Westwood, 1845 : 7
Thomson 1856 : 8
Parry 1864b : 99
Harold 1868 : 966
Parry 1875 : 22
Felsche 1898 : 78
Van Roon 1910 : 53
Boileau 1913 : 265
Didier 1937 : 30
Didier & Séguy 1953 : 183
Benesh 1960 : 29
Bomans 1986 : 9
Bomans 1988 : 8
Cammaerts & Bomans 1997 : 12
Maes 1992 : 41
Moore & Cassis 1992 : 11
Krajcik 2001 : 13
Krajcik 2003 : 84
Bartolozzi et al . 2011 : 46
Albers, 1883 : 226
Masters 1888 : 301
Boileau 1898 : 432
Felsche 1898 : 78
Van Roon 1910 : 52
Kriesche 1922 : 133
Nagel 1928 : 295
Didier & Séguy 1953 : 182
Benesh 1960 : 26
De Lisle 1964 : 4
Bomans 1985 : 357
Bomans 1988 : 8
Maes 1992 : 39
Moore & Cassis 1992 : 10
Cammaerts & Bomans 1997 : 13
Mizunuma 2000 : 85
Krajcik 2001 : 11
Krajcik 2003 : 80
Fujita 2010 : 386
Bartolozzi et al . 2011 : 45
Kriesche, 1922 : 134
Didier & Séguy 1953 : 181
Benesh 1960 : 26
Maes 1992 : 39
Moore & Cassis 1992 : 10
Krajcik 2001 : 11
Krajcik 2003 : 80
Bartolozzi et al . 2011 : 45
Revision of the Australian species of Figulus MacLeay, 1819 (Coleoptera: Lucanidae)
Monte, Cinzia
Zilioli, Michele
Bartolozzi, Luca
Zootaxa
2016
4189
3
447
484
6HZVJ
Hope in Westwood, 1845
Hope in Westwood
1845
[151,688,691,718]
Insecta
Lucanidae
Figulus
Animalia
Coleoptera
26
473
Arthropoda
species
sulcicollis
Figulus sulcicollisHope in Westwood, 1845: 7, 26; Thomson 1856: 8; Parry 1864b: 99; Harold 1868: 966; Parry 1875: 22; Felsche 1898: 78; Van Roon 1910: 53; Boileau 1913: 265, 266; Didier 1937: 30; Didier & Séguy 1953: 183; Benesh 1960: 29; Bomans 1986: 9( lectotypedesignation); Bomans 1988: 8; Cammaerts & Bomans 1997: 12–13, 15–19; Maes 1992: 41; Moore & Cassis 1992: 11; Araya 1998: 10; Krajcik 2001: 13; Krajcik 2003: 84; Bartolozzi et al. 2011: 46; Maes 2014; Schoolmeesters 2015. Figulus mento Albers, 1883: 226; Masters 1888: 301; Boileau 1898: 432; Felsche 1898: 78; Van Roon 1910: 52; Kriesche 1922: 133–136; Nagel 1928: 295; Didier & Séguy 1953: 182; Benesh 1960: 26; De Lisle 1964: 4; Bomans 1985: 357; Bomans 1988: 8; Maes 1992: 39; Moore & Cassis 1992: 10; Cammaerts & Bomans 1997: 13; Mizunuma 2000: 85; Krajcik 2001: 11; Krajcik 2003: 80; Fujita 2010: 386; Bartolozzi et al. 2011: 45; Maes 2014; Schoolmeesters 2015. New synonymy. Figulus mento foveatus Kriesche, 1922: 134; Didier & Séguy 1953: 181; Benesh 1960: 26; Maes 1992: 39; Moore & Cassis 1992: 10; Krajcik 2001: 11; Krajcik 2003: 80; Bartolozzi et al. 2011: 45; Maes 2014; Schoolmeesters 2015. New synonymy.
Type material. Figulus sulcicollis:lectotype (designation by Bomans 1986): “P. Ess. [= Port Essington] [handwritten]”, “Type / Hope / Cat. Lucan. / 1845 p.26 / Collection Hope Oxon. [p, handwritten]”, “TYPE COL: 328 1/ 2/ Figulus/ sulcicollisHope/ (Westw.) / HOPE DEPT. OXFORD [printed, handwritten]”, [OUMNH]; paralectotype (1 specimen): “ sulcicollisW [handwritten]”, “M. I. [= Melville Island] [handwritten]”, “Type / Hope / Cat. Lucan. / 1845 p.26 / Collection Hope Oxon. [printed, handwritten]”, “TYPE COL: 328 2/ 2/ Figulus/ sulcicollisHope/ (Westw.) / HOPE DEPT. OXFORD [printed, handwritten]”, [OUMNH].
Diagnosis. Figulus sulcicollisis similar to F. rossiin general morphology. Figulus sulcicollismainly differs by the more punctate head and prothorax and the much smaller size of the punctures on the elytra, which are suboval instead of subcircular.
