Trox batesi Harold, 1872: 126 Arrow 1912: 54 Blackwelder 1944: 128 Vaurie 1962: 154 Trox ( Omorgus ) batesi : Scholtz 1982: 8 Omorgus (Haroldomorgus) batesi : Scholtz 1986: 362 Zídek 2013: 7 Strümpher et al. 2014b: 549 Strümpher et al. 2016: 62 Zídek 2017: 98 Hielkema and Hielkema 2019: 10 Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae) Costa-Silva, Vinícius Strümpher, Werner P. Vaz-de-Mello, Fernando Z. Journal of Natural History 2021 2021-02-16 54 31 - 32 1993 2024 49DQD (Harold, 1872) : Scholtz 1986 Harold 1872 [380,935,443,469] Insecta Trogidae Omorgus Animalia Coleoptera 5 1997 Arthropoda species batesi Haroldomorgus  ( Figure 1a-c, 8)      Trox batesi Harold, 1872: 126(original description);  Arrow 1912: 54(catalogue);  Blackwelder 1944: 128(catalogue);  Vaurie 1962: 154(redescription).     Trox( Omorgus) batesi: Scholtz 1982: 8(catalogue).     Omorgus (Haroldomorgus) batesi: Scholtz 1986: 362(phylogenetics); Scholtz 1990: 1418 (redescription);  Zídek 2013: 7(catalogue);  Strümpher et al. 2014b: 549(systematics);  Strümpher et al. 2016: 62(systematics);  Zídek 2017: 98(catalogue);  Hielkema and Hielkema 2019: 10(checklist for the Guianas).   Type material examined  Holotype, by monotypy: (  ♂, MNHN, through photographs, Figure 1a-c). First label [blue]: ‘Batesi/typ./Harold’ [Harold’s handwriting]. Second label [white label]: ‘Amaz.’ [Bates’ handwriting]. Third label [white with black border, typeset]: ‘Ex-Musaeo/H.W. Bates/1892’ [vertically]. Fourth label [white, typeset]: ‘Muséum Paris/ex Coll./R. Oberthür/1952’. Fifth label [red, typeset]: ‘TYPE’. Sixth label [red, with a black border, typeset]: ‘HOLOTYPE’. Seventh label [white, typeset]: ‘ HOLOTYPE/ Omorgus/ batesi ( Harold 1872)’. Eighth label [white, typeset]: ‘MNHN/EC9646’. Type locality:  BRAZIL: Amazonia(cited by Harold as ‘Amazonestrom’).   Figure 1.Holotype of  Trox batesi Harold, 1872(now  Omorgus batesi). A, dorsal view; B, lateral view, C, holotype labels (Photos by A. Mantilleri, copyright MNHN).  Note. The lectotypedesignated by Scholtz (1990, p. 1418) from RBINS for  Trox batesidoes not belong to the original type series of this species (see Hielkema and Hielkema 2019).  Additional material examined    BOLIVIA: Santa Cruz, El Cidral, 1–28 .i.1962, R . Golbachleg. (1 IFML).   BRAZIL: Roraima, Caracaraí, vii.1997, F . Vaz-de-Melloleg . (1 CEMT); Rio Uraricoera/ Ilha de Maraca, 2–13 .v.1987, malaise [trap], J .A. Rafael, J.E.B . Brasiland L.S. Aquino leg . ( 1 ♀ CEMT).   Pará, Jacareacanga, x.1959, M . Alvarengaleg . (2 CEMT); Jacareacanga, xii.1968, M . Alvarengaleg . (6 DZUP); Parauapebas, 31 .x.2010, R .A. Zampaulo leg . (2 CEMT, 1 IEXA); Parauapebas, 5 .v.2011, R .A. Zampaulo leg. (1 CEMT).   Maranhão, Bom Jardim, Reserva Biológica do Gurupi, 3°14’S 46° 41 ʹW, 7–15 .i.2011, light trap, F . Limeira-de-Oliveira, M.M . Abreuand E.A.S. Barbosa leg . ( 1 ♂ CEMT).   Amazonas, Barcelos, near Rio Unini, 1°35’S 61°33 ʹW, 19–20 .xi.1995, mercury vapour/ black light, C . Mottaand Freireleg . (1 CEMT); Parque Nacional do Jaú, Rio Carabinani, 1°59’S 61°32 ʹW, 16–17 .iv.1994, mercury vapour/black light, C.  