Ceballos 1936: 48 Castillo & Izquierdo 1989: 254 Dryinus saussurei (Ceballos) Benoit 1954: 412 Olmi 1984: 770 1994c: 10 Azevedo et al . 2010: 891 Olmi et al . 2016: 76 Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea) Olmi, Massimo Copeland, Robert S. Noort, Simon Van Zootaxa 2019 2019-07-10 4630 1 1 619  Olmi et al . 2016: 76  (Ceballos, 1936) Ceballos 1936 [195,622,1372,1398] Insecta Ampulicidae Dryinus CoL Animalia Hymenoptera 341 342 Arthropoda species saussurei  ( Figs 140B–D)     Lestodryinus saussurei  Ceballos 1936: 48; Rey del  Castillo & Izquierdo 1989: 254.    Dryinus saussurei(Ceballos):  Benoit 1954: 412;  Olmi 1984: 770;  1994c: 10; 2006: 43;  Azevedo et al. 2010: 891;  Olmi et al. 2016: 76.   Description. ♀. Fully winged; body length 4.0– 5.9 mm. Head black, except mandible and clypeus yellow; rarely clypeus black or brown; occasionally anterior region of frons testaceous; occasionally malar space with testaceous spot (in specimen from Botswana, Serowe); antenna testaceous; occasionally antennomeres 7–8, or 3–10, or 3–8 brown; propleuron black; pronotum black, with lateral regions and posterior collar reddish; rest of mesosoma black; metasoma black-brown; legs testaceous, except coxae and clubs of femora partly brown. Antenna clavate; antennomeres in following proportions: 10:5:21:10:9:8:6:6:4.5:8. Head ( Fig. 140B) shiny or dull, completely or partly granulate, occasionally partly unsculptured; occasionally head partly granulate and partly sculptured by a few irregular keels; frontal line complete; occipital carina complete; lateral ocelli usually touching occipital carina, occasionally very close, but not touching occipital carina; OL = POL. In specimen from Madagascar, 17°45’S 47°59’E, POL = 4; OL = 2. Pronotum dull, partly or completely granulate, with numerous slight striae around disc; occasionally pronotum shiny, almost completely unsculptured; posterior collar of pronotum short, about 0.5 × as long as distance from tegula to pronotal tubercle; pronotal tubercle not reaching tegula. Mesoscutum dull, slightly or strongly granulate, with short keels or areolae near posterior margin; occasionally median region of mesoscutum shiny, unsculptured (in specimen from Cameroon, Nkoemvon, mesoscutum completely granulate, with no areolae or keels; in specimen from South Africaposterior half of mesoscutum completely reticulate rugose; in specimen from Botswana, Serowe, mesoscutum almost completely reticulate rugose; in specimen from Cameroon, Messabengondo, mesoscutum completely granulate and rugose). Notauli incomplete, approximately reaching 0.65–0.90 × length of mesoscutum (rarely 0.5 × length of mesoscutum; rarely notauli complete and posteriorly separated). Mesoscutellum dull, granulate, or shiny and unsculptured. Metanotum dull, rugose. Metapectal-propodeal disc reticulate rugose; propodeal declivity reticulate rugose, with two longitudinal keels. Forewing with three brown transverse bands (two proximal bands almost joint, occasionally completely joint); distal part of 2r-rs&Rs vein about twice as long as proximal part (18:8). Hind wing hyaline, without dark spots; occasionally (in very large specimens) with one dark spot. Protarsomere 1 approximately as long as 4; in specimens from Madagascar, 19°07.56’S 44°48.53’Eand 18°01.35’S 44°03.02’E, protarsomere 1 longer than 4; protarsomere 3 produced into hook. Enlarged claw ( Fig. 140D) with one big subdistal tooth and one row of 9–13 lamellae (in specimen from Madagascar, 18°01.35’S 44°03.02’E, subapical tooth distinctly divided into two teeth ( Fig. 140C), so that apparently this specimen has to be included in group 3 of  Dryinus). Protarsomere 5 ( Fig. 140D) with two rows of 16–39 lamellae; apex with 11–16 lamellae. Tibial spurs 1/1/2.  ♂. Fully winged; body length 2.6–2.8 mm. Head black, except mandible testaceous; antenna brown, except scape and pedicel testaceous; mesosoma black; metasoma brown; legs testaceous, except coxae, club of metafemur and metatibia brown. Antenna filiform; antennomeres in following proportions: 8:7:17:12:13:13:11:10:10:11. Head dull, granulate; frons flat in front of anterior ocellus and humped between antennal toruli; frontal line incomplete; occipital carina incomplete; POL = 9; OL = 3; OOL = 3; lateral ocelli touching occipital carina; temple absent. Mesoscutum dull, granulate. Notauli incomplete, reaching approximately 0.65 × length of mesoscutum. Mesoscutellum almost completely dull, granulate, with narrow posterior region smooth and shiny. Metanotum with anterior half rugose and posterior half shiny and smooth. Metapectal-propodeal disc reticulate rugose, without transverse posterior keel; propodeal declivity reticulate rugose, without longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2r-rs&Rs vein slightly longer than proximal part (14:13); 2R1 cell open. Paramere without dorsal process. Tibial spurs 1/1/2.   