Ceballos 1936: 48
Castillo & Izquierdo 1989: 254
Dryinus saussurei (Ceballos)
Benoit 1954: 412
Olmi 1984: 770
1994c: 10
Azevedo et al . 2010: 891
Olmi et al . 2016: 76
Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea)
Olmi, Massimo
Copeland, Robert S.
Noort, Simon Van
Zootaxa
2019
2019-07-10
4630
1
1
619
Olmi et al . 2016: 76
(Ceballos, 1936)
Ceballos
1936
[195,622,1372,1398]
Insecta
Ampulicidae
Dryinus
CoL
Animalia
Hymenoptera
341
342
Arthropoda
species
saussurei
( Figs 140B–D)
Lestodryinus saussurei Ceballos 1936: 48; Rey del Castillo & Izquierdo 1989: 254. Dryinus saussurei(Ceballos): Benoit 1954: 412; Olmi 1984: 770; 1994c: 10; 2006: 43; Azevedo et al. 2010: 891; Olmi et al. 2016: 76.
Description. ♀. Fully winged; body length 4.0– 5.9 mm. Head black, except mandible and clypeus yellow; rarely clypeus black or brown; occasionally anterior region of frons testaceous; occasionally malar space with testaceous spot (in specimen from Botswana, Serowe); antenna testaceous; occasionally antennomeres 7–8, or 3–10, or 3–8 brown; propleuron black; pronotum black, with lateral regions and posterior collar reddish; rest of mesosoma black; metasoma black-brown; legs testaceous, except coxae and clubs of femora partly brown. Antenna clavate; antennomeres in following proportions: 10:5:21:10:9:8:6:6:4.5:8. Head ( Fig. 140B) shiny or dull, completely or partly granulate, occasionally partly unsculptured; occasionally head partly granulate and partly sculptured by a few irregular keels; frontal line complete; occipital carina complete; lateral ocelli usually touching occipital carina, occasionally very close, but not touching occipital carina; OL = POL. In specimen from Madagascar, 17°45’S 47°59’E, POL = 4; OL = 2. Pronotum dull, partly or completely granulate, with numerous slight striae around disc; occasionally pronotum shiny, almost completely unsculptured; posterior collar of pronotum short, about 0.5 × as long as distance from tegula to pronotal tubercle; pronotal tubercle not reaching tegula. Mesoscutum dull, slightly or strongly granulate, with short keels or areolae near posterior margin; occasionally median region of mesoscutum shiny, unsculptured (in specimen from Cameroon, Nkoemvon, mesoscutum completely granulate, with no areolae or keels; in specimen from South Africaposterior half of mesoscutum completely reticulate rugose; in specimen from Botswana, Serowe, mesoscutum almost completely reticulate rugose; in specimen from Cameroon, Messabengondo, mesoscutum completely granulate and rugose). Notauli incomplete, approximately reaching 0.65–0.90 × length of mesoscutum (rarely 0.5 × length of mesoscutum; rarely notauli complete and posteriorly separated). Mesoscutellum dull, granulate, or shiny and unsculptured. Metanotum dull, rugose. Metapectal-propodeal disc reticulate rugose; propodeal declivity reticulate rugose, with two longitudinal keels. Forewing with three brown transverse bands (two proximal bands almost joint, occasionally completely joint); distal part of 2r-rs&Rs vein about twice as long as proximal part (18:8). Hind wing hyaline, without dark spots; occasionally (in very large specimens) with one dark spot. Protarsomere 1 approximately as long as 4; in specimens from Madagascar, 19°07.56’S 44°48.53’Eand 18°01.35’S 44°03.02’E, protarsomere 1 longer than 4; protarsomere 3 produced into hook. Enlarged claw ( Fig. 140D) with one big subdistal tooth and one row of 9–13 lamellae (in specimen from Madagascar, 18°01.35’S 44°03.02’E, subapical tooth distinctly divided into two teeth ( Fig. 140C), so that apparently this specimen has to be included in group 3 of Dryinus). Protarsomere 5 ( Fig. 140D) with two rows of 16–39 lamellae; apex with 11–16 lamellae. Tibial spurs 1/1/2. ♂. Fully winged; body length 2.6–2.8 mm. Head black, except mandible testaceous; antenna brown, except scape and pedicel testaceous; mesosoma black; metasoma brown; legs testaceous, except coxae, club of metafemur and metatibia brown. Antenna filiform; antennomeres in following proportions: 8:7:17:12:13:13:11:10:10:11. Head dull, granulate; frons flat in front of anterior ocellus and humped between antennal toruli; frontal line incomplete; occipital carina incomplete; POL = 9; OL = 3; OOL = 3; lateral ocelli touching occipital carina; temple absent. Mesoscutum dull, granulate. Notauli incomplete, reaching approximately 0.65 × length of mesoscutum. Mesoscutellum almost completely dull, granulate, with narrow posterior region smooth and shiny. Metanotum with anterior half rugose and posterior half shiny and smooth. Metapectal-propodeal disc reticulate rugose, without transverse posterior keel; propodeal declivity reticulate rugose, without longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2r-rs&Rs vein slightly longer than proximal part (14:13); 2R1 cell open. Paramere without dorsal process. Tibial spurs 1/1/2.
