Westwood, 1845 : 7 Thomson 1856 : 8 Parry 1864b : 99 Harold 1868 : 966 Parry 1875 : 22 Felsche 1898 : 78 Van Roon 1910 : 53 Boileau 1913 : 265 Didier 1937 : 30 Didier & Séguy 1953 : 183 Benesh 1960 : 29 Bomans 1986 : 9 Bomans 1988 : 8 Cammaerts & Bomans 1997 : 12 Maes 1992 : 41 Moore & Cassis 1992 : 11 Krajcik 2001 : 13 Krajcik 2003 : 84 Bartolozzi et al . 2011 : 46 Albers, 1883 : 226 Masters 1888 : 301 Boileau 1898 : 432 Felsche 1898 : 78 Van Roon 1910 : 52 Kriesche 1922 : 133 Nagel 1928 : 295 Didier & Séguy 1953 : 182 Benesh 1960 : 26 De Lisle 1964 : 4 Bomans 1985 : 357 Bomans 1988 : 8 Maes 1992 : 39 Moore & Cassis 1992 : 10 Cammaerts & Bomans 1997 : 13 Mizunuma 2000 : 85 Krajcik 2001 : 11 Krajcik 2003 : 80 Fujita 2010 : 386 Bartolozzi et al . 2011 : 45 Kriesche, 1922 : 134 Didier & Séguy 1953 : 181 Benesh 1960 : 26 Maes 1992 : 39 Moore & Cassis 1992 : 10 Krajcik 2001 : 11 Krajcik 2003 : 80 Bartolozzi et al . 2011 : 45 Revision of the Australian species of Figulus MacLeay, 1819 (Coleoptera: Lucanidae) Monte, Cinzia Zilioli, Michele Bartolozzi, Luca Zootaxa 2016 4189 3 447 484 6HZVJ Hope in Westwood, 1845 Hope in Westwood 1845 [151,688,691,718] Insecta Lucanidae Figulus Animalia Coleoptera 26 473 Arthropoda species sulcicollis     Figulus sulcicollisHope in  Westwood, 1845: 7, 26;  Thomson 1856: 8;  Parry 1864b: 99;  Harold 1868: 966;  Parry 1875: 22;  Felsche 1898: 78;  Van Roon 1910: 53;  Boileau 1913: 265, 266;  Didier 1937: 30;  Didier & Séguy 1953: 183;  Benesh 1960: 29;  Bomans 1986: 9( lectotypedesignation);  Bomans 1988: 8;  Cammaerts & Bomans 1997: 12–13, 15–19;  Maes 1992: 41;  Moore & Cassis 1992: 11; Araya 1998: 10;  Krajcik 2001: 13;  Krajcik 2003: 84;  Bartolozzi et al. 2011: 46; Maes 2014; Schoolmeesters 2015.    Figulus mento  Albers, 1883: 226;  Masters 1888: 301;  Boileau 1898: 432;  Felsche 1898: 78;  Van Roon 1910: 52;  Kriesche 1922: 133–136;  Nagel 1928: 295;  Didier & Séguy 1953: 182;  Benesh 1960: 26;  De Lisle 1964: 4;  Bomans 1985: 357;  Bomans 1988: 8;  Maes 1992: 39;  Moore & Cassis 1992: 10;  Cammaerts & Bomans 1997: 13;  Mizunuma 2000: 85;  Krajcik 2001: 11;  Krajcik 2003: 80;  Fujita 2010: 386;  Bartolozzi et al. 2011: 45; Maes 2014; Schoolmeesters 2015. New synonymy.    Figulus mento foveatus  Kriesche, 1922: 134;  Didier & Séguy 1953: 181;  Benesh 1960: 26;  Maes 1992: 39;  Moore & Cassis 1992: 10;  Krajcik 2001: 11;  Krajcik 2003: 80;  Bartolozzi et al. 2011: 45; Maes 2014; Schoolmeesters 2015. New synonymy.   Type material.  Figulus sulcicollis:lectotype (designation by Bomans 1986): “P. Ess. [= Port Essington] [handwritten]”, “Type / Hope / Cat. Lucan. / 1845 p.26 / Collection Hope Oxon. [p, handwritten]”, “TYPE COL: 328 1/ 2/  Figulus/  sulcicollisHope/ (Westw.) / HOPE DEPT. OXFORD [printed, handwritten]”, [OUMNH]; paralectotype (1 specimen): “  sulcicollisW [handwritten]”, “M. I. [= Melville Island] [handwritten]”, “Type / Hope / Cat. Lucan. / 1845 p.26 / Collection Hope Oxon. [printed, handwritten]”, “TYPE COL: 328 2/ 2/  Figulus/  sulcicollisHope/ (Westw.) / HOPE DEPT. OXFORD [printed, handwritten]”, [OUMNH].   Diagnosis.  Figulus sulcicollisis similar to  F. rossiin general morphology.  Figulus sulcicollismainly differs by the more punctate head and prothorax and the much smaller size of the punctures on the elytra, which are suboval instead of subcircular.    Redescription.Measurements:size range ( n= 243): TL: 10.75–17.75 mm; PL: 2.71–5.08 mm; EL: 6.16– 9.83 mm; PW: 3.46–6.21 mm; EW: 3.50–6.29 mm.  Male( Fig. 25). Body elongate, with subparallel sides; black to dark brown or reddish brown, glossy.  Head:transverse, about 3 times wider than long, slightly depressed. Frontoclypeal region slightly concave at the anterior margin; minutely punctate, with a small, medial elevation and 2 large, glossy areas situated before the eyes. Labrum broad and short with long, golden setae. Vertex inclined anteriorly, slightly concave in the middle, with 2 oval and transversal elevations moderately developed between eyes. Two moderately developed longitudinal carinae, are present along the inner margin of eyes. Eyes large, divided into 2 parts by the canthi, the upper part smaller than the lower, with distinct facets. Canthus protruding, largely rounded laterally, evenly rounded or more angled behind; upper surface slightly depressed and almost smooth, with