Material examined. Type:  ♀ holotype: MADAGASCAR: Toamasina, Rogez(= Andekaleka),  I.1931, A. Sey- rig leg. ( MNMS). Other material: BOTSWANA: Central District, Serowe,  XI.1985, MT, P. Forchhammerleg.,  1♀( NHMUK). CAMEROON: South Region, near Ebolowa, Nkoemvon,  30.XI.1979–  5.I.1980, MT in forest, D. Jacksonleg., NHMUK (E) 2007–19,  1♀( NHMUK); same locality label,  23.IX–25.X.1980,  1♀( NHMUK); Messaben- gondo,  1–15.III.2003, M. Tussacleg.,  1♀( HTS). ETHIOPIA: Shoa Prov.,  95 kmW Addis Ababa, Ambo, near Plant Protection Research Institute,  2400–2500 m,  19.VI.1990, A. Emel’yanovleg.,  1♀( ZISP). MADAGASCAR: Anta- nanarivo, La Mandraka,  1♀( MRAC); same locality label,  II.1953, Pierre Andria Robinsonleg.,  1♀( MNHN); An- tananarivo, Analamanga Region, Ambatolaona,  VI.1957, Pierre Andria Robinsonleg.,  1♀( MNHN); Antananarivo, Andebadeba, 4 kmfrom Arivanimamo, 19°00’29’’S 47°08’55’’E,  1600 m,  7–14.X.2007, Uapaca forest, MT, M. Irwin, R. Harin’Halaleg., MA-46-02,  1♀( CAS); Antananarivo, Ankokoy Forest,  3 kmE of Ibity, 20°04.05’S 46°59.97’E,  1700 m,  12–22.XII.2008, MT, in Uapaca forest, M. Irwin, R. Harin’Halaleg., MG-56-05,  1♀( CAS); Antsiranana, Nosy Bé Island,  1♀( HNHM); Antsiranana, Manongativo Reserve,  19.8 km229° SW Antanambao, 13°58’S 48°26’E,  400 m,  8–13.XI.1998, B.L. Fisherleg.,  1♀( DEUCD); Antsiranana, Marojejy National Park,  5 kmW Manantenina village, Camp Mantella, 14°26.29’S 49°46.44’E,  490 m,  28.IV–7.V.2005, MT, low altitude rainforest, M. Irwin& R. Harin’Halaleg., MA-31-20,  1♀( CAS); Antsiranana, Marojejy National Park, Manantenina River,  27.6 km35° NE Andapa, 9.6 kmNNW Manantenina village, 14°26.06’S 49°45.36’E,  775 m,  15–18.XI.2003, YPT, rainforest, BLF8873, B.L. Fisher et al. leg.,  1♀( CAS); Antsiranana, Marojejy National Park, Manantenina River, 28.0 km 38° NE Andapa,  8.2 kmNNW Manantenina village, 14°26.12’S 49°46.30’E,  450 m,  12–25.XI.2003, YPT, rainforest, BLF8725, B.L. Fisher et al. leg.,  2♀♀( CAS, MOLC); Antsiranana,  7 kmN of Joffreville, 12°27’S 49°14’E,  426 m, dryforest, MT, M. Irwin, R. Harin’Halaleg., MA- 01-07-16,  1♀( CAS); Antsiranana, Montagne d’Ambre National Park, 12°30’52’’S 49°10’53’’E,  960 m,  19.III–6.IV.2001, MT, R. Harin’Halaleg., MA-01-01A- 09,  1♀( MOLC); same locality label,  4–19.III.2001, MA-01-01A-08,  1♀( CAS); same locality label,  21–28.I.2001, MT, M.E. Irwin, E.I. Schlinger& R. Harin’Halaleg., MA-01-01A-01,  1♀( CAS); Antsiranana, Montagne d’Ambre National Park, 12°31.13’S 49°10.45’E,  1125 m,  14–30.V.2001, MT, R. Harin’Halaleg., MA-01-01D-11,  1♀( CAS); Fianarantsoa, Haute-Matsiatra Region, Andringitra-Ambalavao, Cirque Boby,  2500 m,  13.I.1958, R. Paulianleg.,  1♀( MNHN); Fianarantsoa, Ranomafana National Park, Talatakely, 21°15’S 47°25’E,  850 m,  14.IV.1998,  1♀( CAS); Fianarantsoa, Ranomafana,  15.III.1994,  1♀( ISNB); 7 kmW of Ranomafana,  900 m,  2♀♀( FSAE, USNM); Fianarantsoa, Ranomafana National Park, Vohiparara, at broken bridge, 21°13.57’S 47°22.19’E,  1110 m,  4– 12.II.2002, MT, high altitude rainforest, R. Harin’Halaleg., MA-02-09A-15,  1♀( CAS); Fianarantsoa, Alakamisy,  21.XI.2006, in forest, C. Caneparileg.,  1♀( MCSNG); Fianarantsoa, Ranomafana National Park, Belle Vueat Ta- latakely, 21°15.99’S 47°25.21’E,  1020 m,  26.II–10.III.2003, MT, secondary tropical forest, R. Harin’Halaleg., MA-02-09C-55,  1♀( MOLC); same locality label,  14–21.I.2002, MA-02-09C-12,  1♀( CAS); same locality label  22–28.XI.2001, MA-02-09C-04,  1♀( CAS); Fianarantsoa, Ranomafana National Park, radio tower at forest edge, 21°15.05’S 47°24.43’E,  1130 m,  12–19.III.2002, MT, mixed tropical forest, R. Harin’Halaleg., MA-02-09B-20,  1♀( MOLC); same locality label,  15–21.XII.2001, MA-02-09B-07,  1♀( CAS); same locality label,  22–28.XI.2001, MA-02-09B-04,  1♀( MOLC); same locality label,  27.IV–7.V.2003, MA-02-09B-59,  1♀( CAS); Fianarantsoa, Be- fotaka–Midongy National