Material examined. Type: ♀ holotype: MADAGASCAR: Toamasina, Rogez(= Andekaleka), I.1931, A. Sey- rig leg. ( MNMS). Other material: BOTSWANA: Central District, Serowe, XI.1985, MT, P. Forchhammerleg., 1♀( NHMUK). CAMEROON: South Region, near Ebolowa, Nkoemvon, 30.XI.1979– 5.I.1980, MT in forest, D. Jacksonleg., NHMUK (E) 2007–19, 1♀( NHMUK); same locality label, 23.IX–25.X.1980, 1♀( NHMUK); Messaben- gondo, 1–15.III.2003, M. Tussacleg., 1♀( HTS). ETHIOPIA: Shoa Prov., 95 kmW Addis Ababa, Ambo, near Plant Protection Research Institute, 2400–2500 m, 19.VI.1990, A. Emel’yanovleg., 1♀( ZISP). MADAGASCAR: Anta- nanarivo, La Mandraka, 1♀( MRAC); same locality label, II.1953, Pierre Andria Robinsonleg., 1♀( MNHN); An- tananarivo, Analamanga Region, Ambatolaona, VI.1957, Pierre Andria Robinsonleg., 1♀( MNHN); Antananarivo, Andebadeba, 4 kmfrom Arivanimamo, 19°00’29’’S 47°08’55’’E, 1600 m, 7–14.X.2007, Uapaca forest, MT, M. Irwin, R. Harin’Halaleg., MA-46-02, 1♀( CAS); Antananarivo, Ankokoy Forest, 3 kmE of Ibity, 20°04.05’S 46°59.97’E, 1700 m, 12–22.XII.2008, MT, in Uapaca forest, M. Irwin, R. Harin’Halaleg., MG-56-05, 1♀( CAS); Antsiranana, Nosy Bé Island, 1♀( HNHM); Antsiranana, Manongativo Reserve, 19.8 km229° SW Antanambao, 13°58’S 48°26’E, 400 m, 8–13.XI.1998, B.L. Fisherleg., 1♀( DEUCD); Antsiranana, Marojejy National Park, 5 kmW Manantenina village, Camp Mantella, 14°26.29’S 49°46.44’E, 490 m, 28.IV–7.V.2005, MT, low altitude rainforest, M. Irwin& R. Harin’Halaleg., MA-31-20, 1♀( CAS); Antsiranana, Marojejy National Park, Manantenina River, 27.6 km35° NE Andapa, 9.6 kmNNW Manantenina village, 14°26.06’S 49°45.36’E, 775 m, 15–18.XI.2003, YPT, rainforest, BLF8873, B.L. Fisher et al. leg., 1♀( CAS); Antsiranana, Marojejy National Park, Manantenina River, 28.0 km 38° NE Andapa, 8.2 kmNNW Manantenina village, 14°26.12’S 49°46.30’E, 450 m, 12–25.XI.2003, YPT, rainforest, BLF8725, B.L. Fisher et al. leg., 2♀♀( CAS, MOLC); Antsiranana, 7 kmN of Joffreville, 12°27’S 49°14’E, 426 m, dryforest, MT, M. Irwin, R. Harin’Halaleg., MA- 01-07-16, 1♀( CAS); Antsiranana, Montagne d’Ambre National Park, 12°30’52’’S 49°10’53’’E, 960 m, 19.III–6.IV.2001, MT, R. Harin’Halaleg., MA-01-01A- 09, 1♀( MOLC); same locality label, 4–19.III.2001, MA-01-01A-08, 1♀( CAS); same locality label, 21–28.I.2001, MT, M.E. Irwin, E.I. Schlinger& R. Harin’Halaleg., MA-01-01A-01, 1♀( CAS); Antsiranana, Montagne d’Ambre National Park, 12°31.13’S 49°10.45’E, 1125 m, 14–30.V.2001, MT, R. Harin’Halaleg., MA-01-01D-11, 1♀( CAS); Fianarantsoa, Haute-Matsiatra Region, Andringitra-Ambalavao, Cirque Boby, 2500 m, 13.I.1958, R. Paulianleg., 1♀( MNHN); Fianarantsoa, Ranomafana National Park, Talatakely, 21°15’S 47°25’E, 850 m, 14.IV.1998, 1♀( CAS); Fianarantsoa, Ranomafana, 15.III.1994, 1♀( ISNB); 7 kmW of Ranomafana, 900 m, 2♀♀( FSAE, USNM); Fianarantsoa, Ranomafana National Park, Vohiparara, at broken bridge, 21°13.57’S 47°22.19’E, 1110 m, 4– 12.II.2002, MT, high altitude rainforest, R. Harin’Halaleg., MA-02-09A-15, 1♀( CAS); Fianarantsoa, Alakamisy, 21.XI.2006, in forest, C. Caneparileg., 1♀( MCSNG); Fianarantsoa, Ranomafana National Park, Belle Vueat Ta- latakely, 21°15.99’S 47°25.21’E, 1020 m, 26.II–10.III.2003, MT, secondary tropical forest, R. Harin’Halaleg., MA-02-09C-55, 1♀( MOLC); same locality label, 