Muma 1951 : 48
Eremopus titania ( Muma 1951 )
Muma & Muma 1988 : 11
Eremocosta titania ( Muma 1951 )
Harvey 2002 : 451
Revision of the camel spider genus Eremocosta Roewer and a description of the female Eremocosta gigas Roewer (Arachnida, Solifugae)
Cushing, Paula E.
Channiago, Felix
Brookhart, Jack O.
Zootaxa
2018
2018-03-29
4402
3
443
466
6GGKZ
(Muma 1951)
Muma
1951
[151,554,980,1006]
Arachnida
Eremobatidae
Eremocosta
GBIF
Animalia
Solifugae
17
460
Arthropoda
species
titania
Eremorhax titania Muma 1951: 48–50, figs. 34–38. Eremopus titania( Muma 1951): Muma & Muma 1988: 11–12. Eremocosta titania( Muma 1951): Harvey 2002: 451.
Typematerial. Male holotypeand 2 male paratypesfrom Twenty-nine Palms, California, USA( AMNH). Female allotypefrom 12 kmnorthwest of Las Vegas, Nevada, USA( AMNH). We were able to examine the male holotypeand female allotype. Otherspecimens examined. Males(16). USA: California, Imperial County, Ogilby Rd. N. of Highway8, N 32.77447°, W 114.83692°, 74 melev., 2002, coll. Dustin A. Wood( DMNSZA.16317); USA: California, Imperial County, Highway 534 ( Ogilby Rd.) 3.7 miS. junction Highway78, N 33.03207°, W 114.90521°, 271 melev., 22 May 2000, coll. Dustin A. Wood, collected at night while driving ( DMNSZA.16322); USA: California, Kern County, Dove Springs1, N 35.4308°, W 118.0244°, 1017 melev., March 2003, coll. USGS, pitfall trap( DMNSZA.16315); USA: California, Kern County, Dove Springs1, N 35.436°, W 118.0202°, 1081 melev., June 2004, coll. USGS, pitfall trap( DMNSZA.16316); USA: California, Kern County, Dove Springs1, N 35.4378°, W 118.039°, 1057 melev., June 2003, coll. USGS, pitfall trap( DMNSZA.17170); USA: California, Riverside County, San Jacinto Mountains, Highway74, Pinyon Pines Fire Station, N 33.88°, W 116.71°, 1219 melev., 5 July 2005, coll. Wendell Icenogle, pavement near lighted building ( DMNSZA.17218); USA: California, San Bernardino County, 5 milesNW Sunshine Peak, 29 Palms Marine Base, N 34.678°, W 116.463°, 1007 melev., 17 May 2004, coll. G. Pratt( DMNSZA.15426); USA: California, San Bernardino County, 5 milesNW Sunshine Peak, 29 Palms Marine Base, N 34.678°, W 116.463°, 1007 melev., 17 May 2004, coll. G. Pratt( DMNSZA.16320); USA: California, San Bernardino County, Marine Corps Air Ground Combat Center, N 34.3033°, W 116.30°, 773 melev., July 1999, coll. USGS( DMNSZA.16327); USA: California, San Bernardino County, Joshua Tree National Park, N 33.90925°, W 115.85617°, 538 melev., August 1999, coll. USGS, pitfall trap( DMNSZA.19260); USA: Nevada, Nye County, N 36.661°, W 115.994°, 1158 melev., 20 July 1965, coll. Martin H. Muma, on road ( DMNSZA.15427); USA: Nevada, Nye County, Mercury, N 36.65869°, W 115.99426°, 1149 melev., 18 May 2010, coll. Paul Greger( DMNSZA.26870); USA: Nevada, Nye County, NevadaTest Site, Mercury, Jackass Flatsoff Jackass Flats Rd., N 36.63619°, W 116.0756°, 878 melev., 30 June-3 August 2011, coll. Paula E. Cushing, pitfall trap( DMNSZA.26887); USA: Nevada, Nye County, NevadaTest Site, Mercury, Bldg#23-652, N 36.6569°, W 115.99426°, 1149 melev., 18 May 2010, coll. Paul Greger, in building ( DMNSZA.26888); USA: California, Riverside County, Blythe, 