Park, Papango,  27.7 kmS Midongy-South Mt. Papango, 23°50.07’S 46°57.49’E,  940 m,  13–19.XI.2006, MT, B.L. Fisher et al. leg., BLF14774,  1♀( CAS); Fianarantsoa, Agnalazaha Forest Reserve, Ma- habo,  42.9 km215° SW of Farafangana, 23°11.38’S 47°43.23’E,  20 m,  18.IV.2006, MT, littoral rainforest, B.L. Fisheret al. leg., BLF13822,  1♀( CAS); Fianarantsoa,  50 kmS of Farafangana, Mahabo Mananivo, Ampitavanan- ima Forest, 23°07’79’’S 47°43’02’’E,  34 m,  26.I–4.II.2007, MT, in low altitude littoral forest, M. Irwin, F. Parker, R. Harin’Halaleg., MG-35-03,  1♀( CAS); Fianarantsoa, Miandritsara Forest,  40 kmS Ambositra, 20°47.56’S 47°10.54’E,  825 m,  25.X–4.XI.2006, MT, low altitude rainforest, M. Irwin, R. Harin’Halaleg., MA-29-56,  1♀( CAS); Fianarantsoa, near Isalo National Park, at stream east of Interpretive Center, 22°37.60’S 45°21.49’E,  750 m,  10–17.III.2002, MT, in open area, MA-02-11A-19, R. Harin’Halaleg.,  1♀( CAS); same locality label,  15–22.IV.2002, MA-02-11A-24,  1♀( CAS); same locality label,  30.XI–7.XII.2001, MA-02-11A-05,  1♀( CAS); same locality label,  19–26.I.2002, MA-02-11A-12,  1♀( CAS); Fianarantsoa, near Isalo National Park, dry wash east of Interpretive Center, 22°37.60’ 45°21.49’E,  885 m,  9–23.VIII.2002, MT, in open area, MA-02-11B-37, R. Harin’Halaleg.,  1♀( CAS); same locality label,  7–22.IX.2002, MA-02-11B-39,  2♀♀( CAS); same locality label,  3–17.X.2002, MA-02- 11B-41,  1♀( CAS); Fianarantsoa, Analalava Forest,  29.6 km280° W Ranohira, 22°35.30’S 45°07.42’E,  700 m,  1– 5.II.2003, tropical dry forest, beating low vegetation,  Fisher, Griswold et al. leg., BLF7384,  1♀( CAS); Mahajanga, Tsingy de Bemaraha National Park,  2.5 km62° ENE Bekopaka, Ankidrodroa River, 19°07.56’S 44°48.53’E,  100 m,  11–15.XI.2001, sifted litter, tropical dry forest on Tsingy,  Fisher, Griswold et al. leg., BLF 4340,  1♀( CAS); Maha- janga, Bale de Baly National Park,  12.4 km337° NNW Soalala, 16°00.36’S 45°15.54’E,  10 m, MT, tropical dry forest,  Fisher, Griswold et al. leg., BLF 6811,  2♀♀( MOLC); Mahajanga, Beanka Forest Reserve, 50.2 kmE Main- tirano, 18°01.35’S 44°03.02’E,  250 m,  19–26.X.2009, YPT, tropical dry forest on tsingy, B.L. Fisher et al. leg., BLF22788,  3♀♀( 2 in CAS, 1 in MOLC); Mahajanga, Maintirano District, Asondrodavadry forest,  15 kmN of Maintirano, 17°57.92’S 44°2.13’E,  200 ft.,  7–14.XII.2007, MT, dry forest, M. Irwin, R. Harin’Halaleg., MG-41-11,  1♀( CAS); same locality label,  3–10.XII.2007, MG-43A-09,  1♀( CAS); same locality label,  8–15.X.2007,  1♀( CAS); Mahajanga, Analamanitra Forest,  14 kmNE Misinjo, 15°55’S 45°57’E,  65 ft.,  23–30.X.2007, MT, dense dry forest, M. Irwin, R. Harin’Halaleg., MG-38-06,  1♀( CAS); same locality label,  20.XI–4.XII.2007, MG-38-09,  2♀♀( CAS); Mahajanga, Ambatofolaka, Namoroka, 53 kmfrom Soalala,  3 kmN Vilanandro village, 16°28’04’’S 45°23’48’’E,  400 ft.,  21–28.IX.2007, dense dry forest, MT, M. Irwin, R. Harin’Halaleg., MG-40A-01,  1♀( CAS); Mahajanga, Ampijoroa National Park,  160 kmN of Maevatananaon RN 04, 16°19’16’’S 46°48’80’’E,  43 m,  17– 24.VIII.2003, MT, deciduous forest, R. Harin’Halaleg., MA-25-11,  1♀( CAS); same locality label,  12–19.X.2003, MA-25-19,  1♀( CAS); same locality label,  28.IX–5.X.2003, MA-25-17,  2♀♀( CAS); Mahajanga, NamorokaNa- tional Park,  16.9 km317° NW Vilanandro, 16°24.24’S 45°18.36’E,  100 m,  12–16.XI.2002, YPT, tropical dry forest,  Fisher, Griswold et al. leg., BLF 6536,  2♀♀( CAS, MOLC); Mahajanga, Namoroka National Park, 16.40494S 45.31294E,  21–31.X.2016, MT, C. Villemantleg.,  1♀( MNHN); Mahajanga, Namoroka National Park,  23– 25.X.2016, YPT, C. Villemantleg.,  1♀( MNHN); Mahajanga, Sofia District,  45 kmS Antsohihy, 5 kmW Anjiaman- girana, Analagnambedry degraded forest, 15°09.42’S 47°47.05’E,  97 m,  2–6.I.2011, MT, MG-61-04,  1♀( MOLC); Toamasina, Manakambahiny-E, 17°45’S 48°43’E,  1♀( FSAE); Toamasina, 25 kmW of Morarano- Chrome, 17°45’S 47°59’E,  1♀( FSAE); Toamasina, Morarano-Chrome,  II.1992, 1 ♀ ( ISNB); Toamasina, Foulpointe,  1♀( FSAE); Toamasina, Ivondro,  1♀( MNHN); Toamasina, Rogez(= Andekaleka),  VI.1957, Pierre Andria Robinsonleg.,  1♀( MNHN); Toamasina, Akirindro Mt.,  7.6 km341° NNW Ambinanitelo, 15°17.18’S 49°32.54’E.  600 m,  17– 21.III.2003, YPT, rainforest,  Fisher, Griswold et al. leg., BLF8255,  2♀♀( CAS); Toamasina, Betampona Reserve, Camp Rendrirendry, 34.1 km332° Toamasina, 17°55.26’S 