14–21.I.2002, MA-02-09C-12, 1♀( CAS); same locality label 22–28.XI.2001, MA-02-09C-04, 1♀( CAS); Fianarantsoa, Ranomafana National Park, radio tower at forest edge, 21°15.05’S 47°24.43’E, 1130 m, 12–19.III.2002, MT, mixed tropical forest, R. Harin’Halaleg., MA-02-09B-20, 1♀( MOLC); same locality label, 15–21.XII.2001, MA-02-09B-07, 1♀( CAS); same locality label, 22–28.XI.2001, MA-02-09B-04, 1♀( MOLC); same locality label, 27.IV–7.V.2003, MA-02-09B-59, 1♀( CAS); Fianarantsoa, Be- fotaka–Midongy National Park, Papango, 27.7 kmS Midongy-South Mt. Papango, 23°50.07’S 46°57.49’E, 940 m, 13–19.XI.2006, MT, B.L. Fisher et al. leg., BLF14774, 1♀( CAS); Fianarantsoa, Agnalazaha Forest Reserve, Ma- habo, 42.9 km215° SW of Farafangana, 23°11.38’S 47°43.23’E, 20 m, 18.IV.2006, MT, littoral rainforest, B.L. Fisheret al. leg., BLF13822, 1♀( CAS); Fianarantsoa, 50 kmS of Farafangana, Mahabo Mananivo, Ampitavanan- ima Forest, 23°07’79’’S 47°43’02’’E, 34 m, 26.I–4.II.2007, MT, in low altitude littoral forest, M. Irwin, F. Parker, R. Harin’Halaleg., MG-35-03, 1♀( CAS); Fianarantsoa, Miandritsara Forest, 40 kmS Ambositra, 20°47.56’S 47°10.54’E, 825 m, 25.X–4.XI.2006, MT, low altitude rainforest, M. Irwin, R. Harin’Halaleg., MA-29-56, 1♀( CAS); Fianarantsoa, near Isalo National Park, at stream east of Interpretive Center, 22°37.60’S 45°21.49’E, 750 m, 10–17.III.2002, MT, in open area, MA-02-11A-19, R. Harin’Halaleg., 1♀( CAS); same locality label, 15–22.IV.2002, MA-02-11A-24, 1♀( CAS); same locality label, 30.XI–7.XII.2001, MA-02-11A-05, 1♀( CAS); same locality label, 19–26.I.2002, MA-02-11A-12, 1♀( CAS); Fianarantsoa, near Isalo National Park, dry wash east of Interpretive Center, 22°37.60’ 45°21.49’E, 885 m, 9–23.VIII.2002, MT, in open area, MA-02-11B-37, R. Harin’Halaleg., 1♀( CAS); same locality label, 7–22.IX.2002, MA-02-11B-39, 2♀♀( CAS); same locality label, 3–17.X.2002, MA-02- 11B-41, 1♀( CAS); Fianarantsoa, Analalava Forest, 29.6 km280° W Ranohira, 22°35.30’S 45°07.42’E, 700 m, 1– 5.II.2003, tropical dry forest, beating low vegetation, Fisher, Griswold et al. leg., BLF7384, 1♀( CAS); Mahajanga, Tsingy de Bemaraha National Park, 2.5 km62° ENE Bekopaka, Ankidrodroa River, 19°07.56’S 44°48.53’E, 100 m, 11–15.XI.2001, sifted litter, tropical dry forest on Tsingy, Fisher, Griswold et al. leg., BLF 4340, 1♀( CAS); Maha- janga, Bale de Baly National Park, 12.4 km337° NNW Soalala, 16°00.36’S 45°15.54’E, 10 m, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 6811, 2♀♀( MOLC); Mahajanga, Beanka Forest Reserve, 50.2 kmE Main- tirano, 18°01.35’S 44°03.02’E, 250 m, 19–26.X.2009, YPT, tropical dry forest on tsingy, B.L. Fisher et al. leg., BLF22788, 3♀♀( 2 in CAS, 1 in MOLC); Mahajanga, Maintirano District, Asondrodavadry forest, 15 kmN of Maintirano, 17°57.92’S 44°2.13’E, 200 ft., 7–14.XII.2007, MT, dry forest, M. Irwin, R. Harin’Halaleg., MG-41-11, 1♀( CAS); same locality label, 3–10.XII.2007, MG-43A-09, 1♀( CAS); same locality label, 8–15.X.2007, 1♀( CAS); Mahajanga, Analamanitra Forest, 14 kmNE Misinjo, 15°55’S 45°57’E, 65 ft., 23–30.X.2007, MT, dense dry forest, M. Irwin, R. Harin’Halaleg., MG-38-06, 1♀( CAS); same locality label, 20.XI–4.XII.2007, MG-38-09, 2♀♀( CAS); Mahajanga, Ambatofolaka, Namoroka, 53 kmfrom Soalala, 3 kmN Vilanandro village, 16°28’04’’S 45°23’48’’E, 