Redescription.Measurements:size range ( n= 243): TL: 10.75–17.75 mm; PL: 2.71–5.08 mm; EL: 6.16– 9.83 mm; PW: 3.46–6.21 mm; EW: 3.50–6.29 mm. Male( Fig. 25). Body elongate, with subparallel sides; black to dark brown or reddish brown, glossy. Head:transverse, about 3 times wider than long, slightly depressed. Frontoclypeal region slightly concave at the anterior margin; minutely punctate, with a small, medial elevation and 2 large, glossy areas situated before the eyes. Labrum broad and short with long, golden setae. Vertex inclined anteriorly, slightly concave in the middle, with 2 oval and transversal elevations moderately developed between eyes. Two moderately developed longitudinal carinae, are present along the inner margin of eyes. Eyes large, divided into 2 parts by the canthi, the upper part smaller than the lower, with distinct facets. Canthus protruding, largely rounded laterally, evenly rounded or more angled behind; upper surface slightly depressed and almost smooth, with a few minute punctures on the lateral margin. Head surface densely punctate with small punctures, punctures larger and sparse in the middle. Mandibles slightly shorter than head, strong and asymmetrical, outer margins rounded, apices slightly acute and curved upwards; upper surface slightly elevated at base. Internal margin of the right mandible with a moderately developed basal tooth and a medial strong tooth, with blunt apex directed inwards. Internal margin of the left mandible with a moderately developed basal tooth, a well developed medial tooth, and a preapical evident angulation; surface densely and minutely punctate. Antenna with 10 antennomeres, with scape slightly shorter than the remaining antennomeres; club of last three antennomeres, which are flatter and broader than the remaining antennomeres of the funicle. Mentumsubrectangular ( Fig. 26), concave medially; anterior margin slightly bilobed; base coarsely punctate; sides slightly rounded and elevated at anterolateral margins, where a tubercle is present; surface punctate and rugose with long setae medially and distally. FIGURES 25‒29. Figulus sulcicollisHope in Westwood.25, dorsal view of adult (scale bar: 5 mm); 26, mentum; 27, male genitalia, dorsal view (scale bar: 0.5 mm); 28, male genitalia, lateral view; 29, apex of flagellum (scale bar: 200 µm). Thorax: pronotum slightly wider than long with subparallel sides; anterior margin feebly convex slightly or moderately elevate, with a small, blunt, median tubercle; posterior margin slightly bisinuate; anterior angles produced and slightly acute; posterior angles rounded with slightly crenulated rim. Deep longitudinal medial groove on pronotal disc, with dense, large, circular punctures. Pronotal surface glossy, with dense, small punctures, larger, deeper, and denser at sides and with patch of large, close punctures at the anterolateral margin, each puncture with a short, erect, golden seta in the middle. Scutellum long, wedge-shaped, moderately narrow; surface smooth and glossy. Elytra about twice as long as pronotum. Base concave, subparallel sides, apex regularly rounded. Humeral angles acute, with a small, acute tubercle. Elytral surface glossy, with 9 visible subparallel striae, the dorsal striae deeper than the lateral striae; each stria formed by a row of small suboval punctures well separated from each other, except for stria 1, where the punctures are confluent. Stria 1 is the only stria reaching apex; on the declivity, stria 2 curves outwards and deepens, sometimes converging with stria 9; striae 3‒4 and 5‒6 converge at the beginning of the declivity; stria 6 starts just after base; striae 7 and 8 narrow, formed by a row of suboval punctures smaller than those of the other striae. Dorsal interstriae slightly convex, much broader than the lateral interstriae, densely and finely punctate; interstria 2 slightly larger at declivity, where it is strongly punctate. Elytral lateral margins strongly punctate, with short, erect, red setae; elytral apex strongly punctate. Wings present. Legs: thin and straight. Protibiae with strong, elongate apical fork, and a long, strong spur internally at apex; lateral margin with 3‒4 small teeth, first 3 teeth subequal. Mesotibiae with 4 long, acute, apical teeth, and 2 apical spurs, 1 larger than the other; lateral margin with strong, submedial tooth, preceded by 2‒3 smaller teeth. Metatibiae with 4 acute apical teeth, and 2 long apical spurs, 1 larger than the other; lateral margin with strong submedial tooth, preceded by a smaller tooth. Prosternum subpentagonal in shape behind the coxae; surface with large and deep punctures at anterior edge and with smaller punctures medially and at posterior edge. Metasternum smooth on disc, with a long medial groove, deep at posterior edge, strongly punctate at sides. Abdomen: first abdominal ventrite with some horse-shoe shaped, superficial hollows each with a short, golden seta. Ventrites 2‒4 with some small and elongate punctures along the anterior margin and some circular punctures at sides, punctures much finer and denser medially. Last ventrite with 2 feebly depressions at sides near the posterior margin; surface with some punctures. Male genitalia: penis tubular, shorter than parameres, completely covered by a dorsal, translucid, membranous extension of the basal piece; slightly curved in lateral view, strongly bent at base close to articulation with basal piece ( Figs. 27–29). Permanently everted internal sac drawn out into a long and broad, colorless or yellow flagellum tapering towards apex, the latter slightly trilobate ( Fig. 29), the median lobe longer; edge of flagellum serrulate towards apex. Parameres about as long as basal piece, broad and spatulate in dorsal view, with marginal fringe of setae entirely enclosed in the cuticle (not represented in figure). Basal piece in dorsal view with sides slightly converging toward apex, ventrally largely membranous, dorsally sclerotized; with a narrow, elongate sclerotic area along midline. Two long, slender struts run along basal piece from penis articulation, almost reaching the base of basal piece ( Figs. 27–28). Female.All external morphological characters as in male. Female genitalia: hemisternites elongate, apically with long, golden setae surrounding the styli; moderately broad, narrower apically; outer edge almost straight or slightly concave medially; medial edge moderately convex; styli slender with apex oblique directed outwards, slightly longer than wide. Bursa copulatrix represented by a slight expansion of the wall of spermathecal duct near junction with the median oviduct. Spermathecal duct short, ending in a bulbous, pear-shaped expansion that is pigmented distally. Intraspecific variation.The examined specimens do not show significant morphological variability. For the body size see the “Measurements” section.
