Mollusks from Pliocene and Pleistocene seep deposits in Leyte, Philippines Kiel, Steffen Aguilar, Yolanda M. Kase, Tomoki Acta Palaeontologica Polonica 2020 2020-08-28 65 3 589 627 7RYTP Kiel & Aguilar & Kase, 2020 Kiel & Aguilar & Kase 2020 [862,1089,414,442] Bivalvia Lucinidae Lucinoma Animalia Lucinida 14 603 Mollusca species velosoi sp. nov.   Fig. 15.  Zoobank LSID: urn:lsid:zoobank.org:act: 7801672C-99B2-48F7-98AB- EB42257D6E9E  Etymology: In honur of Eduardo Veloso, the landowner of the typelocality at AntipoloPoint, Leyte, Philippines.  Type material: HolotypeNMP-2166, disarticulated right and left valves.    Typelocality: Antipolo Point, Leyte, Philippines.   Typehorizon: Lower Pleistocene part of the Bata Formation.  Material.—  Two specimensfrom the Pleistoceneof Antipolo Point(  Meganodontiabed): holotypeand NMNS PM 28162;  two specimensfrom the Pleistocene of Cambantug Point: NMNS PM 28161, 28163.  Dimensions(in mm).—NMP-2166, L = 70.9, H = 63.8, W = 45.9 (articulated).  Diagnosis.—Well-inflated  Lucinoma, outline rounded-oval, umbones broad, pointed, surface sculpture of numerous densely spaced, sharp commarginal ribs, few indistinct growth increments in interspaces; posterior sulcus indistinct; lunule broad, short; ligament nymph long.  Description.—Strongly and evenly inflated shell (W/H ratio of single valve = 0.37), greatest width at middle of shell; anterodorsal margin slightly concave, with angulation at transition to broadly rounded anterior margin; posterodorsal margin evenly convex, with slight angulation at transition to broadly rounded posterior margin; ventral margin evenly convex, without distinct transitions to anterior and posterior margins; escutcheon lanceolate, greatest width in posterior half; lunule short, broad, deep; hinge plate broad and thick, two thick and short cardinal teeth in each valve, cardinals 1 and 2 pointing downward, cardinals 3 and 4 pointing toward posteroventral margin; ligament nymph lanceolate, about 2/3 of length of posterodorsal shell margin; anterior adductor muscle scar moderately narrow and of moderate length for genus, reaching down to 2/3 of shell height, detached slightly from pallial for ~70% of its length; posterior adductor muscle scar elongate-oval, its lower end situated at about mid-height of shell; shell interior covered by fine striation, most strongly developed at and beyond pallial line.  Table 1. Overview of the characteristics of  Lucinomaspeciesdiscussed here. Abbreviations: H, height; L, length; W, width. Measurements without decimal point were taken from the literature.    Species L × H ×W [mm] outline inflation umbones lunule posterior sulcus    Lucinoma anemiophila 61 × 54 × 29 rounded-pentagonal weak small, pointed narrow, long well developed    Lucinoma annulata 82 [L] roundish weak small, pointed broad, short weak    Lucinoma aokii 76 × 69 × 41 roundish-pointed anterior moderate pointed, prosogyrate narrow, long weak    Lucinoma dulcinea 45 × 41 [L × H] roundish moderate small, pointed narrow, short indistinct    Lucinoma galathea 50 × 48 [L × H] rounded-pentagonal weak small, pointed narrow, long weak    Lucinoma gracilistriata 62 × 51 × 34 roundish moderate pointed, prosogyrate narrow, long weak    Lucinoma canudai 77 × 71.3 × 50.3 angular strong blunt narrow, long well developed    Lucinomaaff. kastoroae 55 × 48.4 [L × H] roundish to rounded- pentagonal weak small, pointed narrow, long weak    Lucinoma kastoroae 34.6 × 28.6 [L × H] pentagonal-pointed anteriorly weak small, pointed narrow, long weak    Lucinoma kosatorea 52.7 × 46 [L × H] oval, pointed anteriorly weak small, pointed narrow, long weak    Lucinoma rhomboidalis 28.0 × 25.2 [L × H] pentagonal weak pointed, elevated broad, short weak    Lucinoma saetheri 67.4 × 63.3 × 37.8 roundish moderate pointed, prosogyrate narrow, short weak    Lucinoma shinokii 60 × 53 × 34 rounded-pentagonal moderate blunt, prosogyrate broad, short well developed    Lucinoma sibogae 52.2 × 44.0 × 25.5 elongate-pentagonal weak small, pointed narrow, long weak    Lucinoma taiwanensis 79.6 × 70.5 × 39.1 rounded-pentagonal strong blunt, prosogyrate narrow, long weak    Lucinoma tinagoensis 57.8 × 57 × 38.9 rounded-pentagonal strong pointed, prosogyrate broad, short weak    Lucinoma velosoi 70.9 × 63.8 × 45.9 rounded-oval strong broad, pointed broad, short indistinct    Lucinoma yoshidai 39 × 35 [L × H] rounded-pentagonal weak pointed, prosogyrate narrow, long well developed  Fig. 15. The lucinid bivalve  Lucinoma velosoi sp. nov.from the Pleistocene Antipolo Point seep deposits, Leyte, Philippines, holotype NMP-2166, outer surfaces of right (A 1) and left (A 2) valves; dorsal side (A 3); inner surfaces of left (A 4) and right (A 5) valves; UV light image of right valve highlighting muscle scars, blood vessel scar, and pallial line (A 6); close-up on hinges of left (A 7) and right (A 8) valves.  Remarks.—  Lucinoma velosoidiffers from the similarly inflated  L. tinagoensisdescribed below by reaching a larger size ( Table 1), its more elongated shell (that of  L. tinagoensisis taller) and its narrower escutcheon. The Pliocene to Recent  Lucinoma galathea Marwick, 1953from New Zealandhas a more pointed umbo than  L. velosoi, broader and longer ligament nymph, reaches only about 50 mmin length, whereas  L. velosoireaches 70 mm, and is less inflated than  L. velosi( Marwick 1953; Campbell et al. 2010).  Stratigraphic and geographic range.—Early Pleistocene, Leyte, Philippines. [862,1081,683,704] Antipolo 14 603 1 Antipolo holotype [1087,1328,683,704] Philippines Leyte 14 603 1 Southern Leyte holotype NMNS Point Pleistocene 14 603 PM 28162 2 Antipolo holotype NMNS Pleistocene of Cambantug Point 14 603 PM 28161, 28163 2 Antipolo holotype