400 ft., 21–28.IX.2007, dense dry forest, MT, M. Irwin, R. Harin’Halaleg., MG-40A-01, 1♀( CAS); Mahajanga, Ampijoroa National Park, 160 kmN of Maevatananaon RN 04, 16°19’16’’S 46°48’80’’E, 43 m, 17– 24.VIII.2003, MT, deciduous forest, R. Harin’Halaleg., MA-25-11, 1♀( CAS); same locality label, 12–19.X.2003, MA-25-19, 1♀( CAS); same locality label, 28.IX–5.X.2003, MA-25-17, 2♀♀( CAS); Mahajanga, NamorokaNa- tional Park, 16.9 km317° NW Vilanandro, 16°24.24’S 45°18.36’E, 100 m, 12–16.XI.2002, YPT, tropical dry forest, Fisher, Griswold et al. leg., BLF 6536, 2♀♀( CAS, MOLC); Mahajanga, Namoroka National Park, 16.40494S 45.31294E, 21–31.X.2016, MT, C. Villemantleg., 1♀( MNHN); Mahajanga, Namoroka National Park, 23– 25.X.2016, YPT, C. Villemantleg., 1♀( MNHN); Mahajanga, Sofia District, 45 kmS Antsohihy, 5 kmW Anjiaman- girana, Analagnambedry degraded forest, 15°09.42’S 47°47.05’E, 97 m, 2–6.I.2011, MT, MG-61-04, 1♀( MOLC); Toamasina, Manakambahiny-E, 17°45’S 48°43’E, 1♀( FSAE); Toamasina, 25 kmW of Morarano- Chrome, 17°45’S 47°59’E, 1♀( FSAE); Toamasina, Morarano-Chrome, II.1992, 1 ♀ ( ISNB); Toamasina, Foulpointe, 1♀( FSAE); Toamasina, Ivondro, 1♀( MNHN); Toamasina, Rogez(= Andekaleka), VI.1957, Pierre Andria Robinsonleg., 1♀( MNHN); Toamasina, Akirindro Mt., 7.6 km341° NNW Ambinanitelo, 15°17.18’S 49°32.54’E. 600 m, 17– 21.III.2003, YPT, rainforest, Fisher, Griswold et al. leg., BLF8255, 2♀♀( CAS); Toamasina, Betampona Reserve, Camp Rendrirendry, 34.1 km332° Toamasina, 17°55.26’S 49°11.59’E, 390 m, 28.XI.2005, YPT, rainforest, Brian L. Fisher et al. leg., BLF13124, 1♀( CAS); Toamasina, Zahamena National Park, Onibe River, 17°45’33’’S 48°51’17’’E, 780 m, 21–23.II.2009, rainforest, YPT, BLF22170, B.L. Fisher et al. leg., 1♀( CAS); Toamasina, 9.2 kmWSW Befingotra, Anjanaharibe-sud Reserve, 14°45’S 49°28’E, 1280 m, 13–21.XI.1994, B.L. Fisherleg., 1♀( DEUCD); Toliara, Bekily, 2♀♀( MNHN, MRAC); Toliara, Beroboka, 1♀( NHMUK); Toliara, Beroboka village, 45 kmNE Morondava, 19°58.65’S 44°39.92’E, 420 ft., 3–11.VIII.2008, MT, Antsarongazagallery forest, MG-45B- 33, M. Irwin, R. Harin’Halaleg., 1♀( MOLC); Toliara, Belalanda, 1♀( FSAE); Toliara, Mahafaly Plateau, 6.2 km74° ENE of Itampolo, 24°39.13’S 43°59.48’E, 80 m, MT, spiny forest thicket, 21–25.II.2002, Fisher, Griswold et al. leg., BLF5759, 1♀( MOLC); Toliara, Androimpano Forest, 5 kmE of Itampolo, 24°39.02’S 43°57.79’E, 130 m, 3–16.VII.2009, MT, path in dry forest, M. Irwin, R. Harin’Halaleg., MG-54C-33, 1♀( CAS); Toliara, Behara, 1♀( MRAC); Toliara, Fort-Dauphin, 1♀( MRAC); Toliara, CapS.te Marie Special Reserve, 74 kmS of Tsihombe, 25°35’26’’S 45°09’78’’E, 37 m, 18–26.V.2003, MT, spiny bush, M. Irwin, F. Parker& R. Harin’Halaleg., MA- 02- 23-27, 1♀( CAS); same locality label, 25.II–7.III.2003, 3♀♀( 2 in CAS, 1 in MOLC); Toliara, Forêt de Kirindy, 15.5 km64° ENE Marofandilia, 10–17 September 2008, 20°04’09’’S 044°39’38’’E, B.L. Fisher, MT, tropical dry forest, 30 m, BLF18285_45, 2♀♀( CAS); Toliara, Mikea Forest, NW of Manombo, 22°54.22’S 43°28.53’E, 30 m, 23.IX–3.X.2002, MT, deciduous dry forest, R. Harin’Halaleg., MA-02-18A-34, 1♀( CAS); same locality label, 6–17.VII.2003, MA-02-18A-61, 1♀( CAS); Toliara, Andohahela National Park, Tsimelahy, parcelle II, 24°56.21’S 46°37.60’E, 180 m, 30.X–9.XI.2003, MT, transitional