Reitter, 1879 : 546 Redescription of Heterocerus fausti Reitter, 1879, bona species (Coleoptera, Heteroceridae) Sazhnev, Alexey S. Zootaxa 2018 2018-06-28 4441 3 597 600 Reitter, 1879 Reitter 1879 [151,540,1886,1912] Insecta Heteroceridae Heterocerus GBIF Animalia Coleoptera 0 597 Arthropoda species fausti     Heterocerus fausti  Reitter, 1879: 546  stat. nov.     Typelocality. Azerbaijan: Baku.   Typematerial examined. Syntypes: “  Heterocerus FaustiRtt” [Reitter's handwriting], “Caucasus” [red label], “к. А. Яковлева” [cоllеctiоn оf А. Jаcоvlеv—in Russiаn], 2♂♂ (ZISP); “Bаku Fаust.”, gоldеn squаrе plаtе, “  Heterocerus FaustiReitt.” [Reitter's handwriting], 2♀♀ [one specimen without head] (ZISP).   Additional material examined.9 exs.  TURKMENISTAN.  BalkanRegion(  Velayat): Uzun-Ada(now the territory of Hazar Nature Reserve),  15.04.1895, leg. Varentsov, 7 exs. ( ZISP); Turkmenbashi ( Krasnovodsk), 0 6.1900, leg. P. Maksimovitsh, 1 ex. ( ZISP); Esenguly, 3.07.1932, unknown collector, 1 ex. (IBIW).   Diagnosis.Body size 3.8–5.0 mm. Body moderately elongated, dorsal-ventrally compressed ( Fig. 1). Males and females do not differ. Aedeagus as in Fig. 3.   FIGURE 1–4.  Heterocerus faustiReitter, 1879, 1—male from Turkmenistan, Uzun-Ada (photo by A.V. Kovalev). Metatarsomers  Heterocerusspp. 2A—  H. parallelusGebler, 1830, South Kazakhstan; 2B—  H. fausti, Turkmenistan; 2C—  H. fossorKiesenwetter, 1843, Russia, Dagestan; 2D—  H. flexuosusStephens, 1828, Kazakhstan; 2E—  H. persicusMascagni, 1989, Iran. Male genitalia of  Heterocerus fausti, 3A—penis, 3B—tegmen. Scale bar—0.5 mm. Tegmens  Heterocerusspp. 4A—  H. parallelus, South Kazakhstan; 4B—  H. fausti, Caucasus; 4C—  H. fossor, Russia, Dagestan; 4D—  H. flexuosus, European part of Russia (Litovkin et al., 2013), 4E—  H. persicus, Iran. Scale bar—0.5 mm.   FIGURE 5.Distribution of  Heterocerus faustiReitter, 1879. Red circles—studied material.   Heterocerus faustiis similar in body form and drawing of elytra on  Heterocerus parallelusbut can be easily distinguished by the shape of its aedeagus ( Fig. 4A–B) and smaller size.   Redescription. Color.Ground color is yellow or red-yellow; a combination of black-brown blotches on the pronotum, and brown blotches of elytra and abdomen form a characteristic pattern. Head and labrum are black-brown. Antennae completely yellow. Pronotum is black-brown with a more or less wide yellow edging on the sides and narrow yellow stripe along the front edge. Scutellum is black-brown. Drawing of elytra characteristic of  flexuosusspecies-group sensu Charpentier (1965). Yellow pattern of each elytron normally forms edging, S-shaped basal mark, M-shaped medial mark, apical spot and suture stripe. Dark pattern of the elytra brownish almost blending with the yellow background. Prosternum black-brown with yellow episternae and yellowish front margin. Mesepisternum and metepisternum blackbrown. Abdomen black-brown with yellow pattern in form of edging. Legs, except dark metacoxa, yellow.  Head.Clypeus transversal, widely incised on the front edge. Antennae 11-segmented with 7-segment club. Hairs whitish. Puncturing uniform.  Thorax.Pronotum not bordered basally; slightly narrower or comparable in width with base of elytra in females, and slightly wider than elytra base in males. Hairs on the disc are double: comprise dense, short, pressed, and long and standing hairs; whitish. Puncturing very dense, fine and uniform. Elytra covered with erect hairs; hair length is comparable with the length of few first spines of mesotibia; hairs slightly longer along sides and near the elytral apex; hairs whitish. Legs of structure typical for family, covered with whitish hairs. Claws of tarsus noticeably longer or equal to the last tarsomere ( Fig. 2).  Abdomen. Abdomen covered with whitish hairs. Postmetacoxal lines absent.   Remarks.The records of  Heterocerus faustifor Uzbekistan( Jacobson, 1913; Zaitsev, 1946) are not confirmed. Specimen from Uzbekistan(Qamashi) signed of F.A. Zaitsev as “  Heterocerus fausti” is  Heterocerus heydeniKuwert, 1890.   Ecology.The long claws are probably an adaptation to the psammophilous way of life.   Distribution.Pre-Caspian Region including Caucasus: Azerbaijan, West Turkmenistan( Fig. 5). 1977784429 [151,503,1950,1973] Azerbaijan Baku 0 597 1 Baku holotype 1977784428 1895-04-15 ZISP Varentsov Turkmenistan Velayat Hazar Nature Reserve Uzun-Ada 1 598 1 Balkan 1977784427 ZISP P. Maksimovitsh Turkmenistan Krasnovodsk 1 598 1 Balkan