21 June 1978( UCDno number); USA: California, Riverside County, Blythe, 18 July 1980( UCDno number). Females(17). USA: California, Imperial County, off California78, 9 miles west of junction with California86, N 33.13247°, W 115.98183°, 20 melev., 7 September 2010, coll. Wendell Icenogle, on highway in car headlights ( DMNSZA.36259); USA: California: Inyo County, Saline Valley, May 1959, Sta.63, coll. B. Banta( CAS Casent9033224); USA: California: Inyo County, Saline Valley, 15 August 1989, coll. B. Banta( CAS Casent9033229); USA: Inyo County, Saline Valley, 15 July 1959, coll. S. B. Banta( CAS Casent9033644); USA: California: Kern County, Yermo(east of Barstow), 19 May 1973, coll. F. Ennik( CAS Casent9033652); USA: California, Riverside County, Coachella Valley/ Sky Valley, near junction of Dillon and Aurora roads, N 33.90343°, W 116.35964°, 331 melev., 18 May 2009, coll. John Rajoa, inside building ( DMNSZA.36256); USA: California, Riverside County, east side of Palm Springs International Airport, on East Tachevah Dr., 1/ 10 milewest of California111, N 33.83768°, W 116.5089°, 122 melev., 2 September 2006, coll. Wendell Icenogle, on pavement near building with lights, 00:30 hr ( DMNSZA.36268); USA: California, Riverside County, Headquarters Salton Sea State Recreation Area, N 33.51062°, W 115.91995°,- 64 melev., 9 September 2004, coll. Wendell Icenogle, on concrete base of flood light feeding on insect at 24:00 hr ( DMNSZA.36900); USA: California, San Bernardino County, Amboy Rd. 8.5 mileseast of 29 Palms( Adobe Rd.), N 34.15°, W 115.90°, 1600 melev., 4 July 2010, coll. Wendell Icenogle, on pavement beside outbuilding with light ( DMNSZA.36270); USA: California, San Diego County, Ocotillo Wellsranger station, north of California78, N 33.15354°, W 116.16731°, 47 melev., 7 June 2008, coll. Wendell Icenogle, on ground near lights ( DMNSZA.36260); USA: California, San Diego County, 1.5 milessouth of center of Borrego Springs, N 33.2384°, W 116.3759°, 183 melev., 8 October 2012, coll. Wendell Icenogle, crawling slowly across road ( DMNSZA. 36269); USA: Nevada, Clark County, Las Vegas, 15 July 1987, coll. B. DePew( CAS Casent9033514); USA: Nevada, Clark County, Kyle Canyon Rd., Spring Mountains, 16 July 1979, coll. L. DeLuca( CAS Casent9033517); USA: Nevada, Clark County, Las Vegas, 9 June 1986, coll. R. Ramsay( CAS Casent9033520); USA: Nevada, Clark County, Las Vegas, 1 August 1985, coll. E. Hamilton( CAS Casent90335); USA: Nevada, Lyon County, Silver Springs, 17 July1963, coll. L. Warren( CAS Casent9033516); USA: Nevada, Nye County, Rock Valley, N 36.633°, W 116.313°, 853 melev., 22 July 1965, coll. Martin H. Muma, kept alive until 3 August 1965(DMNS ZA.15429).
Diagnosis. Eremocosta titaniais related to E. calexicensisbut differs in coloration, especially of appendages; fixed finger longer and narrower than E. calexicensis, MP of movable finger more robust as well (compare Figs. 2W & Xwith Figs. 2E & F). Male E. titaniaare also generally smaller than male E. calexicensis.Female genital operculum club-shaped with long arms; distinct from operculum of E. titania. See also diagnosis above under E. calexicensis.