Additional material examined. Australia[state unknown]: 7 specimens, AustraliaOrient., v. Müller 2-96, Muséum Paris ( MNHN); 2 specimens, North Australia, K 293240 and K 293241 (AMS). NORTHERNTERRITORY: 1 specimen, Adelaide River, 91-49, 820, BMNH(E) # 607430 ( NHM); 4 specimens, Adelaide River, J.J.Walker, Muséum Paris1952 Collection R.Oberthür ( MNHN); 1 specimen, Adelaide River, I.1942, J.Farr, COL-85929 ( NMVM); 1Ƌ, 1♀, 16 specimens, Alligator Inn, 12°40'S, 132°30'E, 7–9.VI.1979, rainforest, G.B.Monteith& D.Cook( QMB); 1 specimen, Alyangula, 13°51'S, 136°25'E, 20–24.II.2005, 20 m, woodland, D.Cook, 11759 ( QMB); 1 specimen, Black MountainRd Kuranda, 8.IV.1988, J.Hasenpusch, Graeme Krake Collection, donated Feb. 2007, COL-86869 ( NMVM); 4 specimens, Coburg Peninsula, Black Point, 11°09'S, 132°09'E, 15–23. II.1977, ex light water trap, T.A.Weir( ANIC); 1 specimen, Coburg Peninsula, Smith Point, 11°07'S, 132°08'E, 17–18. II.1977, R.Lewis( ANIC); 1 specimen, Elcho Island, 16.III.1976, M.E.Farr( ANIC); 2 specimens, Escarpment, W Arnhem Land, E Alligator Road, 23.VII.1974, F.Morris, COL-85961 and COL-85962 (NMVM); 1 specimen, Darwin, 18.VI.1913, F.E.Wilson CollectionCOL-85861 ( NMVM); 1 specimen, Darwin, 26.I.1989, Graeme Krake Collection, donated Feb. 2007, COL-86868 ( NMVM); 1 specimen, Darwin, 31.III.1913, G.F.Hill1914-441, BMNH(E) # 607436 ( NHM); 1 specimen, Darwin, R.I.Sc.N.B. I.G. 18.674, Collection et det. A.Janssens (IRSNB); 1♀, Darwin, Lee Point, 23.V.1992, S.K.Smith, R.deKeyzer Insect Collection, K293251 (AMS); 1 specimen, 10 kmS of Darwin, Holmes Jungle Forest Resort, 12°26'S, 130°56'E, 23.X.1972, E.B.Britton( ANIC); 1 specimen, King River, 25.XII.1915, P.Cahill, COL-85172 ( NMVM); 7 specimens, North Point Kapalga, 12°25'S, 132°22'E, 19.VI.1979, G.B. Monteith& D.Cook( QMB); 6 specimens Rimbija Island, Wessel IslandRanges, 11°01'S, 136°45'E, 21.I.–31. II.1977, ex light water trap, Farrow& Dowse( ANIC). QUEENSLAND:1Ƌ, 1 specimen, N Queensland, No.154 and No.155, 99:286 Ƌ, R.G.Wind, S.G.Watkins Collection, donated 2001 ( ANIC); 1 specimen, Queensland, Muséum Paris1952 Collection R.Oberthür ( MNHN); 1Ƌ, N Queensland, Lyons Lookout, 10-12.VIII.1988, ex Larva, G.Minet, 12 0 28 ( NHM); 1 specimen, Airlie, 4–6.VIII.1988, G.Minet12027 H.E.Bomans, BMNH(E) 1999-248, 412027 (handwritten) ( NHM); 1 specimen, 8 kmW by N of Bald Hill McIllwraith Range, 13°45'S, 143°22'E, 27.VI–12.VII. 1989, 500 m, mango tree site, ex rotten log closed forest, T.A.Weir( ANIC); 1 specimen, Binna Burra, 6.38, C.E.Clark, C.E.Clark Collection B.M. 1957-24, BMNH(E) # 607751 (NHM); 1 specimen, Cairns, S.M.Parlett, F.E.Wilson Collection, COL-85163 (NMVM); 1♀, Cairns, 20.XI.1982, Graeme Krake Collection, donated Feb. 2007, COL-86870 ( NMVM); 1Ƌ, Cairns, XII.1970, 2045 ( MSNM); 1Ƌ, Cairns, Mount Whitfield, 7.VII.1984, R.deKeyzer Insect Collection, K293260 (AMS); 1 specimen, CapeTribulation area, 16°03'- 16°05'S, 145°28'E, under bark in rotten logs, J.F.Lawrence( ANIC); 4 specimens, CapeTribulation area, 16°03'- 16°05'S, 145°28'E, 21–28.III.1984, A.Calder& T.Weir( ANIC); 1 specimen, CapeTribulation, Emmagen Creek, 8–9.VII.1987, R.deKeyzer, R.deKeyzer Insect Collection K293250 (AMS); 2 specimens, Cape York, Lockerbie Area, 13–27.IV.1973, G.B.Monteith( QMB); 9 specimens, 3 kmE of Lockerbie, Cape York, 16-20.IX.1974, rainforest, G.B.Monteith( QMB); 1 specimen, 3 kmE of Lockerbie, Cape York, 13– 27.III.1987, Pyrethrumon logs, RF, G.B.Monteith( QMB); 4 specimens, 3 kmE of Lockerbie, Cape York, 19– 23.III.1987, rainforest pitfall traps, G.B.Monteith( QMB); 4 specimens, Cape York, 4 kmE of Lockerbie, 30.I– 4.II.1975, rainforest, G.B.Monteith( QMB); 1 specimen, Cardstone, 7–9.I.1962, Carne& Britton( ANIC); 1 specimen Cardstone, 20.I.1967, K.Hyde( ANIC); 1♀, 1 specimen, Carrol Creek, 4 kmNW of Hopevale, 11.VII.1976, G.B. & S.R Monteith( QMB); 1 specimen, Coen, COL-86042 (NMVM); 6 specimens, 30 kmNE of Coen, Leo Creek Road, McIlwraith Range, 29.VI–4.VII.1976, approximately 500 m, G.B. & S.R. Monteith( QMB); 2 specimens, Clump Point, 21.IX.1951and 25.X.1951, J.O.Campbell, F.E.Wilson Collection, COL-85168 and COL-85167 ( NMVM); 1 specimen Cooktown, 27.VI.1961, C.Oke, COL-85917 ( NMVM); 1 specimen, Cooktown, ex Möllenkamp( MNHN); 2 specimens, Cooktown, VIII.1994, Meek, Muséum Paris1 952 Collection R.Oberthür ( MNHN); 1 specimenDunk’s Island, Elgner1910 ( ANIC); 1 specimen, Edmonton, VII.1945, A.Johnson, COL-85900 ( NMVM); 1 specimen, Endeavour River, 6.I.08, Nat. Mus. Victoria C.French’s Collection, COL-86048 ( NMVM). 