a few minute punctures on the lateral margin. Head surface densely punctate with small punctures, punctures larger and sparse in the middle. Mandibles slightly shorter than head, strong and asymmetrical, outer margins rounded, apices slightly acute and curved upwards; upper surface slightly elevated at base. Internal margin of the right mandible with a moderately developed basal tooth and a medial strong tooth, with blunt apex directed inwards. Internal margin of the left mandible with a moderately developed basal tooth, a well developed medial tooth, and a preapical evident angulation; surface densely and minutely punctate. Antenna with 10 antennomeres, with scape slightly shorter than the remaining antennomeres; club of last three antennomeres, which are flatter and broader than the remaining antennomeres of the funicle. Mentumsubrectangular ( Fig. 26), concave medially; anterior margin slightly bilobed; base coarsely punctate; sides slightly rounded and elevated at anterolateral margins, where a tubercle is present; surface punctate and rugose with long setae medially and distally.   FIGURES 25‒29. Figulus sulcicollisHope in Westwood.25, dorsal view of adult (scale bar: 5 mm); 26, mentum; 27, male genitalia, dorsal view (scale bar: 0.5 mm); 28, male genitalia, lateral view; 29, apex of flagellum (scale bar: 200 µm).  Thorax: pronotum slightly wider than long with subparallel sides; anterior margin feebly convex slightly or moderately elevate, with a small, blunt, median tubercle; posterior margin slightly bisinuate; anterior angles produced and slightly acute; posterior angles rounded with slightly crenulated rim. Deep longitudinal medial groove on pronotal disc, with dense, large, circular punctures. Pronotal surface glossy, with dense, small punctures, larger, deeper, and denser at sides and with patch of large, close punctures at the anterolateral margin, each puncture with a short, erect, golden seta in the middle. Scutellum long, wedge-shaped, moderately narrow; surface smooth and glossy. Elytra about twice as long as pronotum. Base concave, subparallel sides, apex regularly rounded. Humeral angles acute, with a small, acute tubercle. Elytral surface glossy, with 9 visible subparallel striae, the dorsal striae deeper than the lateral striae; each stria formed by a row of small suboval punctures well separated from each other, except for stria 1, where the punctures are confluent. Stria 1 is the only stria reaching apex; on the declivity, stria 2 curves outwards and deepens, sometimes converging with stria 9; striae 3‒4 and 5‒6 converge at the beginning of the declivity; stria 6 starts just after base; striae 7 and 8 narrow, formed by a row of suboval punctures smaller than those of the other striae. Dorsal interstriae slightly convex, much broader than the lateral interstriae, densely and finely punctate; interstria 2 slightly larger at declivity, where it is strongly punctate. Elytral lateral margins strongly punctate, with short, erect, red setae; elytral apex strongly punctate. Wings present. Legs: thin and straight. Protibiae with strong, elongate apical fork, and a long, strong spur internally at apex; lateral margin with 3‒4 small teeth, first 3 teeth subequal. Mesotibiae with 4 long, acute, apical teeth, and 2 apical spurs, 1 larger than the other; lateral margin with strong, submedial tooth, preceded by 2‒3 smaller teeth. Metatibiae with 4 acute apical teeth, and 2 long apical spurs, 1 larger than the other; lateral margin with strong submedial tooth, preceded by a smaller tooth. Prosternum subpentagonal in shape behind the coxae; surface with large and deep punctures at anterior edge and with smaller punctures medially and at posterior edge. Metasternum smooth on disc, with a long medial groove, deep at posterior edge, strongly punctate at sides.  Abdomen: first abdominal ventrite with some horse-shoe shaped, superficial hollows each with a short, golden seta. Ventrites 2‒4 with some small and elongate punctures along the anterior margin and some circular punctures at sides, punctures much finer and denser medially. Last ventrite with 2 feebly depressions at sides near the posterior margin; surface with some punctures.  