Macraspis calcarata Spinola 1835: 130 Pseudodorysthetus calcaratus P. calcaratus calcaratus ( Spinola, 1835 ) P. calcaratus meridionalis Soula, 2005 Pseudodorysthetus calcaratus amazonicus Soula 2005: 379 An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas Hielkema, Auke J. Hielkema, Meindert A. Insecta Mundi 2019 2019-10-25 732 732 1 306 (Spinola, 1835) Spinola 1835 [192,811,943,967] Insecta Rutelidae Pseudodorysthetus Animalia Coleoptera 152 151 Arthropoda species calcaratus      Macraspis calcarata Spinola 1835: 130  Note.Three presently recognized subspecies of  Pseudodorysthetus calcaratushave been described, one of which occurs in the research area. The other two are  P. calcaratus calcaratus( Spinola, 1835)(recorded from Brazil) and  P. calcaratus meridionalis Soula, 2005(recorded from Argentina, Paraguay).   §Subspecies   Pseudodorysthetus calcaratus amazonicus Soula, 2005      Pseudodorysthetus calcaratus amazonicus Soula2005: 379( Brazil(NA))   Distribution. Brazil(NA): Soula 2005: 379(NA).