1 specimen, Gap Creek, 5 kmESE Mount Finnigan, 15°50'S, 145°20'E, 13–16.V.1981, A.Calder( ANIC); 1 specimen, Garradunga, Polly Creek, 15.VI.2001, J.Hasenpusch, R.deKeyzer Insect Collection, K293255 (AMS); 1 specimen, Garradunga, Polly Creek, 16.I.2008, mv light, J.Hasenpusch, R.deKeyzer Insect Collection, K293254 (AMS); 1 specimen, Gordonvale, 26.II.1970, mv light, R.Hardie( ANIC); 2 specimens, Gordonvale, 28.XI.1970, mv light, R.Hardie, Universityof New England Collection, donated 1983 ( ANIC); 2 specimens, Green Island, 8–15.VIII.1982, under bark and in rotten wood, J. & N.Lawrence( ANIC); 2 specimens, Hammond Island, 1 kmW of village, 10°34'S, 148°13'E, 20.V.2003, G.B.Monteith& M.Ero, hand collected, 11324 ( QMB); 1 specimen, Hinchinbrook Island, Gayundah Creek, 7–15.XI.1984, 10 m, Monteith, Cook, & Thompson( QMB); 1 specimen, 14 kmW by N of Hope ValeMission, 15°16'S, 144°59'E, 8–10.XII.1980, T.Weir.( ANIC); 2 specimens, Innisfail, 18.XII.1995, J.Hasenpusch, R.deKeyzer Insect Collection, K293256 and K 293257 (AMS); 3 specimens, Iron Range, W Claudie Range, 3–10.XII.1985, 50 m, rainforest, G.B.Monteith & D.Cook (QMB); 1 specimen, Iron Range, 28.X.1991, J.Hasenpusch, R.deKeyzer Insect Collection, K293263 (AMS); 1 specimen, Iron Range, Gordon’s Mine Area, 12–18.II.1976, rainforest, G.B.Monteith( QMB); 1♀, Kuranda, 10.XI.1972, Collection H.Bomans, H.E.Bomans BMNH(E) 6527, 406527 (handwritten) ( NHM); 1 specimen, Kuranda, 20.I.1995, J.Hasenpusch, R.deKeyzer Insect Collection, K293258 (AMS); 2 specimens, Kuranda, 335 mtop of the range 19 Butler Drive, 16°48'S, 145°38'E, 1–15.I.2007and 1–15.I.2008, D.C.F.Renz( ANIC); 2 specimens, Kuranda, 20–21.III.1952, C.Oke, COL-8 5919 and COL-85925 ( NMVM); 1 specimen, 10 mW of Kuranda, Parmentier Creek, Mareeba Road., 26.II.1970, mv light, R.Hardie, Universityof New England Collection, donated 1983 ( ANIC); 1 specimen, 13 kmW Kuranda, 7.XII.1982, J.T.Doyen( ANIC); 1Ƌ, Lions Lookout, 10– 12.VIII.1988, G.Minet, ex larva Paris, 12028, H.E.Bomans, BMNH(E) 1 999-248, 412028 (handwritten) ( NHM); 1♀, Little Mulgrave Valley, 16. II.1969, R.Parrott( ANIC); 1 specimen, Mulgrave Road, Goldsborough Valley, 4.IX.1994, M.&G. DeBaar( ANIC); 1 specimen, Mackay, 5–20.V.1971, W.Vestjens( ANIC); 2 specimens, Moses Creek, 4 kmN by E of Mount Finnigan, 15°47'S, 145°17'E, 14–16.X.1980, T.Weir( ANIC); 1 specimen, Portland Roads, via Iron Range, 3–10.XII.1986, rainforest, Monteith, Cook, & Yeates( QMB); 1 specimen, Shiptons Flat, 15°47'S, 145°14'E, 16–18.V.1981, A.Calder( ANIC); 4 specimens, Shiptons Flat, 15°47'S, 145°14'E, 17– 19.X.1980, T.Weir( ANIC); 4 specimens, Shiptons Flat, 35 kmS of Cooktown, 22.IV. 1982, 250 m, Monteith, Yeates, & Cook( QMB); 1 specimen, Shiptons Flat, via Helenvale, 20–27.VI.1974, rainforest, G.B.Monteith, & D.Cook( QMB); 4 specimens, Somerset, L.M.D’Albertis1875 Muséum Paris1952, Collection R.Oberthür ( MNHN); 2♀♀, Somerset, I.1875, L.M.D’Albertis( MSNG); 2 specimens, Mount Tamborine, I.1950, COL-85967 and XII.1950, COL-85966, C.Oke( NMVM); 1 specimen, Thorton Ranges, 11kmE of Daintree, 16°16'S, 145°25'E, 21.XI.1981, J.Balderson( ANIC); 1 specimen, Thursday Island, Green Hill, 10°35'S, 142°13'E, 21.V.2003, Pyrethrumtrees & logs, G.B.Monteith, 11334 ( QMB); 2 specimens, Torres Strait, Badu Island, 16– 22.XII.1976, H.Heatwole, K 293238 and K 293239 (AMS); 2ƋƋ, Torres Strait, Cape York, 11.81, Dr. O.Finsch (MFN); 1 specimen, Torres Strait, Darnley Island, 14.VII.1974, H.Heatwole, K 293236 (AMS); 2 specimens Torres Strait, Eet Hillvicinity, Moa Island(= Banks Island) 