forest, M. Irwin, F. Parker& R. Harin’Halaleg., MA- 02-20- 44, 1♀( CAS); same locality label, 5–15.I.2004, MA- 02-20-51, 1♀( CAS); Toliara, Beza Mahafaly Reserve, par- celle I near research station, 23°41.19’S 44°35.46’E, 165 m, 23.III–2.IV.2003, MT, dry deciduous forest, R. Harin’Halaleg., MA-02-14A-54, 1♀( MOLC); same locality label, 1–8.II.2002, MA-02-14A-14, 1♀( CAS); same locality label, 8-15.II.2002, MA-02-14A-15, 1♀( CAS); Toliara, Tsimanampetsotsa National Park, Mitoho Cave, 6.4 km77° ENE Efoetse, 17.4 km170° S Beheloka, 24°02.50’S 43°45.11’E, 40 m, 18–22.III.2002, MT, spiny forest thicket, Fisher, Griswold et al. leg., BLF 6159, 1♀( CAS); Toliara, Berenty Special Reserve, 8 kmNW Amboasary, 25°00.40’S 46°18.20’E, elev 85 m, 17–24.III.2003, M.Irwin, F. Parker& R. Harin’Hala, MT, spiny forest, MA- 02- 22-19, 1♀( CAS); same locality label, 5–15.II.2003, MA- 02-22-15, 1♀( CAS); same locality label, 15–25.II.2003, MA- 02-22-16, 1♀( CAS); same locality label, 16–27.XII.2002, MA- 02-22-10, 1♀( CAS); Toliara, Berenty, 1♀( NHMUK); Toliara, Berenty, 12 kmNW Amboasary, 5–15.V.1983, J.S. Noyes& M.C. Dayleg., 2♀♀( NHMUK); Toliara, Zombitse National Park, near ANGAP office, 22°53.19’S 44°41.53’E, 840 m, 9–19.XI.2001, MT, deciduous spiny forest, M.E. Irwin, F.D. Parker& R. Harin’Halaleg., MA-02-13A-02, 1♀( CAS); Toliara, Kalambatritra Special Reserve, Ampanihy, 23°27’49’’S 46°27’47’’E, 1270 m, 9–10.II.2009, YPT, montane rainforest, B.L. Fisher et al. leg., BLF21567, 1♀( CAS); Toliara, Ranobe, 23°02.783’S 43°36.617’E, 20 m, 27.X–2.XI.2003, MT, spiny forest thicket, MGF081, Frontier Wilderness Project, 1♀( CAS); Toliara, Ranobe, 23°02.367’S 43°36.617’E, 20 m, 8–15.XI.2003,MT, gallery forest,MGF084, Frontier Wilderness Project, 1♀( CAS); Toliara, Fiherenana, 23°13.351’S 43°52.853’E, 65 m, 5–10.VIII.2003, MT, degraded riparian forest close to water, MGF075, Frontier Wilderness Project, 1♀( CAS); Toliara, Fiherenana, 23°10.37’S 43°57.39’E, 100 m, 31.X–6.XI.2002, MT, gallery forest, MGF045, Frontier Wilderness Project, 4♀♀( CAS); same locality label, 22–31.X.2002, MGF042, 2♀♀( CAS, MOLC); Toliara, Manderano, 23°31.39’S 44°5.18’E, 70 m, 15–29.V.2002, MT, edge of marsh, 5 mfrom road, gallery forest, MGF032, Frontier Wilderness Project, 2♀♀( CAS); Toliara, Makay Mts., 21°13.11’S, 45°19.26’E, 500 m, 24.XI-12.XII.2010, MT, gallery forest, sandy soil, B.L. Fisher et al. leg., BLF25357, 1♀( CAS); Toliara, Vohi- basia National Park, 44 kmNE of Sakaraha, Ambakakagallery/palm forest, 22°40’S 44°50’E, 600 m, 26.XII.2010- 3.I.2011, M. Irwin, R. Harin’Halaleg., MG-60-12, 1♀( CAS). SOUTH AFRICA: Limpopo, 13 kmW of Beitbridge, Altyddroog Farm, 22°11’S 29°53’E, 8.III.1990, C.D. Eardleyleg. ( SANC); Western Cape, Walker BayNature Reserve, 34°27.41’S 19°21.39’E, 24.I–21.II.1998, YPT, S. van Noort& B. Fisherleg., 1♀( SAMC); Western Cape, Gamkaberg Nature Reserve, 33°43.663‘S 21°57.600‘E, 940 m, 10.IX–4.XI.2009, MT, Rooiberg Sandstone Fynbos, S. van Noortleg., 1♀( SAMC). UGANDA: Uganda, 1♀( OLL); Kibale National Park, Kanyawara, MakerereUni- versity Biological Field Station, 0°33.836‘N 30°21.700‘E, 1523 m, 4–26.VIII.2008, YPT, primary mid–altitude rainforest, near stream, S. van Noortleg., 2♀♀( SAMC). YEMEN: 12 kmNW Manakhah, 21.VIII–28.X.2002, MT, N. 7143, A. van Hartenleg., 1♀( MOLC).