Male genitalia: penis tubular, shorter than parameres, completely covered by a dorsal, translucid, membranous extension of the basal piece; slightly curved in lateral view, strongly bent at base close to articulation with basal piece ( Figs. 27–29). Permanently everted internal sac drawn out into a long and broad, colorless or yellow flagellum tapering towards apex, the latter slightly trilobate ( Fig. 29), the median lobe longer; edge of flagellum serrulate towards apex. Parameres about as long as basal piece, broad and spatulate in dorsal view, with marginal fringe of setae entirely enclosed in the cuticle (not represented in figure). Basal piece in dorsal view with sides slightly converging toward apex, ventrally largely membranous, dorsally sclerotized; with a narrow, elongate sclerotic area along midline. Two long, slender struts run along basal piece from penis articulation, almost reaching the base of basal piece ( Figs. 27–28).  Female.All external morphological characters as in male.  Female genitalia: hemisternites elongate, apically with long, golden setae surrounding the styli; moderately broad, narrower apically; outer edge almost straight or slightly concave medially; medial edge moderately convex; styli slender with apex oblique directed outwards, slightly longer than wide. Bursa copulatrix represented by a slight expansion of the wall of spermathecal duct near junction with the median oviduct. Spermathecal duct short, ending in a bulbous, pear-shaped expansion that is pigmented distally.  Intraspecific variation.The examined specimens do not show significant morphological variability. For the body size see the “Measurements” section.     Additional material examined. Australia[state unknown]: 7 specimens,  AustraliaOrient., v. Müller 2-96, Muséum Paris ( MNHN); 2 specimens,  North Australia, K 293240 and K 293241 (AMS).    NORTHERNTERRITORY: 1 specimen, Adelaide River, 91-49, 820, BMNH(E) # 607430 ( NHM);  4 specimens,  Adelaide River, J.J.Walker, Muséum Paris1952 Collection R.Oberthür ( MNHN);  1 specimen, Adelaide River,  I.1942, J.Farr, COL-85929 ( NMVM); 1Ƌ, 1♀, 16 specimens, Alligator Inn, 12°40'S, 132°30'E,  7–9.VI.1979, rainforest, G.B.Monteith& D.Cook( QMB);  1 specimen, Alyangula, 13°51'S, 136°25'E,  20–24.II.2005, 20 m, woodland, D.Cook, 11759 ( QMB);  1 specimen,  Black MountainRd Kuranda,  8.IV.1988, J.Hasenpusch, Graeme Krake Collection, donated  Feb. 2007, COL-86869 ( NMVM);  4 specimens, Coburg Peninsula, Black Point, 11°09'S, 132°09'E,  15–23. II.1977, ex light water trap, T.A.Weir( ANIC);  1 specimen, Coburg Peninsula, Smith Point, 11°07'S, 132°08'E,  17–18. II.1977, R.Lewis( ANIC);  1 specimen, Elcho Island,  16.III.1976, M.E.Farr( ANIC); 2 specimens, Escarpment, W Arnhem Land, E Alligator Road, 23.VII.1974, F.Morris, COL-85961 and COL-85962 (NMVM);  1 specimen, Darwin,  18.VI.1913, F.E.Wilson CollectionCOL-85861 ( NMVM);  1 specimen, Darwin,  26.I.1989, Graeme Krake Collection, donated  Feb. 2007, COL-86868 ( NMVM);  1 specimen, Darwin,  31.III.1913, G.F.Hill1914-441, BMNH(E) # 607436 ( NHM); 1 specimen, Darwin, R.I.Sc.N.B. I.G. 18.674, Collection et det. A.Janssens (IRSNB);  1♀, Darwin, Lee Point,  23.V.1992, S.K.Smith, R.deKeyzer Insect Collection, K293251 (AMS);  1 specimen,  10 kmS of Darwin, Holmes Jungle Forest Resort, 12°26'S, 130°56'E,  23.X.1972, E.B.Britton( ANIC);  1 specimen, King River,  25.XII.1915, P.Cahill, COL-85172 ( NMVM);  7 specimens, North Point Kapalga, 12°25'S, 132°22'E,  19.VI.1979, G.B. Monteith& D.Cook( QMB);  6 specimens Rimbija Island,  Wessel IslandRanges, 11°01'S, 136°45'E,  21.I.–31. II.1977, ex light water trap, Farrow& Dowse( ANIC).   QUEENSLAND:1Ƌ, 1 specimen, N Queensland, No.154 and No.155, 99:286 Ƌ, R.G.Wind, S.G.Watkins Collection, donated 2001 ( ANIC);  1 specimen, Queensland, Muséum Paris1952 Collection R.Oberthür ( MNHN); 1Ƌ, N Queensland, Lyons Lookout,  10-12.VIII.1988, ex Larva, G.Minet, 12 0 28 ( NHM);  1 specimen, Airlie,  4–6.VIII.1988, G.Minet12027 H.E.Bomans, BMNH(E) 1999-248, 412027 (handwritten) ( NHM);  1 specimen, 8 kmW by N of Bald Hill McIllwraith Range, 13°45'S, 143°22'E,  27.VI–12.VII. 1989, 500 m, mango tree site, ex rotten log closed forest, T.A.Weir( ANIC); 1 specimen, Binna Burra, 6.38, C.E.Clark, C.E.Clark Collection B.M. 1957-24, BMNH(E) # 607751 (NHM); 1 specimen, Cairns, S.M.Parlett, F.E.Wilson Collection, COL-85163 (NMVM);  1♀, Cairns,  20.XI.1982, Graeme Krake Collection, donated  Feb. 2007, COL-86870 ( NMVM); 1Ƌ, Cairns,  