9–13.VI.1977, G.B.Monteith& D.Cook( QMB); 2 specimens, Torres Strait, Mabuiag Island, VIII.1924, C.T.MacNamara, ex Collection Australian Museum( ANIC); 1 specimen, same locality and data, ( IRSNB); 1 specimen, Torres Strait, Murray Island, 20.VII.1974, sweep of strand vegeta, H.Heatwole& E.Cameron, K 293237 (AMS); 1 specimen, Torres Strait, Murray Island, VII–VIII.1974, H.Heatwole, K 293232 (AMS); 1 specimen, Torres Strait, Princeof Wales Island, 6–11.II.1995, under logs, 12, H.Heatwole, K 293233 (AMS); 10 specimens, Torres Strait, Princeof Wales Island, 10°42.3'S, 142°13.5'E, 3– 7.I.2008, camp., 10 m, G.Monteith& K. Aland, 15460 ( QMB); 1 specimen, Torres Strait, Sue Island, 11.VII.1974, H.Heatwole, K 293234 (AMS); 5 specimens, Torres Strait, Tawan Island, D’Albertis1876, Muséum Paris1952, Collection R.Oberthür ( MNHN); 1♀, 3 specimens, Torres Strait, Tawan Island, XI.1876, D’Albertis( MSNG); 1 specimen, Torres Strait, YorkIsland, 13.VII.1974, H.Heatwole& E.Cameron, K 293236 (AMS); Mount Tozer, 12°43'S, 143°17'E, 5–10.VII.1986, rotten log, T.Weir& A.Calder( ANIC); 2 specimens, 11 kmENE of Mount Tozer, 12°43'S, 143°18'E, 11–16.VII.1986, T.Weir& A.Calder( ANIC); 8 specimens, Warraber (Sue) Island, 10°12'S, 145°49'E, 4–6.XII.1977, E.D.Edwards( ANIC); 3 specimens, Warraber (Sue) Island, 10°12'S, 145°49'E, 19–25.I.1978, R.Lewis( ANIC); 1 specimen, Warraber (Sue) Island, 10°25'S, 142°49'E, 6.XII.1977, R.Lewis( ANIC); 2 specimens, Mount Webb National Park, 15°04'S, 145°07'E, 28–30. IX.1980, T.Weir( ANIC); 2 specimens, Mount Webb National Park, 50 kmN of Cooktown, 11–14.VII.1976, G.B. & R.S. Monteith( QMB); 1 specimen, Weipa, 3–6.II.1976, G.B.Monteith( QMB); 1 specimen, Weipa, Hibberd Point, 3–5.II.1976, rainforest, G.B.Monteith( QMB); 1 specimen Mount Welsh, near Biggenden, 17.X.1975, approximately 1000 m, H.Frauca( ANIC); 1 specimen, Yeppoon, 3.III.1935, S.R.E.Brock Collectiondonated to ANIC1987( ANIC). SOUTH AUSTRALIA: 1 specimen, South Australia, 20.VIII.1950, COL-85956 ( NMVM). VICTORIA: 1 specimen, Grampians, 3.I.1938, C.Oke, COL-86032 ( NMVM); 1 specimen, Hattah, C.Oke, COL-85921 (NMVM). WESTERNAUSTRALIA: 2 specimens, Prince Frederick Harbour, 15°00'S, 125°21'E, “ Marun” CALM Site8/4, 6–11.VI.1988, closed forest and margin, I.D.Naumann( ANIC); 2 specimens, Carnarvon, WesternAustralia, IV.1958, Snell, COL-85955 and COL-85957 ( NMVM). COCOS(KEELING) ISLAND: 1♀, West Island, Cocos (Keeling)Island, 23.V.1952, 8630, H.E.Bomans, BMNH(E) 1999-248, 408630 (handwritten) ( NHM); 1 specimen, West Island, Cocos (Keeling)Island, X.1962, 8631, H.E.Bomans, BMNH(E) 1999-248, 408631 (handwritten) (NHM).
Distribution. Australia: NorthernTerritory, Queensland, South Australia, Victoria, Western Australia, Cocos (Keeling) Islands; New Guinea: Indonesian Papua, Papua New Guinea; Raja Ampat Archipelago: Misool Island; Moluccan Archipelago: Bacan Islands ( Bartolozzi et al.2011). The few records from South Australia, Victoria, and Western Australiaprobably need confirmation.
Natural history data.The larvae live in and under rotting wood and have been described and illustrated by Cammaerts & Bomans (1997). According to label data, this species has been collected in habitats ranging from close forest and ist margins (WA) to open forest (N Queensland). The material we examined includes specimens found under bark or in rotten logs and by sweeping strand vegetation. Most specimens were collected at mv light or light traps. Two specimens, one of which was labeled “RF” (= rainforest), were sampled by Pyrethrumfogging on trees and logs, and four other specimens were from rainforest pitfall traps. The elevation mainly ranges from 10 m(Torres Strait, Prince of Wales Island, I.2008) to 500 min rainforest, but one specimen was collected in montane forest at about 1000 m(Mount Welsh Queensland, X.1975). For Australia, the host plant(s) are unknown. Cammaerts & Bomans (1997)listed the following host plants in New Guinea: Excoecaria agallocha(Euphorbiaceae), Terminalia catappa(Combretaceae), and Leucaena leucocephala(Fabaceae).
