Neoathyreus Howden and Martínez 1963: 350 Neoathyreus ( Mesoathyreus ) Howden 2006: 48 An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas Hielkema, Auke J. Hielkema, Meindert A. Insecta Mundi 2019 2019-10-25 732 732 1 306 Howden and Martinez, 1963 Howden and Martinez 1963 [192,808,1497,1521] Insecta Bolboceratidae Neoathyreus Animalia Coleoptera 9 8 Arthropoda genus      Neoathyreus Howden and Martínez 1963: 350  Synopsis. Howden 1985  Subgenus  MESOATHYREUS Howden, 2006      Neoathyreus( Mesoathyreus) Howden 2006: 48