Stechow, 1919 : 89 Vervoort & Vasseur, 1977 : 52 Stechow, 1919 : 101 Sertularia ephemera Jarvis , 1922 : 341 Millard & Bouillon, 1973 : 71 Billard, 1924 : 648 Billard, 1925 : 139 Leloup, 1932 : 161 Leloup, 1935 : 45 Fraser, 1944 : 266 Pennycuik, 1959 : 195 Hoogeveen, 1965 : 34 Cooke, 1975 : 98 Gibbons & Ryland, 1989 : 417 Watson, 2002 : 343 Mammen, 1965 : 37 Calamphora parvula Allman, 1888 Sertularia stechowi Hirohito, 1995: 215 Sertularella cumberlandica Stechow, 1913 : 140 Sertularella Symplectoscyphus cumberlandica Jäderholm, 1905 Symplectoscyphus cumberlandicus Yamada, 1959 : 60 Symplectoscyphus cumberlandicus ( Jäderholm, 1905 ) Sertularella tongenis Stechow, 1919 Sertularia Linnaeus, 1758 Sertularia tongensis Stechow, 1919 Sertularella tongensis Sertularia tongensis Sertularia ephemera Sertularia ephemera S. maldivensis ( Borradaile, 1905 ) Additional shallow-water thecate hydroids (Cnidaria: Hydrozoa) from Guadeloupe and Les Saintes, French Lesser Antilles Galea, Horia R. Zootaxa 2010 2570 1 40 4WYLD Stechow, 1919 Stechow 1919 [151,595,1802,1828] Hydrozoa Sertulariidae Sertularia Animalia Leptothecata 17 18 Cnidaria species tongensis     Sertularella tongensis  Stechow, 1919: 89, fig. F1, G1.―  Vervoort & Vasseur, 1977: 52, fig. 23. not  Sertularia tongensis  Stechow, 1919: 101, fig. N1 (=  Sertularia ephemera nom. nov.).  Sertularia thecocarpa   Jarvis, 1922: 341, pl. 24 fig. 10, syn. nov.―  Millard & Bouillon, 1973: 71, fig. 9C–D.― Morri et    al., 2009: 54, figs 2E–F, 4A–D.    Sertularella minuscula  Billard, 1924: 648, fig. 2F, syn. nov.―  Billard, 1925: 139, fig. 9.―  Leloup, 1932: 161, figs 26, 27.―  Leloup, 1935: 45.―  Fraser, 1944: 266, pl. 57 fig. 254.―  Pennycuik, 1959: 195, pl. 6 fig. 2.―Van Gemerden-  Hoogeveen, 1965: 34, figs 8–12.―Hirohito, 1974: 18, fig. 7.―  Cooke, 1975: 98.―  Gibbons & Ryland, 1989: 417, fig. 33.―  Watson, 2002: 343, fig. 3.    Sertularella parvula  Mammen, 1965: 37, fig. 69, syn. nov.(not  Calamphora parvula Allman, 1888).   Sertularia stechowiHirohito, 1995: 215, figs 72D–F, 73A–C, pl. 12 fig. C, syn. nov.    Sertularella cumberlandica―  Stechow, 1913: 140, fig. 115 [not  Sertularella(=  Symplectoscyphus)  cumberlandica Jäderholm, 1905].    Symplectoscyphus cumberlandicus―  Yamada, 1959: 60[not  Symplectoscyphus cumberlandicus( Jäderholm, 1905)].   Material examined. Stn.9: 21.11.2009, 21 m—several sterile stems to 7 mmhigh, on concretions and stem of  Eudendriumsp. (MNHN-IK.2009-818). Stn.10: 19.11.2009, 17 m—several stems to 7 mmhigh, of which one bears a gonotheca and another two side branches, on sponge; 0 2.12.2009, 15–20 m—several sterile stems, to 9 mmhigh, on sponge and  Dictyotasp. (MHNG-INVE-68724) Stn.12: 30.11.2009, 12–15 m—several sterile stems, to 6 mmhigh, on sponge and worm tube.   TABLE 2. Comparative measurements of  Sertularia tongensis(Stechow, 1919), in µm.    Jarvis (1922), as  Sertularella thecocarpa Billard (1925), as  Sertularella minuscula van Gemerden- Hoogeveen (1965), as  Sertularella minuscula Millard & Bouillon (1973), as  Sertularella thecocarpa Hirohito (1974), as  Sertularella minuscula  Internode  - length 270–340 230–630 105–175 – 300–350  - diameter at node – 80–115 60–110 – –  Hydrotheca  - free adcauline side 230–260 170–250 170–235 140–280 210–230  - adnate adcauline side 170–200 150–180 105–190 160–210 170–180  - abcauline side 230–285 230–330 200–280 270–300  - diameter at aperture 120–140 105–115 105–125 120–150 110–120  Gonotheca  - length 950 – 1060–1220 830 700–850 (3)  - max. width 800 – 575–720 530 500–550 (3) contiuned.   