XII.1970, 2045 ( MSNM); 1Ƌ, Cairns, Mount Whitfield,  7.VII.1984, R.deKeyzer Insect Collection, K293260 (AMS); 1 specimen, CapeTribulation area, 16°03'- 16°05'S, 145°28'E, under bark in rotten logs, J.F.Lawrence( ANIC);  4 specimens, CapeTribulation area, 16°03'- 16°05'S, 145°28'E,  21–28.III.1984, A.Calder& T.Weir( ANIC);  1 specimen, CapeTribulation, Emmagen Creek,  8–9.VII.1987, R.deKeyzer, R.deKeyzer Insect Collection K293250 (AMS);  2 specimens, Cape York, Lockerbie Area,  13–27.IV.1973, G.B.Monteith( QMB);  9 specimens, 3 kmE of Lockerbie, Cape York,  16-20.IX.1974, rainforest, G.B.Monteith( QMB);  1 specimen, 3 kmE of Lockerbie, Cape York,  13– 27.III.1987,  Pyrethrumon logs, RF, G.B.Monteith( QMB);  4 specimens, 3 kmE of Lockerbie, Cape York,  19– 23.III.1987, rainforest pitfall traps, G.B.Monteith( QMB);  4 specimens, Cape York, 4 kmE of Lockerbie,  30.I– 4.II.1975, rainforest, G.B.Monteith( QMB);  1 specimen, Cardstone,  7–9.I.1962, Carne& Britton( ANIC);  1 specimen Cardstone,  20.I.1967, K.Hyde( ANIC);  1♀, 1 specimen, Carrol Creek,  4 kmNW of Hopevale,  11.VII.1976, G.B. & S.R Monteith( QMB); 1 specimen, Coen, COL-86042 (NMVM);  6 specimens,  30 kmNE of Coen, Leo Creek Road, McIlwraith Range,  29.VI–4.VII.1976, approximately  500 m, G.B. & S.R. Monteith( QMB);  2 specimens, Clump Point,  21.IX.1951and 25.X.1951, J.O.Campbell, F.E.Wilson Collection, COL-85168 and COL-85167 ( NMVM);  1 specimen Cooktown,  27.VI.1961, C.Oke, COL-85917 ( NMVM);  1 specimen, Cooktown, ex Möllenkamp( MNHN);  2 specimens, Cooktown,  VIII.1994, Meek, Muséum Paris1 952 Collection R.Oberthür ( MNHN);  1 specimenDunk’s Island, Elgner1910 ( ANIC);  1 specimen, Edmonton,  VII.1945, A.Johnson, COL-85900 ( NMVM);  1 specimen, Endeavour River,  6.I.08, Nat. Mus. Victoria C.French’s Collection, COL-86048 ( NMVM).  1 specimen, Gap Creek,  5 kmESE Mount Finnigan, 15°50'S, 145°20'E,  13–16.V.1981, A.Calder( ANIC);  1 specimen, Garradunga, Polly Creek,  15.VI.2001, J.Hasenpusch, R.deKeyzer Insect Collection, K293255 (AMS);  1 specimen, Garradunga, Polly Creek,  16.I.2008, mv light, J.Hasenpusch, R.deKeyzer Insect Collection, K293254 (AMS);  1 specimen, Gordonvale,  26.II.1970, mv light, R.Hardie( ANIC);  2 specimens, Gordonvale,  28.XI.1970, mv light, R.Hardie, Universityof New England Collection, donated 1983 ( ANIC);  2 specimens, Green Island,  8–15.VIII.1982, under bark and in rotten wood, J. & N.Lawrence( ANIC);  2 specimens, Hammond Island, 1 kmW of village, 10°34'S, 148°13'E,  20.V.2003, G.B.Monteith& M.Ero, hand collected, 11324 ( QMB);  1 specimen, Hinchinbrook Island, Gayundah Creek,  7–15.XI.1984, 10 m, Monteith, Cook, & Thompson( QMB);  1 specimen, 14 kmW by N of Hope ValeMission, 15°16'S, 144°59'E,  8–10.XII.1980, T.Weir.( ANIC);  2 specimens, Innisfail,  18.XII.1995, J.Hasenpusch, R.deKeyzer Insect Collection, K293256 and K 293257 (AMS); 3 specimens, Iron Range, W Claudie Range, 3–10.XII.1985, 50 m, rainforest, G.B.Monteith & D.Cook (QMB);  1 specimen, Iron Range,  28.X.1991, J.Hasenpusch, R.deKeyzer Insect Collection, K293263 (AMS);  1 specimen, Iron Range, Gordon’s Mine Area,  12–18.II.1976, rainforest, G.B.Monteith( QMB);  1♀, Kuranda,  10.XI.1972, Collection H.Bomans, H.E.Bomans BMNH(E) 6527, 406527 (handwritten) ( NHM);  1 specimen, Kuranda,  20.I.1995, J.Hasenpusch, R.deKeyzer Insect Collection, K293258 (AMS);  2 specimens, Kuranda,  335 mtop of the range 19 Butler Drive, 16°48'S, 145°38'E,  1–15.I.2007and 1–15.I.2008, D.C.F.Renz( ANIC);  2 specimens, Kuranda,  20–21.III.1952, C.Oke, COL-8 5919 and COL-85925 ( NMVM);  1 specimen,  10 mW of Kuranda, Parmentier Creek, Mareeba Road.,  26.II.1970, mv light, R.Hardie, Universityof New England Collection, donated 1983 ( ANIC);  1 specimen, 13 kmW Kuranda,  7.XII.1982, J.T.Doyen( ANIC); 1Ƌ, Lions Lookout,  10– 12.VIII.1988, G.Minet, ex larva Paris, 12028, H.E.Bomans, BMNH(E) 1 999-248, 412028 (handwritten) ( NHM);  1♀, Little Mulgrave Valley,  16. II.1969, R.Parrott( ANIC);  1 specimen, Mulgrave Road, Goldsborough Valley,  4.IX.1994, M.