Hosts. Unknown.
Distribution. Botswana, Cameroon, Ethiopia, Madagascar, South Africa, Uganda, Yemen.
2275277563
1931-01
1985-11
1931-01
MNMS
P. Forchhammer
Madagascar
Rogez
Serowe
Andekaleka
342
343
2
1
Central
holotype
2275277135
1979-11-30
1980-01-05
1979-11-30
NHMUK
D. Jackson
Cameroon
South Region
Nkoemvon
Ebolowa
342
343
1
1
holotype
2275276253
[519,1123,1196,1223]
1980-09-23
1980-10-25
1980-09-23
NHMUK
D. Jackson
Cameroon
South Region
Nkoemvon
Ebolowa
342
343
1
1
holotype
2275276045
2003-03-01
2003-03-15
2003-03-01
NHMUK
M. Tussac
Cameroon
342
343
1
1
holotype
2275277054
1990-06-19
HTS
A. Emel'yanov
Ethiopia
Shoa Prov.
2450
Plant Protection Research Institute
Ambo
342
343
1
1
Adis Abeba
holotype
2275275899
ZISP
La Mandraka
Madagascar
MADAGASCAR
342
343
1
1
holotype
2275276540
[428,1221,1304,1331]
1953-02
MRAC
Pierre Andria Robinson
Madagascar
MADAGASCAR
342
343
1
1
holotype
2275276688
1957-06
MNHN
Pierre Andria Robinson
Madagascar
Ambatolaona
Analamanga Region
342
343
1
1
holotype
2275276449
2007-10-07
2007-10-14
2007-10-07
MNHN
M. Irwin & R. Harin'Hala
Madagascar
Andebadeba
1600
-19.008055
Uapaca forest
21
47.148613
Arivanimamo
342
343
1
1
Antananarivo
holotype
2275275851
2008-12-12
2008-12-22
2008-12-12
CAS
M. Irwin & R. Harin'Hala
Madagascar
1700
-20.0675
Uapaca forest
12
46.9995
Ankokoy Forest
342
343
1
1
Antananarivo
holotype
2275276662
CAS
Madagascar
Nosy Be Island
342
343
1
1
Antsiranana
holotype
2275276611
1998-11-08
1998-11-13
1998-11-08
HNHM
B. L. Fisher
Madagascar
400
-13.966666
Manongativo Reserve
1289
48.433334
342
343
1
1
Antsiranana
holotype
2275276352
2005-04-28
2005-05-07
2005-04-28
DEUCD
M. Irwin & R. Harin'Hala
Madagascar
490
-14.438167
Camp Mantella
13
49.774
Marojejy National Park
342
343
1
1
Antsiranana
holotype
2275276548
2003-11-15
2003-11-18
2003-11-15
CAS
B. L. Fisher
Madagascar
Marojejy National Park
775
-14.434334
Manantenina village
13
49.756
Manantenina River
342
343
1
1
Antsiranana
holotype
2275276501
2003-11-12
2003-11-25
2003-11-12
CAS
B. L. Fisher
Madagascar
Marojejy National Park
450
-14.435333
Andapa
13
49.771667
Manantenina River
342
343
1
1
Antsiranana
holotype
2275276199
CAS, MOLC
M. Irwin & R. Harin'Hala
Madagascar
426
-12.45
7 km N of Joffreville
1293
49.233334
342
343
2
2
Antsiranana
holotype
2275277011
2001-03-19
2001-04-06
2001-03-19
CAS
R. Harin'Hala
Madagascar
960
-12.514444
Montagne d'Ambre National Park
21
49.18139
342
343
1
1
Antsiranana
holotype
2275276685
[193,918,1845,1871]
2001-03-04
2001-03-19
2001-03-04
MOLC
R. Harin'Hala
Madagascar
960
-12.514444
Montagne d'Ambre National Park
21
49.18139
342
343
1
1
Antsiranana
holotype
2275276254
2001-01-21
2001-01-28
2001-01-21
CAS
M. E. Irwin & E. I. Schlinger & R. Harin'Hala
Madagascar
960
-12.514444
Montagne d'Ambre National Park
21
49.18139
342
343
1
1
Antsiranana
holotype
2275276715
2001-05-14
2001-05-30
2001-05-14
CAS
R. Harin'Hala
Madagascar
1125
-12.518833
Montagne d'Ambre National Park
13
49.174168
342
343
1
1
Antsiranana