Remarks. This remarkable hydroid is characterized by: 1) its alternate hydrothecae (fig. 5H), instead of (sub)opposite as in the majority of species assigned to  Sertularia Linnaeus, 1758; 2) a hydrothecal aperture closed by two peculiar opercular flaps, both pleated along a median line, giving the impression of a fourflapped operculum when the hydrotheca is seen apically (fig. 5K); 3) its gonothecae (fig. 5L) and, when present, side branches (fig. 5I) arising from within the stem hydrothecae. In spite of its peculiar morphology, four additional nominal species from the Indo-Pacific have been erected for the present hydroid, viz.  Sertularia thecocarpa  Jarvis, 1922,  Sertularella minuscula Billard, 1924,  Sertularella parvula Mammen, 1965, and  Sertularia stechowiHirohito, 1995. All the available descriptions and illustrations referring to them leave no doubt about their conspecificity with Stechow’s (1919) species. Comparison of measurements from various sources (table 2) show that all fall into a common range of variation. The length of both abcauline and free adcauline sides of the hydrotheca appears highly variable and is in relation to whether the renovations (sometimes numerous, though often inconspicuous) have been included when measuring the size of the thecae. The differences in the size of the gonotheca may be due to different stages of maturation and/or to possible sexual dimorphism, though their shape is substantially the same in all the available accounts. Two typesof nematocysts occur in the present material (undischarged capsules): microbasic mastigophores (6.1–6.4 × 1.8–2.0 µm), and larger, unidentified capsules (10.8–11.4 × 3.1–3.3 µm).  Since  Sertularella tongenis Stechow, 1919clearly belongs to  Sertularia Linnaeus, 1758(see SEM images in Morri et al. 2009), it appears that this species and  Sertularia tongensis Stechow, 1919become secondary homonyms. As  Sertularella tongensiswas described on p. 89, and  Sertularia tongensison p. 101 of the same publication ( Stechow 1919), I consider the former specific name as the senior homonym (ICZN Art. 24.2, Principle of the First Reviser), and I propose the new name  Sertularia ephemera nom. nov.for the latter (ICZN Art. 60). Better knowledge of the taxonomical status of  Sertularia ephemerais actually needed, especially with respect to its relationship with  S. maldivensis( Borradaile, 1905).  Caribbean records. Bonaire( Leloup 1935, Van Gemerden-Hogeveen 1965), Farallón Centinela ( Venezuela) ( Van Gemerden-Hoogeveen 1965).  World distribution. Kingdom of Tonga( Stechow 1919), Chagos Archipelago (  Jarvis1922), Indonesia( Billard 1924, 1925), Gulf of Manaar ( Leloup 1932), Queensland ( Australia) ( Pennycuik 1959, Watson 2002), southern India( Mammen 1965), Seychelles( Millard & Bouillon 1973), Bonin Islands (Hirohito 1974), Marshall Islands( Cooke 1975), Moorea ( Vervoort & Vasseur 1977), Fiji( Gibbons & Ryland 1989), Japan( Stechow 1913, Yamada 1959, Hirohito 1995), Levant Sea ( Morri et al. 2009).