&G. DeBaar( ANIC);  1 specimen, Mackay,  5–20.V.1971, W.Vestjens( ANIC);  2 specimens, Moses Creek, 4 kmN by E of Mount Finnigan, 15°47'S, 145°17'E,  14–16.X.1980, T.Weir( ANIC);  1 specimen, Portland Roads, via Iron Range,  3–10.XII.1986, rainforest, Monteith, Cook, & Yeates( QMB);  1 specimen, Shiptons Flat, 15°47'S, 145°14'E,  16–18.V.1981, A.Calder( ANIC);  4 specimens, Shiptons Flat, 15°47'S, 145°14'E,  17– 19.X.1980, T.Weir( ANIC);  4 specimens, Shiptons Flat,  35 kmS of Cooktown,  22.IV. 1982, 250 m, Monteith, Yeates, & Cook( QMB);  1 specimen, Shiptons Flat, via Helenvale,  20–27.VI.1974, rainforest, G.B.Monteith, & D.Cook( QMB);  4 specimens, Somerset, L.M.D’Albertis1875 Muséum Paris1952, Collection R.Oberthür ( MNHN);  2♀♀, Somerset,  I.1875, L.M.D’Albertis( MSNG);  2 specimens, Mount Tamborine,  I.1950, COL-85967 and  XII.1950, COL-85966, C.Oke( NMVM);  1 specimen, Thorton Ranges,  11kmE of Daintree, 16°16'S, 145°25'E,  21.XI.1981, J.Balderson( ANIC);  1 specimen, Thursday Island, Green Hill, 10°35'S, 142°13'E,  21.V.2003,  Pyrethrumtrees & logs, G.B.Monteith, 11334 ( QMB);  2 specimens, Torres Strait, Badu Island,  16– 22.XII.1976, H.Heatwole, K 293238 and K 293239 (AMS); 2ƋƋ, Torres Strait,  Cape York, 11.81, Dr. O.Finsch (MFN); 1 specimen, Torres Strait, Darnley Island,  14.VII.1974, H.Heatwole, K 293236 (AMS);  2 specimens Torres Strait, Eet Hillvicinity, Moa Island(= Banks Island)  9–13.VI.1977, G.B.Monteith& D.Cook( QMB);  2 specimens, Torres Strait, Mabuiag Island,  VIII.1924, C.T.MacNamara, ex Collection Australian Museum( ANIC);  1 specimen, same locality and data, ( IRSNB);  1 specimen, Torres Strait, Murray Island,  20.VII.1974, sweep of strand vegeta, H.Heatwole& E.Cameron, K 293237 (AMS);  1 specimen, Torres Strait, Murray Island,  VII–VIII.1974, H.Heatwole, K 293232 (AMS);  1 specimen, Torres Strait, Princeof Wales Island,  6–11.II.1995, under logs, 12, H.Heatwole, K 293233 (AMS);  10 specimens, Torres Strait, Princeof Wales Island, 10°42.3'S, 142°13.5'E,  3– 7.I.2008, camp.,  10 m, G.Monteith& K. Aland, 15460 ( QMB);  1 specimen, Torres Strait, Sue Island,  11.VII.1974, H.Heatwole, K 293234 (AMS);  5 specimens, Torres Strait, Tawan Island, D’Albertis1876, Muséum Paris1952, Collection R.Oberthür ( MNHN);  1♀, 3 specimens, Torres Strait, Tawan Island,  XI.1876, D’Albertis( MSNG);  1 specimen, Torres Strait, YorkIsland,  13.VII.1974, H.Heatwole& E.Cameron, K 293236 (AMS); Mount Tozer, 12°43'S, 143°17'E,  5–10.VII.1986, rotten log, T.Weir& A.Calder( ANIC);  2 specimens, 11 kmENE of Mount Tozer, 12°43'S, 143°18'E,  11–16.VII.1986, T.Weir& A.Calder( ANIC);  8 specimens, Warraber (Sue) Island, 10°12'S, 145°49'E,  4–6.XII.1977, E.D.Edwards( ANIC);  3 specimens, Warraber (Sue) Island, 10°12'S, 145°49'E,  19–25.I.1978, R.Lewis( ANIC);  1 specimen, Warraber (Sue) Island, 10°25'S, 142°49'E,  6.XII.1977, R.Lewis( ANIC);  2 specimens, Mount Webb National Park, 15°04'S, 145°07'E,  28–30. IX.1980, T.Weir( ANIC);  2 specimens, Mount Webb National Park,  50 kmN of Cooktown,  11–14.VII.1976, G.B. & R.S. Monteith( QMB);  1 specimen, Weipa,  3–6.II.1976, G.B.Monteith( QMB);  1 specimen, Weipa, Hibberd Point,  3–5.II.1976, rainforest, G.B.Monteith( QMB);  1 specimen Mount Welsh, near Biggenden,  17.X.1975, approximately  1000 m, H.Frauca( ANIC);  1 specimen, Yeppoon,  3.III.1935, S.R.E.Brock Collectiondonated to  ANIC1987( ANIC).   SOUTH AUSTRALIA: 1 specimen,  South Australia,  20.VIII.1950, COL-85956 ( NMVM).   VICTORIA: 1 specimen, Grampians,  3.I.1938, C.Oke, COL-86032 ( NMVM); 1 specimen, Hattah, C.Oke, COL-85921 (NMVM).    WESTERNAUSTRALIA: 2 specimens, Prince Frederick Harbour, 15°00'S, 125°21'E, “ Marun” CALM Site8/4,  6–11.VI.1988, closed forest and margin, I.D.Naumann( ANIC);  2 specimens, Carnarvon,  WesternAustralia,  IV.1958, Snell, COL-85955 and COL-85957 ( NMVM).    COCOS(KEELING) ISLAND: 1♀, West Island, Cocos (Keeling)Island,  23.V.1952, 8630, H.E.Bomans,  BMNH(E) 1999-248, 408630 (handwritten) ( NHM); 1 specimen, West Island, Cocos (Keeling)Island,  X.1962, 8631, H.E.Bomans, BMNH(E) 1999-248, 408631 (handwritten) (NHM).   Distribution. Australia:  NorthernTerritory, Queensland, South Australia, Victoria,  Western Australia,  Cocos (Keeling) Islands; New Guinea: Indonesian Papua, Papua New Guinea; Raja Ampat Archipelago: Misool Island; Moluccan Archipelago: Bacan Islands ( Bartolozzi et al.2011). The few records from South Australia, Victoria, and  Western Australiaprobably need confirmation.   