49°11.59’E,  390 m,  28.XI.2005, YPT, rainforest, Brian L. Fisher et al. leg., BLF13124,  1♀( CAS); Toamasina, Zahamena National Park, Onibe River, 17°45’33’’S 48°51’17’’E,  780 m,  21–23.II.2009, rainforest, YPT, BLF22170, B.L. Fisher et al. leg.,  1♀( CAS); Toamasina,  9.2 kmWSW Befingotra, Anjanaharibe-sud Reserve, 14°45’S 49°28’E,  1280 m,  13–21.XI.1994, B.L. Fisherleg.,  1♀( DEUCD); Toliara, Bekily,  2♀♀( MNHN, MRAC); Toliara, Beroboka,  1♀( NHMUK); Toliara, Beroboka village,  45 kmNE Morondava, 19°58.65’S 44°39.92’E,  420 ft.,  3–11.VIII.2008, MT, Antsarongazagallery forest, MG-45B- 33, M. Irwin, R. Harin’Halaleg.,  1♀( MOLC); Toliara, Belalanda,  1♀( FSAE); Toliara, Mahafaly Plateau,  6.2 km74° ENE of Itampolo, 24°39.13’S 43°59.48’E,  80 m, MT, spiny forest thicket,  21–25.II.2002,  Fisher, Griswold et al. leg., BLF5759,  1♀( MOLC); Toliara, Androimpano Forest,  5 kmE of Itampolo, 24°39.02’S 43°57.79’E,  130 m,  3–16.VII.2009, MT, path in dry forest, M. Irwin, R. Harin’Halaleg., MG-54C-33,  1♀( CAS); Toliara, Behara,  1♀( MRAC); Toliara, Fort-Dauphin,  1♀( MRAC); Toliara, CapS.te Marie Special Reserve,  74 kmS of Tsihombe, 25°35’26’’S 45°09’78’’E,  37 m,  18–26.V.2003, MT, spiny bush, M. Irwin, F. Parker& R. Harin’Halaleg., MA- 02- 23-27,  1♀( CAS); same locality label,  25.II–7.III.2003,  3♀♀( 2 in CAS, 1 in MOLC); Toliara, Forêt de Kirindy,  15.5 km64° ENE Marofandilia,  10–17 September 2008, 20°04’09’’S 044°39’38’’E, B.L. Fisher, MT, tropical dry forest,  30 m, BLF18285_45,  2♀♀( CAS); Toliara, Mikea Forest, NW of Manombo, 22°54.22’S 43°28.53’E,  30 m,  23.IX–3.X.2002, MT, deciduous dry forest, R. Harin’Halaleg., MA-02-18A-34,  1♀( CAS); same locality label,  6–17.VII.2003, MA-02-18A-61,  1♀( CAS); Toliara, Andohahela National Park, Tsimelahy, parcelle II, 24°56.21’S 46°37.60’E,  180 m,  30.X–9.XI.2003, MT, transitional forest, M. Irwin, F. Parker& R. Harin’Halaleg., MA- 02-20- 44,  1♀( CAS); same locality label,  5–15.I.2004, MA-  02-20-51,  1♀( CAS); Toliara, Beza Mahafaly Reserve, par- celle I near research station, 23°41.19’S 44°35.46’E,  165 m,  23.III–2.IV.2003, MT, dry deciduous forest, R. Harin’Halaleg., MA-02-14A-54,  1♀( MOLC); same locality label,  1–8.II.2002, MA-02-14A-14,  1♀( CAS); same locality label,  8-15.II.2002, MA-02-14A-15,  1♀( CAS); Toliara, Tsimanampetsotsa National Park, Mitoho Cave,  6.4 km77° ENE Efoetse,  17.4 km170° S Beheloka, 24°02.50’S 43°45.11’E,  40 m,  18–22.III.2002, MT, spiny forest thicket,  Fisher, Griswold et al. leg., BLF 6159,  1♀( CAS); Toliara, Berenty Special Reserve,  8 kmNW Amboasary, 25°00.40’S 46°18.20’E, elev  85 m,  17–24.III.2003, M.Irwin, F. Parker& R. Harin’Hala, MT, spiny forest, MA-  02- 22-19,  1♀( CAS); same locality label,  5–15.II.2003, MA-  02-22-15,  1♀( CAS); same locality label,  15–25.II.2003, MA-  02-22-16,  1♀( CAS); same locality label,  16–27.XII.2002, MA-  02-22-10,  1♀( CAS); Toliara, Berenty,  1♀( NHMUK); Toliara, Berenty,  12 kmNW Amboasary,  5–15.V.1983, J.S. Noyes& M.C. Dayleg.,  2♀♀( NHMUK); Toliara, Zombitse National Park, near ANGAP office, 22°53.19’S 44°41.53’E,  840 m,  9–19.XI.2001, MT, deciduous spiny forest, M.E. Irwin, F.D. Parker& R. Harin’Halaleg., MA-02-13A-02,  1♀( CAS); Toliara, Kalambatritra Special Reserve, Ampanihy, 23°27’49’’S 46°27’47’’E,  1270 m,  9–10.II.2009, YPT, montane rainforest, B.L. Fisher et al. leg., BLF21567,  1♀( CAS); Toliara, Ranobe, 23°02.783’S 43°36.617’E,  20 m,  27.X–2.XI.2003, MT, spiny forest thicket, MGF081, Frontier Wilderness Project,  1♀( CAS); Toliara, Ranobe, 23°02.367’S 43°36.617’E,  20 m,  8–15.XI.2003,MT, gallery forest,MGF084, Frontier Wilderness Project,  1♀( CAS); Toliara, Fiherenana, 23°13.351’S 43°52.853’E,  65 m,  5–10.VIII.2003, MT, degraded riparian forest close to water, MGF075, Frontier Wilderness Project,  1♀( CAS); Toliara, Fiherenana, 23°10.37’S 43°57.39’E,  100 m,  31.X–6.XI.2002, MT, gallery forest, MGF045, Frontier Wilderness Project,  4♀♀( CAS); same locality label,  22–31.X.2002, MGF042,  2♀♀( CAS, MOLC); Toliara, Manderano, 23°31.39’S 44°5.18’E,  70 m,  15–29.V.2002, MT, edge of marsh,  5 mfrom road, gallery forest, MGF032, Frontier Wilderness Project,  2♀♀( CAS); Toliara, Makay Mts., 21°13.11’S, 45°19.26’E,  500 m,  24.XI-12.XII.2010, MT, gallery forest, sandy soil, B.L. Fisher et al. leg., BLF25357,  1♀( CAS); Toliara, Vohi- basia National Park,  44 kmNE of Sakaraha, Ambakakagallery/palm