Revealing the diversity of ant-eating spiders in Colombia I: morphology, distribution and taxonomy of the barronus group of the genus Tenedos O. Pickard-Cambridge, 1897 (Araneae: Zodariidae) Martínez, Leonel Brescovit, Antonio D. Quijano, Luis G. Zootaxa 2022 2022-05-02 5130 1 1 154 99BM7 Martínez & Brescovit & Quijano, 2022 Martínez & Brescovit & Quijano 2022 [151,374,1662,1689] Arachnida Zodariidae Tenedos Animalia Araneae 93 94 Arthropoda species santarosa sp. nov.   Figs 66–68; 107.   Type material.  Holotype: COLOMBIA. Risaralda: Santa Rosa de Cabal, Vereda Campo Alegrito, Parque Municipal Natural Campo Alegre, Finca La Albania, Secondary forest, Pitfall trap,  2490m[ 4°52′03″N, 75°32′48″W], A. Pulido, Y. Martínez& E. Henaoleg.,  22-24.II.2004, 1 ♂(IAvH-I-651).  Paratypes: same data as the holotype, 1 ♂(IAvH-I-654), 1 ♂(IAvH-I-653), 1 ♀(IAvH-I-655), 1 ♀(IAvH-I-656).   Etymology.The specific name is a noun in apposition taken from the typelocality.   Diagnosis.Males of  Tenedos santarosa  sp. n., resemble those of  T. eberhardi  sp. n.and  T. tatama  sp. n.by the bifid retrolateral tibial apophysis (RTA) and large median apophysis (MA) ( Figs 64A–D; 65A–B; 67A–D; 68A–B; 70A–D; 71A–B), but are distinguished by having thirteen dorsal abdominal spots; wider and laminar posterior branch of the retrolateral tibial apophysis (pRTA); thin, laminar, and long anterior branch (aRTA) with rounded apex; large bifid median apophysis with short, curved anterior branch (aMA), triangular-shaped posterior branch (pMA); large ventral tibial apophysis (VTA) ( Figs 67A–D; 68A–B). Females are similar those of  Tenedos eberhardi  sp. n.and  T. tatama  sp. n.by lateral borders (LB) reaching the medial region of median field plate (MFP); medial field plate globose in dorsal surface; seminal receptacles forming bowl-shaped in dorsal view ( Figs 64E–F; 65C–D; 67E–F; 68C–D; 70E–F; 71C–D).   Description. Male( Holotype, IAvH-I-651). Coloration ( Fig. 66A–B): carapace uniformly dark brown. Chelicerae with paturon brown, fangs brown-reddish. Endites brown, white on anterior region. Labium and sternum light brown. Legs: Coxae I–IV light yellow. Femora I–IV with base brown, white on medial region and distally brown. Patellae dark brown. Tibia I pale yellow, II–IV brown with light patches. Metatarsi-tarsi I–IV dark brown. Abdomen: dorsally dark gray with thirteen white guanine spots organized as follows: two elongated spots, anteriorly positioned; two oval spots smaller than previous ones, sub-medially positioned; four transversal and large spots, medially positioned; two sub-rounded spots, in posteromedial position; three transversal spots, liked in medial region, posteriorly positioned. Laterally dark gray without spots. Ventrally dark gray with small spots on lateral sides and a long and wide central spot. Spinnerets light brown. Measurements: total length 6.21, carapace length 3.28, width 2.05, height 1.54. Clypeus height 0.72. Eye diameters and interdistances: AME 0.10, ALE 0.12, PME 0.11, PLE 0.13; AME–AME 0.19, AME–ALE 0.21, AME–PME 0.26, PME–PME 0.31, PME–PLE 0.38, ALE–PLE 0.25. Chelicerae 0.71 length. Sternum length 1.13, width 1.14. Legs: I—femur 1.86/ patella 0.78/ tibia 1.88/ metatarsus 1.65/ tarsus 1.12/ total 