Natural history data.The larvae live in and under rotting wood and have been described and illustrated by Cammaerts & Bomans (1997). According to label data, this species has been collected in habitats ranging from close forest and ist margins (WA) to open forest (N Queensland). The material we examined includes specimens found under bark or in rotten logs and by sweeping strand vegetation. Most specimens were collected at mv light or light traps. Two specimens, one of which was labeled “RF” (= rainforest), were sampled by Pyrethrumfogging on trees and logs, and four other specimens were from rainforest pitfall traps. The elevation mainly ranges from 10 m(Torres Strait, Prince of Wales Island, I.2008) to 500 min rainforest, but one specimen was collected in montane forest at about 1000 m(Mount Welsh Queensland, X.1975). For Australia, the host plant(s) are unknown. Cammaerts & Bomans (1997)listed the following host plants in New Guinea:  Excoecaria agallocha(Euphorbiaceae),  Terminalia catappa(Combretaceae), and  Leucaena leucocephala(Fabaceae).   Remarks.This species is as common and widespread as  Figulus regularis.We found one specimen in the MNHN collection labeled “New Zealand” (Belligny, 3-43), but we consider this locality as erroneous, as the genus  Figulusis not known from this country.  Kriesche (1922)described the subspecies foveatusof  Figulus mentofrom northern Queensland. The type specimens in Hannover Museum were destroyed during World War II ( Krajcik 2003). Since we never found any specimen corresponding to this subspecies, nor to  F. mento mento, coming from Australia, we believe that the record from Australia is erroneous and that the species should be removed from any checklists of Australian Lucanidae. We also consided  F. mento mento(recorded from New Guinea: Papua New Guinea; Moluccan Archipelago: Aru Islands [ Bartolozzi et al. 2011]) as a synonym of  F. sulcicollis( new synonymy). The species was also recorded from the Philippines (Luzon, Mindanao) and Palau ( De Lisle 1964). According to Cammaerts & Bomans (1997), the species does not occur in Aru and in Palau. 1324963954 [199,1048,1196,1221] Australia Australia 28 475 7 1324963987 Australia MNHN 28 475 2 MNHN 1324963968 [636,1255,1231,1256] Australia North Australia 28 475 1 North Australia 1324963994 BMNH, NHM Australia Adelaide River 28 475 1 NORTHERN TERRITORY 1324963982 Adelaide River, J. J. & Museum Paris & Collection R. Australia Adelaide River 28 475 4 MNHN 1324963984 1942-01 NMVM J. Farr Australia Adelaide River 28 475 1 MNHN 1324963976 1979-06-07 1979-06-09 1979-06-07 QMB G. B. Monteith & D. Cook Australia -12.666667 Alligator Inn 1292 132.5 28 475 17 1 MNHN 1324963981 2005-02-20 2005-02-24 2005-02-20 QMB D. Cook Australia -13.85 Alyangula 1289 136.41667 28 475 1 MNHN 1324963957 1988-04-08 2007-02 1988-04-08 NMVM J. Hasenpusch & Graeme Krake Collection Australia Black Mountain Rd Kuranda Mountain Rd Kuranda Black Mountain 28 475 1 MNHN 1324963967 1977-02-15 1977-02-23 1977-02-15 ANIC T. A. Weir Australia -11.15 Black Point 1296 132.15 Coburg Peninsula 28 475 4 MNHN 1324963964 1977-02-17 1977-02-18 1977-02-17 ANIC R. Lewis Australia -11.116667 Smith Point 1296 132.13333 Coburg Peninsula 28 475 1 MNHN 1324964002 [757,1408,1519,1544] 1976-03-16 ANIC M. E. Farr Iceland Island 28 475 1 1324963988 [277,1185,1591,1616] 1913-06-18 NMVM F. E. Wilson Collection Iceland Darwin 28 475 1 1324963993 1989-01-26 2007-02 1989-01-26 NMVM Graeme Krake Collection Iceland Darwin 28 475 1 1324963974 1913-03-31 BMNH, NHM G. F. Hill Iceland Darwin 28 475 1 1324964018 1992-05-23 S. K. Smith & Insect Collection, K Iceland Lee Point Darwin 28 475 1 1 1324964006 1972-10-23 ANIC E. B. Britton Iceland -12.433333 Holmes Jungle Forest Resort 1293 130.93333 28 475 1 1324963996 [252,1036,1772,1797] 1915-12-25 NMVM P. Cahill Iceland King River 28 475 1 1324964013 1979-06-19 QMB G. B. Monteith & D. Cook Iceland -12.416667 North Point Kapalga 1293 132.36667 28 475 7 1324963991 1977-01-21 1977-02-31 1977-01-21 ANIC Farrow & Dowse Iceland Rimbija Island -11.016666 Wessel Island 1296 136.75 Wessel Island Ranges 28 475 6 1324964003 ANIC No. & D, R. G. Wind, S. G. Iceland QUEENSLAND 28 475 1 QUEENSLAND 1324964010 [252,1137,1916,1941] Museum Paris & Collection R. Iceland Queensland 28 475 1 Queensland 1324964001 1988-08-10 1988-08-12 1988-08-10 NHM Larva, G. Iceland Lyons Lookout 28 475 1 Queensland 1324963990 1988-08-04 1988-08-06 1988-08-04 BMNH, NHM G. Minet & H. E. Bomans Iceland Airlie 28 475 1 Queensland 1324964021 