holotype
2275276720
1958-01-13
CAS
R. Paulian
Madagascar
Haute-Matsiatra Region
2500
Cirque Boby
Andringitra-Ambalavao
342
343
1
1
Fianarantsoa
holotype
2275276986
[151,1385,1988,2015]
1998-04-14
MNHN
Madagascar
850
-21.25
Talatakely
1264
47.416668
Ranomafana National Park
342
343
1
1
Fianarantsoa
holotype
2275276013
1994-03-15
CAS
Madagascar
343
344
Ranomafana
342
343
1
1
Fianarantsoa
holotype
2275276491
[688,1174,153,179]
ISNB
Madagascar
900
Ranomafana
343
344
1
1
Fianarantsoa
holotype
2275276465
2002-02-04
2002-02-12
2002-02-04
FSAE, USNM
R. Harin'Hala
Madagascar
1110
-21.226166
Vohiparara
12
47.369835
Ranomafana National Park
343
344
2
2
Fianarantsoa
holotype
2275276115
2006-11-21
CAS
C. Canepari
Madagascar
Alakamisy
343
344
1
1
Fianarantsoa
holotype
2275275820
2003-02-26
2003-03-10
2003-02-26
MCSNG
R. Harin'Hala
Madagascar
1020
-21.2665
Belle Vue
12
47.420166
Ranomafana National Park
343
344
1
1
Fianarantsoa
holotype
2275275619
[343,1076,333,359]
2002-01-14
2002-01-21
2002-01-14
MOLC
R. Harin'Hala
Madagascar
1020
-21.2665
Belle Vue
12
47.420166
Ranomafana National Park
343
344
1
1
Fianarantsoa
holotype
2275277536
2001-11-22
2001-11-28
2001-11-22
CAS
R. Harin'Hala
Madagascar
1020
-21.2665
Belle Vue
12
47.420166
Ranomafana National Park
343
344
1
1
Fianarantsoa
holotype
2275276230
2002-03-12
2002-03-19
2002-03-12
CAS
R. Harin'Hala
Madagascar
1130
-21.250834
Ranomafana National Park
12
47.407166
343
344
1
1
Fianarantsoa
holotype
2275277340
2001-12-15
2001-12-21
2001-12-15
MOLC
R. Harin'Hala
Madagascar
1130
-21.250834
Ranomafana National Park
12
47.407166
343
344
1
1
Fianarantsoa
holotype
2275276046
2001-11-22
2001-11-28
2001-11-22
CAS
R. Harin'Hala
Madagascar
1130
-21.250834
Ranomafana National Park
12
47.407166
343
344
1
1
Fianarantsoa
holotype
2275277406
[341,1095,476,503]
2003-04-27
2003-05-07
2003-04-27
MOLC
R. Harin'Hala
Madagascar
1130
-21.250834
Ranomafana National Park
12
47.407166
343
344
1
1
Fianarantsoa
holotype
2275276543
2006-11-13
2006-11-19
2006-11-13
CAS
B. L. Fisher
Madagascar
940
-23.8345
Papango
12
46.958168
National Park
343
344
1
1
Fianarantsoa
holotype
2275276123
2006-04-18
CAS
B. L. Fisher
Madagascar
20
-23.189667
Agnalazaha Forest Reserve
12
47.7205
343
344
1
1
Fianarantsoa
holotype
2275276955
2007-01-26
2007-02-04
2007-01-26
CAS
M. Irwin & F. Parker & R. Harin'Hala
Madagascar
34
Forest
47.717224
Mahabo Mananivo
343
344
1
1
Fianarantsoa
holotype
2275275554
2006-10-25
2006-11-04
2006-10-25
CAS
M. Irwin & R. Harin'Hala
Madagascar
825
-20.792667
Miandritsara Forest
12
47.175667
343
344
1
1
Fianarantsoa
holotype
2275276291
2002-03-10
2002-03-17
2002-03-10
CAS
R. Harin'Hala
Madagascar
750
-22.626667
Interpretive Center
12
45.358166
Isalo National Park
343
344
1
1
Fianarantsoa
holotype
2275276234
2002-04-15
2002-04-22
2002-04-15
CAS
R. Harin'Hala
Madagascar
750
-22.626667
Interpretive Center
12
45.358166
Isalo National Park
343
344
1
1
Fianarantsoa
holotype
2275276551
[340,1082,836,863]
2001-11-30
2001-12-07