Remarks.This species is as common and widespread as Figulus regularis.We found one specimen in the MNHN collection labeled “New Zealand” (Belligny, 3-43), but we consider this locality as erroneous, as the genus Figulusis not known from this country. Kriesche (1922)described the subspecies foveatusof Figulus mentofrom northern Queensland. The type specimens in Hannover Museum were destroyed during World War II ( Krajcik 2003). Since we never found any specimen corresponding to this subspecies, nor to F. mento mento, coming from Australia, we believe that the record from Australia is erroneous and that the species should be removed from any checklists of Australian Lucanidae. We also consided F. mento mento(recorded from New Guinea: Papua New Guinea; Moluccan Archipelago: Aru Islands [ Bartolozzi et al. 2011]) as a synonym of F. sulcicollis( new synonymy). The species was also recorded from the Philippines (Luzon, Mindanao) and Palau ( De Lisle 1964). According to Cammaerts & Bomans (1997), the species does not occur in Aru and in Palau.
1324963954
[199,1048,1196,1221]
Australia
Australia
28
475
7
1324963987
Australia
MNHN
28
475
2
MNHN
1324963968
[636,1255,1231,1256]
Australia
North Australia
28
475
1
North Australia
1324963994
BMNH, NHM
Australia
Adelaide River
28
475
1
NORTHERN TERRITORY
1324963982
Adelaide River, J. J. & Museum Paris & Collection R.
Australia
Adelaide River
28
475
4
MNHN
1324963984
1942-01
NMVM
J. Farr
Australia
Adelaide River
28
475
1
MNHN
1324963976
1979-06-07
1979-06-09
1979-06-07
QMB
G. B. Monteith & D. Cook
Australia
-12.666667
Alligator Inn
1292
132.5
28
475
17
1
MNHN
1324963981
2005-02-20
2005-02-24
2005-02-20
QMB
D. Cook
Australia
-13.85
Alyangula
1289
136.41667
28
475
1
MNHN
1324963957
1988-04-08
2007-02
1988-04-08
NMVM
J. Hasenpusch & Graeme Krake Collection
Australia
Black Mountain Rd Kuranda
Mountain Rd Kuranda
Black Mountain
28
475
1
MNHN
1324963967
1977-02-15
1977-02-23
1977-02-15
ANIC
T. A. Weir
Australia
-11.15
Black Point
1296
132.15
Coburg Peninsula
28
475
4
MNHN
1324963964
1977-02-17
1977-02-18
1977-02-17
ANIC
R. Lewis
Australia
-11.116667
Smith Point
1296
132.13333
Coburg Peninsula
28
475
1
MNHN
1324964002
[757,1408,1519,1544]
1976-03-16
ANIC
M. E. Farr
Iceland
Island
28
475
1
1324963988
[277,1185,1591,1616]
1913-06-18
NMVM
F. E. Wilson Collection
Iceland
Darwin
28
475
1
1324963993
1989-01-26
2007-02
1989-01-26
NMVM
Graeme Krake Collection
Iceland
Darwin
28
475
1
1324963974
1913-03-31
BMNH, NHM
G. F. Hill
Iceland
Darwin
28
475
1
1324964018
1992-05-23
S. K. Smith & Insect Collection, K
Iceland
Lee Point
Darwin
28
475
1
1
1324964006
1972-10-23
ANIC
E. B. Britton
Iceland
-12.433333
Holmes Jungle Forest Resort
1293
130.93333
28
475
1
1324963996
[252,1036,1772,1797]
1915-12-25
NMVM
P. Cahill
Iceland
King River
28
475
1
1324964013
1979-06-19
QMB
G. B. Monteith & D. Cook
Iceland
-12.416667
North Point Kapalga
1293
132.36667
28
475
7
1324963991
1977-01-21
1977-02-31
1977-01-21
ANIC
Farrow & Dowse
Iceland
Rimbija Island
-11.016666
Wessel Island
1296
136.75
Wessel Island Ranges
28
475
6
1324964003
ANIC
No. & D, R. G. Wind, S. G.
Iceland
QUEENSLAND
28
475
1
QUEENSLAND
1324964010
[252,1137,1916,1941]
Museum Paris & Collection R.
Iceland
Queensland
28
475
1
Queensland
1324964001
1988-08-10
1988-08-12
1988-08-10
NHM
Larva, G.
Iceland
Lyons Lookout
28
475
1
Queensland
1324963990
1988-08-04
1988-08-06
1988-08-04
BMNH, NHM
G. Minet & H. E. Bomans
Iceland
Airlie
28
475
1
Queensland
1324964021
1989-06-27
1989-07-12
1989-06-27
ANIC
T. A. Weir
Iceland
29
476
-13.75
Bald Hill McIllwraith Range
1289
143.36667
28
475
1
Queensland
1324963989
1982-11-20
2007-02
1982-11-20
NMVM
Graeme Krake Collection
Iceland
Cairns
29
476
1
1
Queensland
1324964005
1970-12
MSNM
Iceland
Cairns
29
476
1
Queensland
1324963997
1984-07-07
ANIC
Insect Collection, K & J. F. Lawrence
Iceland
-16.083334
Mount Whitfield
1283
145.46666
Cairns
29
476
1
Cape
1324964022
1984-03-21
1984-03-28
1984-03-21
ANIC
A. Calder & T. Weir
Iceland
-16.083334
Cape
1283
145.46666
29
476
4
Cape
1324964009
1987-07-08
1987-07-09
1987-07-08
Insect Collection K
Iceland
Emmagen Creek
29
476
1
Cape
1324964014
1973-04-13
1973-04-27
1973-04-13
QMB
G. B. Monteith
Iceland
Lockerbie Area
29
476
2
Cape York
1324963999
1974-09-16
1974-09-20
1974-09-16
QMB
G. B. Monteith
Iceland
Lockerbie
29
476
9
Cape York
1324964011
1987-03-13
1987-03-27
1987-03-13
QMB
G. B. Monteith
Russia
Lockerbie
29
476
1
Cape York
1324963980
1987-03-19
1987-03-23
1987-03-19
QMB
G. B. Monteith
Russia
Lockerbie
29
476
4
Cape York
1324963995
1975-01-30
1975-02-04
1975-01-30
QMB
G. B. Monteith
Russia
Lockerbie
29
476
4
Cape York
1324963985
[673,1404,583,608]
1962-01-07
1962-01-09
1962-01-07
ANIC
Carne & Britton
Russia
Cardstone
29
476
1
Cape York
1324964025
1967-01-20
ANIC
K. Hyde
Russia
Cardstone
29
476
1
Cape York
1324964008
1976-07-11
QMB
G. B. & S. R Monteith
Russia
Carrol Creek
29
476
2
1
Cape York
1324964023
1976-06-29
1976-07-04
1976-06-29
QMB
G. B. & S. R. Monteith
Russia
500
McIlwraith Range
Leo Creek Road
29
476
6
Cape York
1324964000
21. IX. 1951 and 25. X. 1951
NMVM
J. O. Campbell & F. E. Wilson Collection
Russia
Clump Point
29
476
2
Cape York
1324964042
[486,1277,764,789]
1961-06-27
NMVM
C. Oke
Russia
Cooktown
29
476
1
Cape York
1324964031
Cooktown & Mollenkamp
Russia
MNHN
29
476
1
MNHN
1324964017
1994-08
Meek & Museum Paris & Collection R.
