1970]
Oregon
29
30
1
Oregon
holotype
1917355181
[199,797,182,204]
La Trappe
Canada
Quebec
30
31
2
Quebec
holotype
1917355151
[807,1108,182,204]
Canada
Ontario
30
31
1
Ontario
paratype
1917355169
[199,843,857,879]
Nogares & di Orizaba
Mexico
1540
Messico
30
31
1
holotype