Measurements. Males (n = 16).TL 24.0–33.5; CL 7.1–10.4; CH3.0–6.2; FNL 0.3–0.5; FNH 0.4–0.7; FFH 0.6–0.9; PL 20.0–35.0; PT 1.0–4.1; PMT 5.1–10.0; LI 12.0–25.0; LIV 24.0–33.0; PPL 4.0–5.4; A/CP 4.1–7.8; FNL/FNH 0.5–1.0; FFH/FNH 1.0–2.0; FFH/ CH0.1–0.3. Females (n = 17).TL 27.0–48.0; CL 8.6–13.9; CH4.0–5.9; PL 20.0–37.0; PT 1.7–2.0; PMT 4.6–6.9; LI 12.0– 24.5; LIV 25.5–41.0; PPL 3.9–6.4; A/CP 4.5–6.3. Description. Coloration. Male.Overall light yellow; appendages a bit darker with distal ends of femur and proximal ends of tibia a light purple-violet. Anterior margin of propeltidium dusky violet ( Fig. 5H). Chelicera. Male.Deep VDC, similar to E. calexicensisbut with slightly mesal orientation ( Fig. 1G). Fixed finger slightly thicker than E. calexicensis, with no median dentition. Movable finger: MP-2 MSM-MM; distinct MPL ( Figs. 2W & X). Fondal notch tiny with one RFA and two-three RFA on the ventral region of fixed finger; fondal teeth graded I, III, II, IV retrolaterally; I, II, III, IV prolaterally ( Fig. 3G). Setation. Male. Typical setation. L III, LIV with bacilli on anterior edge of coxa. Coloration. Female.Overall slightly lighter than the male. Appendages the same as male. Chelicera. Female.Fixed finger: FP-1 FSM-FM-FSD-FD. Movable finger: MP-2 MSM, proximal larger-MM; distinct MPL ( Figs. 2Y & Z). One large RFA; fondal teeth graded II, I, III, IV retrolaterally, and I, II, tiny III, serrate IV prolaterally. Setation. Female.Typical female cheliceral setation. Genital operculum. Muma’s illustration is accurate (1951, fig. 43). Club-shaped with long arms ending in oval shaped wings ( Fig. 4F).
Distribution. USA Californiaand Nevada. Eremocosta titaniaranges from the northern Mojave Desert to the Sonoran Desert west of the ColoradoRiver. It is sympatric in part of its range with the closely related E. calexicensis.
Discussion. Specimens from the more northern regions, Nevada, Northern Californiahave a darker coloration than those found in regions of Southern Californiabut male cheliceral dentition and female genital operculum remains relatively constant.
1836936119
AMNH
United States of America
Las Vegas
Male
Twenty-nine Palms
17
460
5
2
California
holotype
1836936122
AMNH
United States of America
Nevada
17
460
2
Nevada
holotype
1836936142
DMNS
A. Wood
United States of America
Other
74
32.77447
Highway
1
-114.83692
Males
17
460
1
California
1836936125
2000-05-22
DMNS
Dustin A. & Wood
United States of America
271
33.03207
Highway
1
-114.90521
Ogilby Rd.
17
460
1
California
1836936128
2003-03
USGS, DMNS
United States of America
Kern County
1017
35.4308
Dove Springs
6
-118.0244
17
460
1
California
1836936143
2004-06
USGS, DMNS
United States of America
Kern County
1081
35.436
Dove Springs
55
-118.0202
17
460
1
California
1836936132
2003-06
USGS, DMNS
United States of America
Kern County
1057
35.4378
Dove Springs
45
-118.039
17
460
1
California
1836936139
2005-07-05
DMNS
Wendell Icenogle
United States of America
Riverside County
1219
33.88
Highway
721
-116.71
San Jacinto Mountains
17
460
1
California
1836936133
2004-05-17
DMNS
G. Pratt
United States of America
San Bernardino County
1007
34.678
Palms Marine Base
71
-116.463
Sunshine Peak
17
460
1
California
1836936124
2004-05-17
DMNS
G. Pratt
United States of America
San Bernardino County
1007
34.678
Palms Marine Base
71
-116.463
Sunshine Peak
17
460
1
California
1836936130
1999-07
USGS, DMNS
United States of America
773
34.3033
Corps Air Ground Combat Center
458
-116.3
17
460
1
California
1836936127
1999-08
USGS, DMNS
United States of America
San Bernardino County
538
33.90925
Joshua Tree National Park
1
-115.85617
17
460
1
California
1836936134
1965-07-20
DMNS
Martin H. & Muma
United States of America
Nye County
1158
36.661
Nye County
70
-115.994
17
460
1
Nevada
1836936140
2010-05-18
DMNS
Paul Greger
United States of America
Nye County
1149
36.65869
Mercury
1
-115.99426
17
460
1
Nevada
1836936136
[813,1121,1965,1991]
United States of America
Nye County
Nye County
17
460
1
Nevada
1836936120
2011-06-30
2011-08-03
2011-06-30
Paula E.