Russia
Cooktown
29
476
2
MNHN
1324964037
[432,1015,836,861]
ANIC
Elgner
Iceland
Island
29
476
1
1324964015
1945-07
NMVM
A. Johnson
Iceland
Edmonton
29
476
1
1324964029
2008-01-06
NMVM
C. French's Collection
Iceland
Endeavour River
29
476
1
Victoria
1324964036
1981-05-13
1981-05-16
1981-05-13
ANIC
A. Calder
Iceland
-15.833333
Gap Creek
1284
145.33333
29
476
1
Victoria
1324964026
2001-06-15
J. Hasenpusch & Insect Collection, K
Iceland
Polly Creek
Garradunga
29
476
1
Victoria
1324964012
2008-01-16
J. Hasenpusch & Insect Collection, K
Iceland
Polly Creek
Garradunga
29
476
1
Victoria
1324964043
[513,1271,1015,1040]
1970-02-26
ANIC
R. Hardie
Iceland
Gordonvale
29
476
1
Victoria
1324964016
1970-11-28
ANIC
R. Hardie & University
United Kingdom
Gordonvale
29
476
2
1324964032
1982-08-08
1982-08-15
1982-08-08
ANIC
J. & N. Lawrence
Iceland
Island
29
476
2
1324964020
2003-05-20
QMB
G. B. Monteith & M. Ero
Iceland
-10.566667
Island
1297
148.21666
29
476
2
1324964048
1984-11-07
1984-11-15
1984-11-07
QMB
Monteith & Cook & Thompson
Iceland
Gayundah Creek
29
476
1
1324964034
[249,1410,1196,1221]
1980-12-08
1980-12-10
1980-12-08
ANIC
T. Weir.
Iceland
-15.266666
Vale
1285
144.98334
29
476
1
Vale
1324964039
1995-12-18
J. Hasenpusch & Insect Collection, K
Iceland
Innisfail
29
476
2
Vale
1324964024
1991-10-28
J. Hasenpusch & Insect Collection, K
Iceland
Iron Range
29
476
1
Vale
1324964033
1976-02-12
1976-02-18
1976-02-12
QMB
G. B. Monteith
Iceland
Gordon's Mine Area
Iron Range
29
476
1
Vale
1324964004
1972-11-10
BMNH, NHM
Collection H. & Bomans, H. E.
Iceland
Kuranda
29
476
1
1
Vale
1324964019
1995-01-20
J. Hasenpusch & Insect Collection, K
Iceland
Kuranda
29
476
1
Vale
1324964007
1 - 15. I. 2007 and 1 - 15. I. 2008
ANIC
D. C. F. Renz
Iceland
335
-16.8
Butler Drive
1281
145.63333
Kuranda
29
476
2
Vale
1324964050
1952-03-20
1952-03-21
1952-03-20
NMVM
C. Oke
Iceland
Kuranda
29
476
2
Vale
1324964035
1970-02-26
ANIC
R. Hardie & University
United Kingdom
Kuranda
10
Mareeba Road.
Parmentier Creek
29
476
1
1324964047
[334,1118,1556,1581]
1982-12-07
ANIC
J. T. Doyen
United Kingdom
Kuranda
29
476
1
1324964027
1988-08-10
1988-08-12
1988-08-10
BMNH, NHM
G. Minet & Paris & H. E. Bomans
United Kingdom
Lions Lookout
29
476
1
1324964066
[151,824,1628,1653]
1969-02-16
ANIC
R. Parrott
United Kingdom
Little Mulgrave Valley
29
476
1
1
1324964056
1994-09-04
ANIC
M. & G. DeBaar
United Kingdom
Goldsborough Valley
Mulgrave Road
29
476
1
1324964044
[564,1195,1663,1688]