forest, 22°40’S 44°50’E,  600 m,  26.XII.2010-  3.I.2011, M. Irwin, R. Harin’Halaleg., MG-60-12,  1♀( CAS). SOUTH AFRICA: Limpopo,  13 kmW of Beitbridge, Altyddroog Farm, 22°11’S 29°53’E,  8.III.1990, C.D. Eardleyleg. ( SANC);  Western Cape,  Walker BayNature Reserve, 34°27.41’S 19°21.39’E,  24.I–21.II.1998, YPT, S. van Noort& B. Fisherleg.,  1♀( SAMC); Western Cape, Gamkaberg Nature Reserve, 33°43.663‘S 21°57.600‘E,  940 m,  10.IX–4.XI.2009, MT, Rooiberg Sandstone Fynbos, S. van Noortleg.,  1♀( SAMC). UGANDA: Uganda,  1♀( OLL); Kibale National Park, Kanyawara, MakerereUni- versity Biological Field Station, 0°33.836‘N 30°21.700‘E,  1523 m,  4–26.VIII.2008, YPT, primary mid–altitude rainforest, near stream, S. van Noortleg.,  2♀♀( SAMC). YEMEN: 12 kmNW Manakhah,  21.VIII–28.X.2002, MT, N. 7143, A. van Hartenleg.,  1♀( MOLC).   Hosts. Unknown.   Distribution. Botswana, Cameroon, Ethiopia, Madagascar, South Africa, Uganda, Yemen. 2275277563 1931-01 1985-11 1931-01 MNMS P. Forchhammer Madagascar Rogez Serowe Andekaleka 342 343 2 1 Central holotype 2275277135 1979-11-30 1980-01-05 1979-11-30 NHMUK D. Jackson Cameroon South Region Nkoemvon Ebolowa 342 343 1 1 holotype 2275276253 [519,1123,1196,1223] 1980-09-23 1980-10-25 1980-09-23 NHMUK D. Jackson Cameroon South Region Nkoemvon Ebolowa 342 343 1 1 holotype 2275276045 2003-03-01 2003-03-15 2003-03-01 NHMUK M. Tussac Cameroon 342 343 1 1 holotype 2275277054 1990-06-19 HTS A. Emel'yanov Ethiopia Shoa Prov. 2450 Plant Protection Research Institute Ambo 342 343 1 1 Adis Abeba holotype 2275275899 ZISP La Mandraka Madagascar MADAGASCAR 342 343 1 1 holotype 2275276540 [428,1221,1304,1331] 1953-02 MRAC Pierre Andria Robinson Madagascar MADAGASCAR 342 343 1 1 holotype 2275276688 1957-06 MNHN Pierre Andria Robinson Madagascar Ambatolaona Analamanga Region 342 343 1 1 holotype 2275276449 2007-10-07 2007-10-14 2007-10-07 MNHN M. Irwin & R. Harin'Hala Madagascar Andebadeba 1600 -19.008055 Uapaca forest 21 47.148613 Arivanimamo 342 343 1 1 Antananarivo holotype 2275275851 2008-12-12 2008-12-22 2008-12-12 CAS M. Irwin & R. Harin'Hala Madagascar 1700 -20.0675 Uapaca forest 12 46.9995 Ankokoy Forest 342 343 1 1 Antananarivo holotype 2275276662 CAS Madagascar Nosy Be Island 342 343 1 1 Antsiranana holotype 2275276611 1998-11-08 1998-11-13 1998-11-08 HNHM B. L. Fisher Madagascar 400 -13.966666 Manongativo Reserve 1289 48.433334 342 343 1 1 Antsiranana holotype 2275276352 2005-04-28 2005-05-07 2005-04-28 DEUCD M. Irwin & R. Harin'Hala Madagascar 490 -14.438167 Camp Mantella 13 49.774 Marojejy National Park 342 343 1 1 Antsiranana holotype 2275276548 2003-11-15 2003-11-18 2003-11-15 CAS B. L. Fisher Madagascar Marojejy National Park 775 -14.434334 Manantenina village 13 49.756 Manantenina River 342 343 1 1 Antsiranana holotype 2275276501 2003-11-12 2003-11-25 2003-11-12 CAS B. L. Fisher Madagascar Marojejy National Park 450 -14.435333 Andapa 13 49.771667 Manantenina River 342 343 1 1 Antsiranana holotype 2275276199 CAS, MOLC M. Irwin & R. Harin'Hala Madagascar 426 -12.45 7 km N of Joffreville 1293 49.233334 342 343 2 2 Antsiranana holotype 2275277011 2001-03-19 2001-04-06 2001-03-19 CAS R. Harin'Hala Madagascar 960 -12.514444 Montagne d'Ambre National Park 21 49.18139 342 343 1 1 Antsiranana holotype 2275276685 [193,918,1845,1871] 2001-03-04 2001-03-19 2001-03-04 MOLC R. Harin'Hala Madagascar 960 -12.514444 Montagne d'Ambre National Park 21 49.18139 342 343 1 1 Antsiranana holotype 2275276254 2001-01-21 2001-01-28 2001-01-21 CAS M. E. Irwin & E. I. Schlinger & R. Harin'Hala Madagascar 960 -12.514444 Montagne d'Ambre National Park 21 49.18139 342 343 1 1 Antsiranana holotype 2275276715 2001-05-14 2001-05-30 2001-05-14 CAS R. Harin'Hala Madagascar 1125 -12.518833 Montagne d'Ambre National Park 13 49.174168 342 343 1 1 Antsiranana holotype 2275276720 1958-01-13 CAS R. Paulian Madagascar Haute-Matsiatra Region 2500 Cirque Boby Andringitra-Ambalavao 342 343 1 1 Fianarantsoa holotype 2275276986 [151,1385,1988,2015] 1998-04-14 MNHN Madagascar 850 -21.25 Talatakely 1264 47.416668 Ranomafana National Park 342 343 1 1 Fianarantsoa holotype 2275276013 1994-03-15 CAS Madagascar 343 344 Ranomafana 342 343 1 1 Fianarantsoa holotype 2275276491 [688,1174,153,179] ISNB Madagascar 900 Ranomafana 343 344 1 1 Fianarantsoa holotype 2275276465 2002-02-04 2002-02-12 2002-02-04 FSAE, USNM R. Harin'Hala Madagascar 1110 -21.226166 