7.29; II—1.67/ 0.64/ 1.34/ 1.20/ 0.74/ 5.59; III—1.30/ 0.70/ 1.16/ 0.87/ 0.45/ 4.48; IV—1.88/ 0.67/ 1.71/ 2.24/ 1.01/ 7.51. Abdomen length 2.98. Legs spines pattern (only the differences from the general pattern): I—tibia v1r-1r-2, metatarsus v1r-1r-2; II—tibia v1r-1r-1r, metatarsus v2-2-2, p0-0-1d, III—tibia v2-2-2. Palp: retrolateral process of cymbium (RPC) long, widening towards base; tegulum (T) rounded; subtegulum (St) large, longer than wide in ventral view; conductor (C) large, with short, thin sclerotized region on distal side; appendix (ApC) short, apically sharp; embolus (E) long, filiform towards apex; base of embolus (EB) approximately as long as basal tegular membrane; basal tegular membrane (BTM) wide, originated basally on tegulum, reticulated, appendix reduced; spermatic ducts (SD) S-shaped, short, wide, with both folds full open; ventral tibial apophysis (VTA) large, anteriorly projected; median apophysis (MA) large, bifid, with small, curved anterior branch (aMA), posterior branch (pMA) triangular, wideat base; retrolateral tibial apophysis (RTA) large, bifid, with thin, laminar anterior branch (aRTA), posterior branch (pRTA) wide with apical notch ( Figs 67A–D; 68A–B).  Female( Paratype, IAvH-I-656). Coloration and abdominal pattern of spots as male ( Fig. 66C–D). Measurements: total length 5.96, carapace length 2.82, width 1.82, height 1.57. Clypeus height 0.60. Eye diameters and interdistances: AME 0.09, ALE 0.12, PME 0.10, PLE 0.12; AME–AME 0.17, AME–ALE 0.26, AME–PME 0.31, PME–PME 0.32, PME–PLE 0.38, ALE–PLE 0.27. Chelicerae 0.91 length. Sternum length 1.26, width 1.20. Legs: I—femur 1.69/ patella 0.69/ tibia 1.38/ metatarsus 1.10/ tarsus 0.87/ total 5.73; II—1.48/ 0.65/ 0.99/ 1.02/ 0.80/ 4.94; III—1.27/ 0.74/ 1.02/ 1.15/ 0.64/ 4.82; IV—1.55/ 0.74/ 1.38/ 1.65/ 0.74/ 6.06. Abdomen length 3.25. Legs spines pattern (only the differences from the general pattern): I—tibia v2-0-1p, metatarsus v1r-0-2; II—tibia v1r-1r-0, metatarsus v1r-1r-2; III—tibia v2-1p- 1p.Epigyne: lateral borders ( LB) short and curved towards posteromedial region of the epigyne, forming medial and large atrium (A); median field plate ( MFP) very large, sub-rounded; copulatory ducts ( CD) very short, wide, almost undistinguished from spermathecae; seminal receptacles (SR) long, wide at base, slightly curved towards median septum; spermathecae (S) large, irregulars, posteriorly positioned; fertilization ducts ( FD) almost as long as spermathecae length ( Figs 67E–F; 68C–D).  Variation. Males (n=3): total length: 5.27–6.21; carapace length: 3.26–3.28; femur I length: 1.09–1.86. Females (n=2): total length: 5.62–5.92; carapace length: 2.82–3.10; femur I length: 1.69–1.77.   Distribution.Known from Risaralda department( Fig. 107). 4027224325 2004-02-22 2004-02-24 2004-02-22 A. Pulido; Y. Martinez & E. Henao Colombia Santa Rosa de Cabal 2490 4.8675003 Parque Municipal Natural Campo Alegre 21 -75.54667 Vereda Campo Alegrito 93 94 1 1 Risaralda holotype 4027224382 2004-02-22 2004-02-24 2004-02-22 A. Pulido; Y. Martinez & E. Henao Colombia Santa Rosa de Cabal 2490 4.8675003 Parque Municipal Natural Campo Alegre 21 -75.54667 Vereda Campo Alegrito 93 94 4 2 2 Risaralda paratype 4027224348 II, III, LB, MFP, CD, FD 97 98 1