1989-06-27 1989-07-12 1989-06-27 ANIC T. A. Weir Iceland 29 476 -13.75 Bald Hill McIllwraith Range 1289 143.36667 28 475 1 Queensland 1324963989 1982-11-20 2007-02 1982-11-20 NMVM Graeme Krake Collection Iceland Cairns 29 476 1 1 Queensland 1324964005 1970-12 MSNM Iceland Cairns 29 476 1 Queensland 1324963997 1984-07-07 ANIC Insect Collection, K & J. F. Lawrence Iceland -16.083334 Mount Whitfield 1283 145.46666 Cairns 29 476 1 Cape 1324964022 1984-03-21 1984-03-28 1984-03-21 ANIC A. Calder & T. Weir Iceland -16.083334 Cape 1283 145.46666 29 476 4 Cape 1324964009 1987-07-08 1987-07-09 1987-07-08 Insect Collection K Iceland Emmagen Creek 29 476 1 Cape 1324964014 1973-04-13 1973-04-27 1973-04-13 QMB G. B. Monteith Iceland Lockerbie Area 29 476 2 Cape York 1324963999 1974-09-16 1974-09-20 1974-09-16 QMB G. B. Monteith Iceland Lockerbie 29 476 9 Cape York 1324964011 1987-03-13 1987-03-27 1987-03-13 QMB G. B. Monteith Russia Lockerbie 29 476 1 Cape York 1324963980 1987-03-19 1987-03-23 1987-03-19 QMB G. B. Monteith Russia Lockerbie 29 476 4 Cape York 1324963995 1975-01-30 1975-02-04 1975-01-30 QMB G. B. Monteith Russia Lockerbie 29 476 4 Cape York 1324963985 [673,1404,583,608] 1962-01-07 1962-01-09 1962-01-07 ANIC Carne & Britton Russia Cardstone 29 476 1 Cape York 1324964025 1967-01-20 ANIC K. Hyde Russia Cardstone 29 476 1 Cape York 1324964008 1976-07-11 QMB G. B. & S. R Monteith Russia Carrol Creek 29 476 2 1 Cape York 1324964023 1976-06-29 1976-07-04 1976-06-29 QMB G. B. & S. R. Monteith Russia 500 McIlwraith Range Leo Creek Road 29 476 6 Cape York 1324964000 21. IX. 1951 and 25. X. 1951 NMVM J. O. Campbell & F. E. Wilson Collection Russia Clump Point 29 476 2 Cape York 1324964042 [486,1277,764,789] 1961-06-27 NMVM C. Oke Russia Cooktown 29 476 1 Cape York 1324964031 Cooktown & Mollenkamp Russia MNHN 29 476 1 MNHN 1324964017 1994-08 Meek & Museum Paris & Collection R. Russia Cooktown 29 476 2 MNHN 1324964037 [432,1015,836,861] ANIC Elgner Iceland Island 29 476 1 1324964015 1945-07 NMVM A. Johnson Iceland Edmonton 29 476 1 1324964029 2008-01-06 NMVM C. French's Collection Iceland Endeavour River 29 476 1 Victoria 1324964036 1981-05-13 1981-05-16 1981-05-13 ANIC A. Calder Iceland -15.833333 Gap Creek 1284 145.33333 29 476 1 Victoria 1324964026 2001-06-15 J. Hasenpusch & Insect Collection, K Iceland Polly Creek Garradunga 29 476 1 Victoria 1324964012 2008-01-16 J. Hasenpusch & Insect Collection, K Iceland Polly Creek Garradunga 29 476 1 Victoria 1324964043 [513,1271,1015,1040] 1970-02-26 ANIC R. Hardie Iceland Gordonvale 29 476 1 Victoria 1324964016 1970-11-28 ANIC R. Hardie & University United Kingdom Gordonvale 29 476 2 1324964032 1982-08-08 1982-08-15 1982-08-08 ANIC J. & N. Lawrence Iceland Island 29 476 2 1324964020 2003-05-20 QMB G. B. Monteith & M. Ero Iceland -10.566667 Island 1297 148.21666 29 476 2 1324964048 1984-11-07 1984-11-15 1984-11-07 QMB Monteith & Cook & Thompson Iceland Gayundah Creek 29 476 1 1324964034 [249,1410,1196,1221] 1980-12-08 1980-12-10 1980-12-08 ANIC T. Weir. Iceland -15.266666 Vale 1285 144.98334 29 476 1 Vale 1324964039 1995-12-18 J. Hasenpusch & Insect Collection, K Iceland Innisfail 29 476 2 Vale 1324964024 1991-10-28 J. Hasenpusch & Insect Collection, K Iceland Iron Range 29 476 1 Vale 1324964033 1976-02-12 1976-02-18 1976-02-12 QMB G. B. Monteith Iceland Gordon's Mine Area Iron Range 29 476 1 Vale 1324964004 1972-11-10 BMNH, NHM Collection H. & Bomans, H. E. Iceland Kuranda 29 476 1 1 Vale 1324964019 1995-01-20 J. Hasenpusch & Insect Collection, K Iceland Kuranda 29 476 1 Vale 1324964007 1 - 15. I. 2007 and 1 - 15. I. 2008 ANIC D. C. F. Renz Iceland 335 -16.8 Butler Drive 1281 145.63333 Kuranda 29 476 2 Vale 1324964050 1952-03-20 1952-03-21 1952-03-20 NMVM C. Oke Iceland Kuranda 29 476 2 Vale 1324964035 1970-02-26 ANIC R. Hardie & University United Kingdom Kuranda 10 Mareeba Road. Parmentier Creek 29 476 1 1324964047 [334,1118,1556,1581] 1982-12-07 ANIC J. T. Doyen United Kingdom Kuranda 29 476 1 1324964027 1988-08-10 1988-08-12 1988-08-10 BMNH, NHM G. Minet & Paris & H. E. Bomans United Kingdom Lions Lookout 29 476 1 1324964066 [151,824,1628,1653] 1969-02-16 ANIC R. Parrott United Kingdom Little Mulgrave Valley 29 476 1 1 1324964056 1994-09-04 ANIC M. & G. DeBaar United Kingdom Goldsborough Valley Mulgrave Road 29 476 1 1324964044 [564,1195,1663,1688] 