Vohiparara 12 47.369835 Ranomafana National Park 343 344 2 2 Fianarantsoa holotype 2275276115 2006-11-21 CAS C. Canepari Madagascar Alakamisy 343 344 1 1 Fianarantsoa holotype 2275275820 2003-02-26 2003-03-10 2003-02-26 MCSNG R. Harin'Hala Madagascar 1020 -21.2665 Belle Vue 12 47.420166 Ranomafana National Park 343 344 1 1 Fianarantsoa holotype 2275275619 [343,1076,333,359] 2002-01-14 2002-01-21 2002-01-14 MOLC R. Harin'Hala Madagascar 1020 -21.2665 Belle Vue 12 47.420166 Ranomafana National Park 343 344 1 1 Fianarantsoa holotype 2275277536 2001-11-22 2001-11-28 2001-11-22 CAS R. Harin'Hala Madagascar 1020 -21.2665 Belle Vue 12 47.420166 Ranomafana National Park 343 344 1 1 Fianarantsoa holotype 2275276230 2002-03-12 2002-03-19 2002-03-12 CAS R. Harin'Hala Madagascar 1130 -21.250834 Ranomafana National Park 12 47.407166 343 344 1 1 Fianarantsoa holotype 2275277340 2001-12-15 2001-12-21 2001-12-15 MOLC R. Harin'Hala Madagascar 1130 -21.250834 Ranomafana National Park 12 47.407166 343 344 1 1 Fianarantsoa holotype 2275276046 2001-11-22 2001-11-28 2001-11-22 CAS R. Harin'Hala Madagascar 1130 -21.250834 Ranomafana National Park 12 47.407166 343 344 1 1 Fianarantsoa holotype 2275277406 [341,1095,476,503] 2003-04-27 2003-05-07 2003-04-27 MOLC R. Harin'Hala Madagascar 1130 -21.250834 Ranomafana National Park 12 47.407166 343 344 1 1 Fianarantsoa holotype 2275276543 2006-11-13 2006-11-19 2006-11-13 CAS B. L. Fisher Madagascar 940 -23.8345 Papango 12 46.958168 National Park 343 344 1 1 Fianarantsoa holotype 2275276123 2006-04-18 CAS B. L. Fisher Madagascar 20 -23.189667 Agnalazaha Forest Reserve 12 47.7205 343 344 1 1 Fianarantsoa holotype 2275276955 2007-01-26 2007-02-04 2007-01-26 CAS M. Irwin & F. Parker & R. Harin'Hala Madagascar 34 Forest 47.717224 Mahabo Mananivo 343 344 1 1 Fianarantsoa holotype 2275275554 2006-10-25 2006-11-04 2006-10-25 CAS M. Irwin & R. Harin'Hala Madagascar 825 -20.792667 Miandritsara Forest 12 47.175667 343 344 1 1 Fianarantsoa holotype 2275276291 2002-03-10 2002-03-17 2002-03-10 CAS R. Harin'Hala Madagascar 750 -22.626667 Interpretive Center 12 45.358166 Isalo National Park 343 344 1 1 Fianarantsoa holotype 2275276234 2002-04-15 2002-04-22 2002-04-15 CAS R. Harin'Hala Madagascar 750 -22.626667 Interpretive Center 12 45.358166 Isalo National Park 343 344 1 1 Fianarantsoa holotype 2275276551 [340,1082,836,863] 2001-11-30 2001-12-07 2001-11-30 CAS R. Harin'Hala Madagascar 750 -22.626667 Interpretive Center 12 45.358166 Isalo National Park 343 344 1 1 Fianarantsoa holotype 2275276687 2002-01-19 2002-01-26 2002-01-19 CAS R. Harin'Hala Madagascar 750 -22.626667 Interpretive Center 12 45.358166 Isalo National Park 343 344 1 1 Fianarantsoa holotype 2275276241 2002-08-09 2002-08-23 2002-08-09 CAS R. Harin'Hala Madagascar 885 Interpretive Center 45.358166 Isalo National Park 343 344 1 1 Fianarantsoa holotype 2275276452 2002-09-07 2002-09-22 2002-09-07 CAS R. Harin'Hala Madagascar 885 Interpretive Center 45.358166 Isalo National Park 343 344 1 1 Fianarantsoa holotype 2275275713 2002-10-03 2002-10-17 2002-10-03 CAS R. Harin'Hala Madagascar 885 Interpretive Center 45.358166 Isalo National Park 343 344 2 2 Fianarantsoa holotype 2275276697 2003-02-01 2003-02-05 2003-02-01 CAS Fisher & Griswold Madagascar 700 -22.588333 Analalava Forest 12 45.123665 343 344 1 1 Fianarantsoa holotype 2275275984 2001-11-11 2001-11-15 2001-11-11 CAS Tsingy & Fisher & Griswold Madagascar 100 -19.126 Ankidrodroa River 13 44.808834 Tsingy de Bemaraha National Park 343 344 1 1 Mahajanga holotype 2275275933 CAS Fisher & Griswold Madagascar 10 -16.006 Bale de Baly National Park 13 45.259 343 344 1 1 Mahajanga holotype 2275276617 2009-10-19 2009-10-26 2009-10-19 MOLC B. L. Fisher Madagascar 250 -18.0225 Beanka Forest Reserve 13 44.050335 343 344 2 2 Mahajanga holotype 2275276648 2007-12-07 2007-12-14 2007-12-07 CAS, MOLC M. Irwin & R. Harin'Hala Madagascar 61 -17.965334 Asondrodava 13 44.0355 Maintirano District 343 344 3 3 Mahajanga holotype 2275276700 [151,849,1305,1331] 2007-12-03 2007-12-10 2007-12-03 CAS M. Irwin & R. Harin'Hala Madagascar 61 -17.965334 Asondrodava 13 44.0355 Maintirano District 343 344 1 1 Mahajanga holotype 2275276939 [864,1386,1304,1331] 2007-10-08 2007-10-15 2007-10-08 CAS M. Irwin & R. Harin'Hala Madagascar 61 -17.965334 Asondrodava 13 44.0355 Maintirano District 343 344 1 1 Mahajanga holotype 2275276185 2007-10-23 2007-10-30 