United States of America
878
36.63619
Jackass Flats Rd.
4
-116.0756
Jackass Flats
17
460
1
Nevada
1836936131
[151,925,151,176]
DMNS
Cushing
United States of America
Nye County
Nye County
18
461
1
Nevada
1836936138
2010-05-18
DMNS
Paul Greger
United States of America
1149
36.6569
Bldg
5
-115.99426
18
461
1
Nevada
1836936129
1978-06-21
UCD
United States of America
Riverside County
Blythe
18
461
1
California
1836936126
1980-07-18
UCD
United States of America
Riverside County
Blythe
18
461
1
California
1836936137
United States of America
Imperial County
20
33.13247
Females
1
-115.98183
18
461
1
California
1836936135
2010-09-07
DMNS
Wendell Icenogle
United States of America
Inyo County
Saline Valley
18
461
1
California
1836936123
1959-05
CAS
Sta. & B. Banta
United States of America
Inyo County
Saline Valley
Casent
18
461
1
California
1836936166
1989-08-15
CAS
B. Banta
United States of America
Inyo County
Saline Valley
Casent
18
461
1
1836936148
1959-07-15
CAS
S. B. Banta
United States of America
Kern County
Yermo
Casent
18
461
1
California
1836936153
1973-05-19
CAS
F. Ennik
United States of America
33.90343
Sky Valley
1
-116.35964
Casent
18
461
1
California
1836936168
2009-05-18
DMNS
John Rajoa
United States of America
Riverside County
331
East Tachevah Dr.
Palm Springs International Airport
18
461
1
California
1836936162
United States of America
122
33.83768
California
4
-116.5089
18
461
1
California
1836936165
2006-09-02
DMNS
Wendell Icenogle
United States of America
Riverside County
33.51062
Headquarters Salton Sea State Recreation Area
1
-115.91995
18
461
1
California
1836936144
2004-09-09
DMNS
Wendell Icenogle
United States of America
San Bernardino County
64
34.15
Palms
720
-115.9
Amboy Rd.
18
461
1
California
1836936163
2010-07-04
DMNS
Wendell Icenogle
United States of America
San Diego County
1600
33.15354
California
1
-116.16731
Ocotillo Wells
18
461
1
California
1836936157
2008-06-07
DMNS
Wendell Icenogle
United States of America
San Diego County
47
33.2384
Borrego Springs
6
-116.3759
18
461
1
California
1836936156
2012-10-08
DMNS
Wendell Icenogle
United States of America
Clark County
183
Las Vegas
18
461
1
Nevada
1836936149
1987-07-15
CAS
B. DePew
United States of America
Clark County
Kyle Canyon Rd.
Casent
18
461
1
Nevada
1836936159
1979-07-16
CAS
L. DeLuca
United States of America
Clark County
Las Vegas
Casent
18
461
1
Nevada
1836936164
1986-06-09
CAS
R. Ramsay
United States of America
Clark County
Las Vegas
Casent
18
461
1
Nevada
1836936167
1985-08-01
CAS
E. Hamilton
United States of America
Lyon County
Silver Springs
Casent
18
461
1
Nevada
1836936145
1963-07-17
CAS
L. Warren
United States of America
Nye County
36.633
Rock Valley
70
-116.313
Casent
18
461
1
Nevada