Mottaet al leg. (1 CVMD)   Tocantins, Pium, CanguÇu/UFT [= Federal University of Tocantins], 01–02 .xi.2016, light, B .C. Ganns leg . ( 1 ♀ CEMT); Porto Nacional, 10°31 ʹ8”S 48°17 ʹ35”W, 7 .xi.2015, light trap, A . Spechtleg. (1 CEMT).   Bahia, Muruci, ii.1971, P .C. Eliasleg (1 MZSP).   Mato Grosso, Poconé, Pousada Araras, 3 .xii.1998, J .L . Morenoand T. Mestre leg . (1 CEMT); Rio Branco, xii.1984, Magno & Alvarenga leg . ( 1 ♂ CEMT); Rosário Oeste, x .1961, no collector (1 DZUP); Santa Terezinha, ‘ Barra do Itapirapé’, xi.1964, B . Malkinleg . (1 MZSP); Várzea Grande, Sesc Pantanal, 10 .xi .2004, light trap, no collector ( 1 ♀ CEMT); Várzea Grande, Sesc Pantanal, 21 .xi .2003, no collector ( 1 ♀ CEMT); Xingu, xi.1961, Alvarenga and Werner leg. (1 MZSP).   Mato Grosso do Sul, Corumbá, Reserva Particular do Patrimônio Natural Acurizal, 17°49’S 57°33 ʹW, 24–26 .vi.2012, light trap, F .R. Tortato leg . ( 1 ♀ CEMT); Corumbá, Fazenda Santa Blanca, 7 .xii.1960, K . Lenkoleg . (2 MZSP); Miranda, EstaÇão Ferroviária Salobra, i .1955, no collector (1 MZSP); Rio Verde de Mato Grosso, x.1962, no collector (1 DZUP).   Espirito Santo, Barra de São Francisco, Corrego Itá, xi.1954, W . Zikanleg . (2 MZSP); Santa Teresa, 19 .xii.1966, C . Elias& C.T. Elias leg . (1 DZUP); Sooretama, xi.1962, A . Martinezleg . (3 SAMC, 1 ♀ MACN); Sooretama, 2 .xi .1964, Werner, Oliveiraand Seabraleg. (1 DZUP).   Rio de Janeiro, Araruama, Represa de Juturnaíba, xi.1981, M . Alvarengaleg . (1 CEMT); Campos dos Goytacazes, i.1942, M . Alvarengaleg. (1 DZUP).   São Paulo, Pradópolis, x.1976, P .S.M. Botelho leg. (1 CEMT).   Paraná, Jussara, 12 .xii .1975, ExpediÇão do Depepartamento de Zoolologiada UFPR (= Federal University of Paraná) leg. (1 DZUP).   PARAGUAY: San Pedro, Rio Ypane, xi.1979 (1 SAMC).   Canindeyú, Salto del Guairá, 13 .xii.1990, G . Arriágadaleg . (1 CVMD); Teniente Enciso, 2–3 .xi.2001, C . Aguilarleg . ( 1 ♂, 1 CVMD).   ARGENTINA: Salta, Oran Urundal, ii.1944, Monrós leg . (1 SAMC); Las Lajitas, ii.1982, Willi leg . (1 CVMD); Salta, Forest, 35 kmfrom Joaquín Víctor González, 12–14 .ii.1980, Turk-L’ Argentiner leg. (1 IFML).   Diagnosis   Omorgus batesiis the only representative of the subgenus. It is very distinctive and can be easily recognised by the smooth and shiny pronotum and elytra without tubercles or ridges. The elytral intercostae is marked with deep round punctures. For a more detailed treatment of the species, see Vaurie (1962)and Scholtz (1990).  Geographic distribution This species is widely distributed, yet rarely collected. It has been known so far from Argentina, Paraguayand Brazil, to which we add the first record for Bolivia. From Brazil, it has been recorded across the Amazon (AM, PA, RR), Cerrado (TO, MA), Pantanal (MT, MS) and Atlantic Forest (SP, RJ, ES) biomes ( Figure 8). [160,578,537,562] El Cidral Bolivia Santa Cruz 6 1998 1 Santa Cruz [578,585,537,562] El Cidral Bolivia true Santa Cruz 6 1998 1 Santa Cruz [681,831,537,562] Golbach Bolivia true Santa Cruz 6 1998 1 Santa Cruz Brazil Roraima 6 1998 1 Roraima [380,573,572,597] Vaz-de-Mello Brazil true Roraima 6 1998 1 Roraima [573,1073,572,597] CEMT Brazil Ilha de Maraca Rio Uraricoera 6 1998 1 Roraima [1072,1089,572,597] CEMT Brazil true Ilha de Maraca Rio Uraricoera 6 1998 1 Roraima [335,356,606,631] CEMT Brazil true Ilha de Maraca Rio Uraricoera 6 1998 1 Roraima [497,637,606,631] Brazil Brasil 6 1998 1 Roraima [793,925,606,631] CEMT Brazil 6 1998 1 1 Roraima Brazil Jacareacanga 6 1998 1 Para [270,443,641,666] Alvarenga Brazil true Jacareacanga 6 1998 1 Para [443,762,641,666] CEMT Brazil Jacareacanga 6 1998 1 Para [861,1033,641,666] CEMT Alvarenga Brazil true Jacareacanga 6 1998 1 Para DZUP Brazil Parauapebas 6 1998 1 Para [340,356,676,701] DZUP Brazil true Parauapebas 6 1998 1 Para [449,470,676,701] DZUP Brazil true Parauapebas 6 1998 1 Para [640,1023,676,701] CEMT Brazil Parauapebas 6 1998 1 Para [1023,1040,676,701] CEMT Brazil true Parauapebas 6 1998 1 Para [1131,1152,676,701] CEMT Brazil true Parauapebas 6 1998 1 Para Brazil -3.2333333 Reserva Biologica do Gurupi 1307 -46.683334 Bom Jardim 6 1998 1 Maranhao [284,291,745,770] Brazil true -3.2333333 Reserva Biologica do Gurupi 1307 -46.683334 Bom Jardim 6 1998 1 Maranhao [493,746,745,770] Limeira-de-Oliveira, M Brazil true -3.2333333 Reserva Biologica do Gurupi 1307 -46.683334 Bom Jardim 6 1998 1 Maranhao [773,923,745,770] Brazil Abreu 6 1998 1 Maranhao CEMT Brazil 6 1998 1 1 Maranhao [240,881,780,805] Brazil -1.5833333 Rio Unini 1308 -61.55 Barcelos 6 1998 1 Amazonas [881,902,780,805] Brazil true -1.5833333 Rio Unini 1308 -61.55 Barcelos 6 1998 1 Amazonas [302,534,814,839] Motta & Freire Brazil true -1.5833333 Rio Unini 1308 -61.55 Barcelos 6 1998 1 Amazonas CEMT Brazil -1.9833333 Rio Carabinani 1307 -61.533333 Parque Nacional do Jau 6 1998 1 Amazonas [329,350,849,874] CEMT Brazil true -1.9833333 Rio Carabinani 1307 -61.533333 Parque Nacional do Jau 6 1998 1 Amazonas [757,1029,849,874] CVMD Motta Brazil 6 1998 1 Amazonas Brazil CanguCu Pium 6 1998 1 Tocantins [837,858,884,909] Brazil true CanguCu Pium 6 1998 1 Tocantins [1009,1030,884,909] Brazil true CanguCu Pium 6 1998 1 Tocantins CEMT Brazil -10.518888 Porto Nacional 21 -48.293056 6 1998 1 1 Tocantins [747,768,918,943] CEMT Brazil true -10.518888 Porto Nacional 21 -48.293056 6 1998 1 Tocantins [984,1120,918,943] CEMT Specht Brazil true -10.518888 Porto Nacional 21 -48.293056 6 1998 1 Tocantins [240,414,953,978] Brazil Muruci 6 1998 1 Bahia [501,522,953,978] Brazil true Muruci 6 1998 1 Bahia Pousada Araras Brazil Pocone 6 1998 1 Mato Grosso [173,200,988,1013] Pousada Araras Brazil true Pocone 6 1998 1 Mato Grosso [286,303,988,1013] Pousada Araras Brazil true Pocone 6 1998 1 Mato Grosso [303,469,988,1013] Brazil Moreno 6 1998 1 Mato Grosso [596,867,988,1013] CEMT Brazil Rio Branco 6 1998 1 Mato Grosso [196,528,1022,1047] CEMT Brazil Rosario Oeste 6 1998 1 1 Mato Grosso DZUP Brazil Barra do Itapirape' Santa Terezinha 6 1998 1 Mato Grosso [384,513,1057,1082] DZUP Malkin Brazil true Barra do Itapirape' Santa Terezinha 6 1998 1 Mato Grosso [514,1009,1057,1082] MZSP Brazil Sesc Pantanal Varzea Grande 6 1998 1 Mato Grosso [1009,1029,1057,1082] MZSP Brazil true Sesc Pantanal Varzea Grande 6 1998 1 Mato Grosso CEMT Brazil Sesc Pantanal Varzea Grande 6 1998 1 1 Mato Grosso [871,892,1092,1117] CEMT Brazil true Sesc Pantanal Varzea Grande 6 1998 1 Mato Grosso CEMT Brazil Xingu 6 1998 1 1 Mato Grosso Brazil -17.816668 Reserva Particular do Patrimonio Natural Acurizal 1277 -57.55 Corumba 6 1998 1 Mato Grosso do Sul [958,979,1161,1186] Brazil true -17.816668 Reserva Particular do Patrimonio Natural Acurizal 1277 -57.55 Corumba 6 1998 1 Mato Grosso do Sul [174,194,1196,1221] Brazil true -17.816668 Reserva Particular do Patrimonio Natural Acurizal 1277 -57.55 Corumba 6 1998 1 Mato Grosso do Sul [334,875,1196,1221] CEMT Brazil Fazenda Santa Blanca Corumba 6 1998 1 1 Mato Grosso do Sul [874,901,1196,1221] CEMT Brazil true Fazenda Santa Blanca Corumba 6 1998 1 Mato Grosso do Sul [995,1121,1196,1221] CEMT Lenko Brazil true Fazenda Santa Blanca Corumba 6 1998 1 Mato Grosso do Sul MZSP Brazil EstaCao Ferroviaria Salobra Miranda 6 1998 1 Mato Grosso do Sul MZSP Brazil Rio Verde de Mato Grosso 6 1998 1 Mato Grosso Corrego Ita Brazil Barra de Sao Francisco 6 1998 1 Espirito Santo [275,389,1300,1325] Zikan Brazil true Barra de Sao Francisco 6 1998 1 Espirito Santo [389,693,1300,1325] MZSP Brazil Santa Teresa 6 1998 1 Espirito Santo [692,719,1300,1325] MZSP Brazil true Santa Teresa 6 1998 1 Espirito Santo [811,918,1300,1325] MZSP Brazil true Elias 6 1998 1 Espirito Santo DZUP Brazil Sooretama 6 1998 1 Espirito Santo [406,561,1334,1359] DZUP Martinez Brazil true Sooretama 6 1998 1 Espirito Santo [561,972,1334,1359] SAMC, MACN Brazil Sooretama 6 1998 1 1 Espirito Santo [972,993,1334,1359] SAMC, MACN Brazil true Sooretama 6 1998 1 Espirito Santo Seabra Brazil Oliveira Werner 6 1998 1 Espirito Santo [527,1091,1369,1394] Brazil Represa de Juturnaiba Araruama 6 1998 1 Rio de Janeiro [183,344,1404,1429] Alvarenga Brazil true Represa de Juturnaiba Araruama 6 1998 1 Rio de Janeiro [344,739,1404,1429] CEMT Brazil Campos dos Goytacazes 6 1998 1 Rio de Janeiro [834,995,1404,1429] CEMT Alvarenga Brazil true Campos dos Goytacazes 6 1998 1 Rio de Janeiro Brazil Pradopolis 6 1998 1 Sao Paulo [490,510,1438,1463] Brazil true Pradopolis 6 1998 1 Sao Paulo [833,1062,1438,1463] Brazil Jussara 6 1998 1 Parana [1062,1089,1438,1463] Brazil true Jussara 6 1998 1 Parana DZUP Brazil Federal University of Parana ExpediCao do Depepartamento de Zoolologia 6 1998 1 Parana [279,720,1508,1533] Paraguay Rio Ypane 6 1998 1 San Pedro Paraguay Salto del Guaira 6 1998 1 Canindeyu [271,298,1542,1567] Paraguay true Salto del Guaira 6 1998 1 Canindeyu [391,552,1542,1567] Arriagada Paraguay true Salto del Guaira 6 1998 1 Canindeyu [552,907,1542,1567] CVMD Paraguay Teniente Enciso 6 1998 1 Canindeyu [907,928,1542,1567] CVMD Paraguay true Teniente Enciso 6 1998 1 Canindeyu [1019,1152,1542,1567] CVMD Aguilar Paraguay true Teniente Enciso 6 1998 1 Canindeyu CVMD Paraguay 6 1998 1 1 Canindeyu [321,715,1577,1602] Argentina Salta 6 1998 1 Salta SAMC Argentina Las Lajitas 6 1998 1 Salta [332,1081,1612,1637] CVMD Argentina Salta Joaquin Victor Gonzalez Forest 6 1998 1 Salta [1081,1091,1612,1637] CVMD Argentina Salta true Joaquin Victor Gonzalez Forest 6 1998 1 Salta