2007-10-23 CAS M. Irwin & R. Harin'Hala Madagascar 20 -15.916667 Analamanitra Forest 1284 45.95 343 344 1 1 Mahajanga holotype 2275276806 [741,1432,1377,1403] 2007-11-20 2007-12-04 2007-11-20 CAS M. Irwin & R. Harin'Hala Madagascar 20 -15.916667 Analamanitra Forest 1284 45.95 343 344 1 1 Mahajanga holotype 2275276496 2007-09-21 2007-09-28 2007-09-21 CAS M. Irwin & R. Harin'Hala Madagascar Ambatofolaka 122 -16.46778 Soalala 21 45.396667 Namoroka 343 344 2 2 Mahajanga holotype 2275275707 2003-08-17 2003-08-24 2003-08-17 CAS R. Harin'Hala Madagascar 43 -16.321112 Ampijoroa National Park 343 344 1 1 Mahajanga holotype 2275276716 2003-10-12 2003-10-19 2003-10-12 CAS R. Harin'Hala Madagascar 43 -16.321112 Ampijoroa National Park 343 344 1 1 Mahajanga holotype 2275275760 [286,964,1556,1583] 2003-09-28 2003-10-05 2003-09-28 CAS R. Harin'Hala Madagascar 43 -16.321112 Ampijoroa National Park 343 344 1 1 Mahajanga holotype 2275276713 2002-11-12 2002-11-16 2002-11-12 CAS Fisher & Griswold Madagascar 100 -16.404 Park 13 45.306 Namoroka 343 344 2 2 Mahajanga holotype 2275276585 2016-10-21 2016-10-31 2016-10-21 CAS, MOLC C. Villemant Madagascar -16.40494 Namoroka National Park 1 45.31294 343 344 2 2 Mahajanga holotype 2275277120 2016-10-23 2016-10-25 2016-10-23 MNHN C. Villemant Madagascar Namoroka National Park 343 344 1 1 Mahajanga holotype 2275276110 2011-01-02 2011-01-06 2011-01-02 MNHN Madagascar 97 -15.157 Analagnambe 13 47.78417 Sofia District 343 344 1 1 Mahajanga holotype 2275276474 MOLC Madagascar -17.75 Toamasina 1277 48.716667 343 344 1 1 Toamasina holotype 2275276411 FSAE Madagascar -17.75 Chrome 1277 47.983334 343 344 1 1 Toamasina holotype 2275276485 [262,889,1809,1835] 1992-02-01 FSAE Madagascar Morarano-Chrome 343 344 1 1 Toamasina holotype 2275276750 [889,1282,1809,1835] ISNB Foulpointe Madagascar Toamasina 343 344 1 Toamasina holotype 2275276290 FSAE Ivondro Madagascar Toamasina 343 344 1 1 Toamasina holotype 2275276327 [392,1388,1844,1872] 1957-06 MNHN Pierre Andria Robinson Madagascar Andekaleka Rogez 343 344 1 1 Toamasina holotype 2275276296 2003-03-17 2003-03-21 2003-03-17 MNHN Fisher & Griswold Madagascar 600 -15.286333 Akirindro Mt. 13 49.54233 343 344 1 1 Toamasina holotype 2275275954 2005-11-28 CAS Brian L. Fisher Madagascar 390 -17.921 Camp Rendrirendry 13 49.193165 Betampona Reserve 343 344 2 2 Toamasina holotype 2275276294 2009-02-21 2009-02-23 2009-02-21 CAS B. L. Fisher Madagascar 780 344 345 -17.759167 Onibe River 21 48.85472 Zahamena National Park 343 344 1 1 Toamasina holotype 2275275869 1994-11-13 1994-11-21 1994-11-13 CAS B. L. Fisher Madagascar 1280 -14.75 Anjanaharibe-sud Reserve 1287 49.466667 344 345 1 1 Toamasina holotype 2275275974 DEUCD Bekily Madagascar Toliara 344 345 1 1 Toliara holotype 2275275959 [468,952,225,251] MNHN, MRAC Beroboka Madagascar Toliara 344 345 2 2 Toliara holotype 2275276616 2008-08-03 2008-08-11 2008-08-03 NHMUK Antsarongaza & M. Irwin & R. Harin'Hala Madagascar 128 -19.9775 Beroboka village 12 44.665333 344 345 1 1 Toliara holotype 2275275780 [531,896,297,323] MOLC Belalanda Madagascar Toliara 344 345 1 1 Toliara holotype 2275276475 2002-02-21 2002-02-25 2002-02-21 FSAE Fisher & Griswold Madagascar 80 -24.652166 Mahafaly Plateau 12 43.991333 344 345 1 1 Toliara holotype 2275275875 2009-07-03 2009-07-16 2009-07-03 MOLC M. Irwin & R. Harin'Hala Madagascar 130 -24.650333 Androimpano Forest 12 43.963165 344 345 1 1 Toliara holotype 2275275915 [1083,1389,405,431] CAS Behara Madagascar Toliara 344 345 1 1 Toliara holotype 2275276495 MRAC Fort-Dauphin Madagascar Toliara 344 345 1 1 Toliara holotype 2275276676 2003-05-18 2003-05-26 2003-05-18 MRAC M. Irwin & F. Parker & R. Harin'Hala Madagascar 37 -25.590557 Marie Special Reserve Cap 344 345 1 1 Toliara holotype 2275276015 [232,776,512,539] 2003-02-25 2003-03-07 2003-02-25 CAS M. Irwin & F. Parker & R. Harin'Hala Madagascar 37 -25.590557 Marie Special Reserve Cap 344 345 1 1 Toliara holotype 2275275945 2008-09-10 2008-09-17 2008-09-10 CAS, MOLC B. L. Fisher Madagascar 30 -20.069168 Foret de Kirindy 21 44.660557 344 345 3 3 Toliara holotype 2275276303 2002-09-23 2002-10-03 2002-09-23 CAS R. Harin'Hala Madagascar 30 -22.903667 Mikea Forest 12 43.4755 344 345 2 2 Toliara holotype 2275276169 2003-07-06 2003-07-17 2003-07-06 CAS R. Harin'Hala Madagascar 30 -22.903667 Mikea Forest 12 43.4755 344 345 1 1 Toliara holotype 2275276461 2003-10-30 2003-11-09 2003-10-30 CAS M. Irwin & F. Parker & R. Harin'Hala Madagascar 180 -24.936832 Tsimelahy 12 46.626667 Andohahela National Park 344 345 1 1 Toliara holotype 2275276775 [195,867,728,755] 1951-02-20 2004-01-15 1951-02-20 CAS M. Irwin & F. Parker & R. Harin'Hala Madagascar 180 -24.936832 Tsimelahy 12 46.626667 Andohahela National Park 344 345 1 1 Toliara holotype 2275275664 2003-03-23 2003-04-02 2003-03-23 CAS R. Harin'Hala Madagascar 165 -23.6865 Beza Mahafaly Reserve 12 44.591 344 345 1 1 Toliara holotype 2275275916 [531,1237,801,827] 2002-02-01 2002-02-08 2002-02-01 MOLC R. Harin'Hala Madagascar 165 -23.6865 Beza Mahafaly Reserve 12 44.591 344 345 1 1 Toliara holotype 2275276020 2002-02-08 2002-02-15 2002-02-08 CAS R. Harin'Hala Madagascar 165 -23.6865 Beza Mahafaly Reserve 12 44.591 344 345 1 1 Toliara holotype 2275275623 2002-03-18 2002-03-22 2002-03-18 CAS Fisher & Griswold Madagascar Tsimanampetsotsa National Park 40 -24.041666 Beheloka 12 43.751835 Mitoho Cave 344 345 1 1 Toliara holotype 2275275805 1922-02-19 2003-03-24 1922-02-19 CAS M. Irwin & F. Parker & Harin'Hala Madagascar 85 -25.006666 Berenty Special Reserve 12 46.303333 344 345 1 1 Toliara holotype 2275276392 [231,904,980,1007] 1922-02-15 2003-02-15 1922-02-15 CAS M. Irwin & F. Parker & Harin'Hala Madagascar 85 -25.006666 Berenty Special Reserve 12 46.303333 344 345 1 1 Toliara holotype 2275276946 1922-02-16 2003-02-25 1922-02-16 CAS M. Irwin & F. Parker & Harin'Hala Madagascar 85 -25.006666 Berenty Special Reserve 12 46.303333 344 345 1 1 Toliara holotype 2275276346 [325,1051,1017,1043] 2002-10-22 2002-12-27 2002-10-22 CAS M. Irwin & F. Parker & Harin'Hala Madagascar 85 -25.006666 Berenty Special Reserve 12 46.303333 344 345 1 1 Toliara holotype 2275275570 [1065,1387,1017,1043] CAS Berenty Madagascar Toliara 344 345 1 1 Toliara holotype 2275276776 1983-05-05 1983-05-15 1983-05-05 NHMUK J. S. Noyes & M. C. Day Madagascar Berenty 344 345 1 1 Toliara holotype 2275275930 2001-11-09 2001-11-19 2001-11-09 NHMUK M. E. Irwin & F. D. Parker & R. Harin'Hala Madagascar 840 -22.8865 Zombitse National Park 12 44.692165 344 345 2 2 Toliara holotype 2275275633 2009-02-09 2009-02-10 2009-02-09 CAS B. L. Fisher Madagascar 1270 -23.463612 Ampanihy 21 46.463055 Kalambatritra Special Reserve 344 345 1 1 Toliara holotype 2275276642 2003-10-27 2003-11-02 2003-10-27 CAS Frontier Wilderness Project Madagascar 20 -23.046383 Ranobe 1 43.610283 344 345 1 1 Toliara holotype 2275275444 2003-11-08 2003-11-15 2003-11-08 CAS Frontier Wilderness Project Madagascar 20 -23.03945 Ranobe 1 43.610283 344 345 1 1 Toliara holotype 2275276845 2003-08-05 2003-08-10 2003-08-05 CAS Frontier Wilderness Project Madagascar 65 -23.222517 Fiherenana 1 43.880882 344 345 1 1 Toliara holotype 2275276137 2002-10-31 2002-11-06 2002-10-31 CAS Frontier Wilderness Project Madagascar 100 -23.172832 Fiherenana 12 43.9565 344 345 1 1 Toliara holotype 2275276679 [605,1284,1377,1403] 2002-10-22 2002-10-31 2002-10-22 CAS Frontier Wilderness Project Madagascar 100 -23.172832 Fiherenana 12 43.9565 344 345 4 4 Toliara holotype 2275276382 2002-05-15 2002-05-29 2002-05-15 CAS, MOLC Frontier Wilderness Project Madagascar 70 -23.523167 Manderano 12 44.086334 344 345 2 2 Toliara holotype 2275276459 2010-11-24 2010-12-12 2010-11-24 CAS B. L. Fisher Madagascar 500 -21.2185 Makay Mts. 12 45.321 344 345 2 2 Toliara holotype 2275276386 2010-12-26 2011-01-03 2010-12-26 CAS M. Irwin & R. Harin'Hala Madagascar 600 -22.666666 National Park 1259 44.833332 344 345 1 1 Toliara holotype 2275276151 1990-03-08 CAS, SANC C. D. Eardley South Africa -22.183332 Altyddroog Farm 1261 29.883333 344 345 1 1 Limpopo holotype 2275275965 1998-01-24 1998-02-21 1998-01-24 S. van Noort & B. Fisher South Africa -34.456833 Walker Bay Nature Reserve 11 19.3565 344 345 1 Western Cape holotype 2275276437 2009-09-10 2009-11-04 2009-09-10 SAMC Rooiberg Sandstone Fynbos & S. van Noort South Africa 940 -33.72772 Gamkaberg Nature Reserve 1 21.96 344 345 1 1 Western Cape holotype 2275276479 [356,737,1701,1727] SAMC Uganda Uganda UGANDA 344 345 1 1 holotype 2275275209 2008-08-04 2008-08-26 2008-08-04 OLL S. van Noort Uganda National Park 1523 0.5639333 Makerere 1 30.361668 Kanyawara 344 345 1 1 Kibale holotype 2275276061 2002-08-21 2002-10-28 2002-08-21 SAMC A. van Harten Yemen Manakhah 344 345 2 2 holotype 2275277056 [474,619,1809,1835] MOLC Yemen 344 345 1 1 holotype