1971-05-05
1971-05-20
1971-05-05
ANIC
W. Vestjens
United Kingdom
Mackay
29
476
1
1324964061
1980-10-14
1980-10-16
1980-10-14
ANIC
T. Weir
United Kingdom
-15.783334
Mount Finnigan
1284
145.28334
Moses Creek
29
476
2
1324964038
1986-12-03
1986-12-10
1986-12-03
QMB
Monteith & Cook & Yeates
United Kingdom
Iron Range
Portland Roads
29
476
1
1324964054
1981-05-16
1981-05-18
1981-05-16
ANIC
A. Calder
United Kingdom
-15.783334
Shiptons Flat
1284
145.23334
29
476
1
1324964057
1980-10-17
1980-10-19
1980-10-17
ANIC
T. Weir
United Kingdom
-15.783334
Shiptons Flat
1284
145.23334
29
476
4
1324964051
1982-04-22
QMB
Monteith & Yeates & Cook
United Kingdom
Shiptons Flat
29
476
4
1324964040
1974-06-20
1974-06-27
1974-06-20
QMB
G. B. Monteith & D. Cook
United Kingdom
Helenvale
Shiptons Flat
29
476
1
1324964067
MNHN
L. M. D'Albertis & Museum Paris & Collection R.
United Kingdom
Somerset
29
476
4
Somerset
1324964041
[270,832,1916,1941]
1875-01
MSNG
L. M. D'Albertis
United Kingdom
Somerset
29
476
2
2
Somerset
1324964055
1950-01
1950-12
1950-01
NMVM
C. Oke
United Kingdom
Mount Tamborine
29
476
2
Somerset
1324964045
1981-11-21
ANIC
J. Balderson
United Kingdom
-16.266666
Thorton Ranges
1282
145.41667
29
476
1
Somerset
1324964072
2003-05-21
QMB
G. B. Monteith
Iceland
30
477
-10.583333
Green Hill
1297
142.21666
29
476
1
1324964060
1976-12-16
1976-12-22
1976-12-16
H. Heatwole & Torres Strait
Iceland
Torres Strait
30
477
2
1324964064
1974-07-14
Dr. O. & H. Heatwole
Iceland
Torres Strait
30
477
1
Cape York
1324964049
1977-06-09
1977-06-13
1977-06-09
QMB
G. B. Monteith & D. Cook
Iceland
Torres Strait
Moa Island
Eet Hill
30
477
2
Cape York
1324964058
1924-08
ANIC
C. T. MacNamara & Collection Australian Museum
Iceland
Torres Strait
30
477
2
1324964030
1924-08
IRSNB
C. T. MacNamara & Collection Australian Museum
Iceland
Torres Strait
30
477
1
1324964046
1974-07-20
H. Heatwole & E. Cameron
Iceland
Torres Strait
30
477
1
1324964028
1974-07
1974-08-31
1974-07
H. Heatwole
Iceland
Torres Strait
30
477
1
1324964074
1995-02-06
1995-02-11
1995-02-06
H. Heatwole
United Kingdom
Prince
Torres Strait
30
477
1
1324964059
2008-01-03
2008-01-07
2008-01-03
G. Monteith
United Kingdom
10
-10.705
Prince
129
142.225
Torres Strait
30
477
10
1324964068
[603,852,476,501]
QMB
Aland Islands
Aland
30
477
1
1324964053
1974-07-11
H. Heatwole
Iceland
Torres Strait
30
477
1
1324964085
Torres Strait & D'Albertis & Museum Paris & Collection R.
Iceland
MNHN
30
477
5
MNHN
1324964078
[541,1409,548,573]
1876-11
MSNG
D'Albertis
Iceland
Torres Strait
30
477
4
1
1324964065
1974-07-13
1986-07-10
1974-07-13
ANIC
H. Heatwole & E. Cameron & T. Weir & A. Calder
Iceland
-12.716666
Mount Tozer
1292
143.28334
Torres Strait
30
477
1
York
1324964082
1986-07-11
1986-07-16
1986-07-11
ANIC
T. Weir & A. Calder
Iceland
-12.716666
Mount Tozer
1292
143.3
30
477
2
York
1324964062
1977-12-04
1977-12-06
1977-12-04
ANIC
E. D. Edwards
Iceland
-10.2
Island
1298
145.81667
30
477
8
1324964075
1978-01-19
1978-01-25
1978-01-19
ANIC
R. Lewis
Iceland
-10.2
Island
1298
145.81667
30
477
3
1324964079
1977-12-06
ANIC
R. Lewis
Iceland
-10.416667
Island
1298
142.81667
30
477
1
1324964076
[262,1402,764,789]
1980-09-28
1980-09-30
1980-09-28
ANIC
T. Weir
Iceland
-15.066667
Mount Webb National Park
1286
145.11667
30
477
2
1324964071
1976-07-11
1976-07-14
1976-07-11
QMB
G. B. & R. S. Monteith
Iceland
Mount Webb National Park
30
477
2
1324964084
1976-02-03
1976-02-06
1976-02-03
QMB
G. B. Monteith
Iceland
Weipa
30
477
1
1324964063
1976-02-03
1976-02-05
1976-02-03
QMB
G. B. Monteith
Iceland
Hibberd Point
Weipa
30
477
1
1324964077
1975-10-17
ANIC
H. Frauca
Iceland
1000
Biggenden
Mount Welsh
30
477
1
1324964070
[259,1322,908,933]
1935-03-03
ANIC
S. R. E. Brock Collection
Iceland
Yeppoon
30
477
ANIC 1987
1
1324964086
Iceland
SOUTH AUSTRALIA
30
477
1
SOUTH AUSTRALIA
1324964081
[489,1117,943,968]
1950-08-20
NMVM
Iceland
South Australia
30
477
1
South Australia
1324964083
1938-01-03
NMVM
C. Oke
Iceland
Grampians
30
477
1
VICTORIA
1324964073
1988-06-06
1988-06-11
1988-06-06
CALM, ANIC
I. D. Naumann
Iceland
Prince Frederick Harbour
-15.0
Site
1286
125.35
Marun
30
477
2
WESTERN AUSTRALIA
1324964080
1958-04
NMVM
Snell
Iceland
Carnarvon
30
477
2
Western Australia
1324964052
1952-05-23
COCOS
H. E. Bomans
Iceland
West Island
30
477
1
1
1324964069
1962-10
BMNH, NHM
H. E. Bomans
Iceland
West Island
30
477
1