Analysis of male Leptagrion Selys, 1876 sensu lato (Odonata: Coenagrionidae) with description of four new genera and a new species Lencioni, F. A. A. Zootaxa 2022 2022-03-02 5105 1 63 104 5CR8 Selys, 1876 Selys 1876 [151,429,223,249] Insecta Coenagrionidae Leptagrion Animalia Odonata 2 65 Arthropoda genus   Figs. A, 1–15  Typespecies.  Leptagrion macrurum( Burmeister, 1839)by original designation (Selys 1876).   Etymology:  Leptagrion= Lepto, Greek (λεπτός) for slender + agrion, neuter form of agrios (ἄγριος) meaning “wild”, which in damselflies is now used as an indicator that a genus belongs to the Coenagrionidae ( Fliedner 2006). In reference to the slender abdomen of the species.  Generic characterization.Medium sized Coenagrionidae( 40–51 mm), pale postocular spots and pale occipital bar absent, rear of the head pale (presumably pale in  L. auricepsand an enlarged area black around the occipital foramen in  L. vilelai  sp. nov.), cerci longer than S10, basal 2/3 “C”–shaped with the concavity internally, superior and inferior internal border of the “C” enlarged and apical extremity of cerci bifurcated ( Fig. 1), the dorsal branch smaller and covered with brush–like setae, ventral one folded in an inverted “T” (  L. bocainense) or “U”–shape (other species), inclined medially; paraproct vestigial (plate–like) and ental surface of penis S3 with a medio–longitudinal laminar process.   Distribution. Brazil(Atlantic Forest in Brazilfrom Bahiato Santa Catarina).   FIGURE A.  Leptagrion: 1—Appendages, lateral view; 2—Appendages, medio–ventral view; 3—Head, posterior view; 4— Penis, lateral view (S1—segment 1, S3—segment 3, TF—terminal fold, IF—internal fold). Species: