New polychaete species collected during the expeditions ANDEEP I, II, and III to the deep Atlantic sector of the Southern Ocean in the austral summers 2002 and 2005 — Ampharetidae, Opheliidae, and Scalibregmatidae Schüller, Myriam Zootaxa 2008 1705 51 68 78GVZ [151,476,1410,1436] Polychaeta Scalibregmatidae Pseudoscalibregma Animalia 11 62 Annelida species papilia sp. nov.    Holotype.ANDEEP I – II, South Sandwich Islands, Sta. 141-10, 23 March 2002, 58°25.08’S, 25°0.77’W, 2258–2313 m, EBS, ZMH P-24761.   Paratypes.ANDEEP I – II, South Sandwich Islands, Sta. 141-10, 23 March 2002, 58°25.08’S, 25°0.77’W, 2258–2313 m, EBS, 2 specimens, ZMH P-24762.  Additional material.42-2 (3), 46-7 (2), 121-11 (1), 142-5 (1), 150-6 (5), 153-7 (1).   Etymology.The species is named after the shape of the posterior parapodia which resemble butterfly wings. Diagnosis. The species can be recognized by prominent, almost foliose dorsal and ventral cirri in its posterior parapodia, distinctly rounded prostomial lobes and a rather smooth to irregularly wrinkled body surface.   Description. Holotype. complete, 6 mmlong and 1 mmwide for 33 chaetigers. A moderately large species of 5–12 mmlength and 0.5–1 mmwidth. Number of chaetigers 26–33 ( Fig. 5A). Color in alcohol white to a light tan. Body sometimes expanded in anterior region to about chaetiger 12.   TABLE 3.Distribution of scalibregmatid species known for the Southern Ocean, (?) depth record uncertain (after Blake 1981; Schüller & Hilbig 2007).    Species Distribution Depth (m)   Ascleirocheilus ashworthiBlake, 1981 off Elephant Island west of Antipodes Island 223–2892 384–397    Hyboscolex equatorialisBlake, 1981 Ecuador, Peru South Sandwich Trench intertidal 2258–2313    Axiokebuita millsiPocklington & Fournier, 1987 South Sandwich Trench 773–752    Axiokebuita minutaHartman, 1967 South Shetland Islands South Orkney Islands Antarctic Peninsula Ross Sea Weddell Sea off Elephant Island South Sandwich Trench 300 593–598 412 589–608 1122–4069 2889–2892 2258–2313    Oligobregma blakeiSchüller & Hilbig, 2007 off Elephant Island 2889–2892    Oligobregma collare(Levenstein, 1975) Drake Passage Ross Sea Weddell Sea Bellinghausen Sea 2889–3806 6070 (?) 1622–4575 2562    Oligobregma hartmanaeBlake, 1981 Weddell Sea 585    Oligobregma notialeBlake, 1981 Antarctic Peninsula Drake Passage Weddell Sea Budd and Knox Coasts, East AntarcticaRoss Sea 18–97 3683–3690 28 909–923    Oligobregma pseudocollareSchüller & Hilbig, 2007 South Sandwich Trench off Elephant Island Weddell Sea 773–752 2889–2892 3049–3050    Oligobregma quadrispinosaSchüller & Hilbig, 2007 Drake Passage Weddell Sea South Sandwich Trench 3683–3690 3049–4069 2258–2313   Pseudosaclibregma  bransfieldium(Hartman, 1967) Antarctic Peninsula Bransfield Strait Weddell Sea off Elephant Island Drake Passage South Sandwich Trench Ross Sea 335 769 412–2086 2889–2892 3683–3690 773–752 916    Pseudoscalibregma papilia  sp. nov. Ona Basin off Elephant Island South Sandwich Trench 3683–3690 2889–2892 2258–2313    Pseudoscalibregma ursapiumBlake, 1981 Ross Sea Drake Passage off Elephant Island Weddell Sea 2143–2154 3683–3690 2889–2892 3049–3050    Scalibregma inflatumRathke, 1843 cosmopolitan intertidal-abyssal    Sclerocheilus antarcticusAshworth, 1915 Antarctic Peninsula Bransfield Strait 45–90 210–426   FIGURE 5.  Pseudoscalibregma papilia  sp. nov.A. entire animal, B. anterior segments, C. posterior parapodium, D. furcate chaetae, E. pygidium. Prostomium with two spherical lobes anterolaterally; no eyes, nuchal organs not apparent. Peristomium a single achaetous ring, well developed ( Fig. 5B). Body surface almost smooth, sometimes irregularly wrinkled, a scheme in annulation not apparent. Anterior parapodia with reduced parapodial lobes, dorsal and ventral cirri, these rapidly increasing in size in median region; posterior dorsal and ventral cirri of large size, almost foliose, ventral cirri larger than dorsal ones; interramal sense organs missing ( Fig. 5C). All parapodia with simple chaetae; furcate chaetae with unequal tynes covered by fine hairs, present from chaetiger 2 ( Fig. 5D). Pygidium terminal, formed by a ring of large tubercles carrying cirri of different lengths ( Fig. 5E).   Remarks.The species is most similar to  Pseudoscalibregma bransfieldium( Hartman, 1967)which is also very common in the Southern Ocean ( Blake 1981). The two species have in common the moderately large size and the lack of a schematic annulation (unlike  P. ursapiume.g.which is covered by prominent tubercles).  Pseudoscalibregma papilia  sp. nov.can easily be distinguished from  P. bransfieldiumby the lack of a prominent nuchal organ dorsally on the prostomium, the distinctly spherical form of the anterolateral prostomial lobes, and the exceptionally large size of the posterior dorsal and ventral cirri.   Distribution.Weddell Sea, AntarcticPeninsula, Drake Passage and South Sandwich Trench, 1970–3690 m