1971-05-05 1971-05-20 1971-05-05 ANIC W. Vestjens United Kingdom Mackay 29 476 1 1324964061 1980-10-14 1980-10-16 1980-10-14 ANIC T. Weir United Kingdom -15.783334 Mount Finnigan 1284 145.28334 Moses Creek 29 476 2 1324964038 1986-12-03 1986-12-10 1986-12-03 QMB Monteith & Cook & Yeates United Kingdom Iron Range Portland Roads 29 476 1 1324964054 1981-05-16 1981-05-18 1981-05-16 ANIC A. Calder United Kingdom -15.783334 Shiptons Flat 1284 145.23334 29 476 1 1324964057 1980-10-17 1980-10-19 1980-10-17 ANIC T. Weir United Kingdom -15.783334 Shiptons Flat 1284 145.23334 29 476 4 1324964051 1982-04-22 QMB Monteith & Yeates & Cook United Kingdom Shiptons Flat 29 476 4 1324964040 1974-06-20 1974-06-27 1974-06-20 QMB G. B. Monteith & D. Cook United Kingdom Helenvale Shiptons Flat 29 476 1 1324964067 MNHN L. M. D'Albertis & Museum Paris & Collection R. United Kingdom Somerset 29 476 4 Somerset 1324964041 [270,832,1916,1941] 1875-01 MSNG L. M. D'Albertis United Kingdom Somerset 29 476 2 2 Somerset 1324964055 1950-01 1950-12 1950-01 NMVM C. Oke United Kingdom Mount Tamborine 29 476 2 Somerset 1324964045 1981-11-21 ANIC J. Balderson United Kingdom -16.266666 Thorton Ranges 1282 145.41667 29 476 1 Somerset 1324964072 2003-05-21 QMB G. B. Monteith Iceland 30 477 -10.583333 Green Hill 1297 142.21666 29 476 1 1324964060 1976-12-16 1976-12-22 1976-12-16 H. Heatwole & Torres Strait Iceland Torres Strait 30 477 2 1324964064 1974-07-14 Dr. O. & H. Heatwole Iceland Torres Strait 30 477 1 Cape York 1324964049 1977-06-09 1977-06-13 1977-06-09 QMB G. B. Monteith & D. Cook Iceland Torres Strait Moa Island Eet Hill 30 477 2 Cape York 1324964058 1924-08 ANIC C. T. MacNamara & Collection Australian Museum Iceland Torres Strait 30 477 2 1324964030 1924-08 IRSNB C. T. MacNamara & Collection Australian Museum Iceland Torres Strait 30 477 1 1324964046 1974-07-20 H. Heatwole & E. Cameron Iceland Torres Strait 30 477 1 1324964028 1974-07 1974-08-31 1974-07 H. Heatwole Iceland Torres Strait 30 477 1 1324964074 1995-02-06 1995-02-11 1995-02-06 H. Heatwole United Kingdom Prince Torres Strait 30 477 1 1324964059 2008-01-03 2008-01-07 2008-01-03 G. Monteith United Kingdom 10 -10.705 Prince 129 142.225 Torres Strait 30 477 10 1324964068 [603,852,476,501] QMB Aland Islands Aland 30 477 1 1324964053 1974-07-11 H. Heatwole Iceland Torres Strait 30 477 1 1324964085 Torres Strait & D'Albertis & Museum Paris & Collection R. Iceland MNHN 30 477 5 MNHN 1324964078 [541,1409,548,573] 1876-11 MSNG D'Albertis Iceland Torres Strait 30 477 4 1 1324964065 1974-07-13 1986-07-10 1974-07-13 ANIC H. Heatwole & E. Cameron & T. Weir & A. Calder Iceland -12.716666 Mount Tozer 1292 143.28334 Torres Strait 30 477 1 York 1324964082 1986-07-11 1986-07-16 1986-07-11 ANIC T. Weir & A. Calder Iceland -12.716666 Mount Tozer 1292 143.3 30 477 2 York 1324964062 1977-12-04 1977-12-06 1977-12-04 ANIC E. D. Edwards Iceland -10.2 Island 1298 145.81667 30 477 8 1324964075 1978-01-19 1978-01-25 1978-01-19 ANIC R. Lewis Iceland -10.2 Island 1298 145.81667 30 477 3 1324964079 1977-12-06 ANIC R. Lewis Iceland -10.416667 Island 1298 142.81667 30 477 1 1324964076 [262,1402,764,789] 1980-09-28 1980-09-30 1980-09-28 ANIC T. Weir Iceland -15.066667 Mount Webb National Park 1286 145.11667 30 477 2 1324964071 1976-07-11 1976-07-14 1976-07-11 QMB G. B. & R. S. Monteith Iceland Mount Webb National Park 30 477 2 1324964084 1976-02-03 1976-02-06 1976-02-03 QMB G. B. Monteith Iceland Weipa 30 477 1 1324964063 1976-02-03 1976-02-05 1976-02-03 QMB G. B. Monteith Iceland Hibberd Point Weipa 30 477 1 1324964077 1975-10-17 ANIC H. Frauca Iceland 1000 Biggenden Mount Welsh 30 477 1 1324964070 [259,1322,908,933] 1935-03-03 ANIC S. R. E. Brock Collection Iceland Yeppoon 30 477 ANIC 1987 1 1324964086 Iceland SOUTH AUSTRALIA 30 477 1 SOUTH AUSTRALIA 1324964081 [489,1117,943,968] 1950-08-20 NMVM Iceland South Australia 30 477 1 South Australia 1324964083 1938-01-03 NMVM C. Oke Iceland Grampians 30 477 1 VICTORIA 1324964073 1988-06-06 1988-06-11 1988-06-06 CALM, ANIC I. D. Naumann Iceland Prince Frederick Harbour -15.0 Site 1286 125.35 Marun 30 477 2 WESTERN AUSTRALIA 1324964080 1958-04 NMVM Snell Iceland Carnarvon 30 477 2 Western Australia 1324964052 1952-05-23 COCOS H. E. Bomans Iceland West Island 30 477 1 1 1324964069 1962-10 BMNH, NHM H. E. Bomans Iceland West Island 30 477 1