2001-11-30
CAS
R. Harin'Hala
Madagascar
750
-22.626667
Interpretive Center
12
45.358166
Isalo National Park
343
344
1
1
Fianarantsoa
holotype
2275276687
2002-01-19
2002-01-26
2002-01-19
CAS
R. Harin'Hala
Madagascar
750
-22.626667
Interpretive Center
12
45.358166
Isalo National Park
343
344
1
1
Fianarantsoa
holotype
2275276241
2002-08-09
2002-08-23
2002-08-09
CAS
R. Harin'Hala
Madagascar
885
Interpretive Center
45.358166
Isalo National Park
343
344
1
1
Fianarantsoa
holotype
2275276452
2002-09-07
2002-09-22
2002-09-07
CAS
R. Harin'Hala
Madagascar
885
Interpretive Center
45.358166
Isalo National Park
343
344
1
1
Fianarantsoa
holotype
2275275713
2002-10-03
2002-10-17
2002-10-03
CAS
R. Harin'Hala
Madagascar
885
Interpretive Center
45.358166
Isalo National Park
343
344
2
2
Fianarantsoa
holotype
2275276697
2003-02-01
2003-02-05
2003-02-01
CAS
Fisher & Griswold
Madagascar
700
-22.588333
Analalava Forest
12
45.123665
343
344
1
1
Fianarantsoa
holotype
2275275984
2001-11-11
2001-11-15
2001-11-11
CAS
Tsingy & Fisher & Griswold
Madagascar
100
-19.126
Ankidrodroa River
13
44.808834
Tsingy de Bemaraha National Park
343
344
1
1
Mahajanga
holotype
2275275933
CAS
Fisher & Griswold
Madagascar
10
-16.006
Bale de Baly National Park
13
45.259
343
344
1
1
Mahajanga
holotype
2275276617
2009-10-19
2009-10-26
2009-10-19
MOLC
B. L. Fisher
Madagascar
250
-18.0225
Beanka Forest Reserve
13
44.050335
343
344
2
2
Mahajanga
holotype
2275276648
2007-12-07
2007-12-14
2007-12-07
CAS, MOLC
M. Irwin & R. Harin'Hala
Madagascar
61
-17.965334
Asondrodava
13
44.0355
Maintirano District
343
344
3
3
Mahajanga
holotype
2275276700
[151,849,1305,1331]
2007-12-03
2007-12-10
2007-12-03
CAS
M. Irwin & R. Harin'Hala
Madagascar
61
-17.965334
Asondrodava
13
44.0355
Maintirano District
343
344
1
1
Mahajanga
holotype
2275276939
[864,1386,1304,1331]
2007-10-08
2007-10-15
2007-10-08
CAS
M. Irwin & R. Harin'Hala
Madagascar
61
-17.965334
Asondrodava
13
44.0355
Maintirano District
343
344
1
1
Mahajanga
holotype
2275276185
2007-10-23
2007-10-30
2007-10-23
CAS
M. Irwin & R. Harin'Hala
Madagascar
20
-15.916667
Analamanitra Forest
1284
45.95
343
344
1
1
Mahajanga
holotype
2275276806
[741,1432,1377,1403]
2007-11-20
2007-12-04
2007-11-20
CAS
M. Irwin & R. Harin'Hala
Madagascar
20
-15.916667
Analamanitra Forest
1284
45.95
343
344
1
1
Mahajanga
holotype
2275276496
2007-09-21
2007-09-28
2007-09-21
CAS
M. Irwin & R. Harin'Hala
Madagascar
Ambatofolaka
122
-16.46778
Soalala
21
45.396667
Namoroka
343
344
2
2
Mahajanga
holotype
2275275707
2003-08-17
2003-08-24
2003-08-17
CAS
R. Harin'Hala
Madagascar
43
-16.321112
Ampijoroa National Park
343
344
1
1
Mahajanga
holotype
2275276716
2003-10-12
2003-10-19
2003-10-12
CAS
R. Harin'Hala
Madagascar
43
-16.321112
Ampijoroa National Park
343
344
1
1
Mahajanga
holotype
2275275760
[286,964,1556,1583]
2003-09-28
2003-10-05
2003-09-28
CAS
R. Harin'Hala
Madagascar
43
-16.321112
Ampijoroa National Park
343
344
1
1
Mahajanga
holotype
2275276713
2002-11-12
2002-11-16
2002-11-12
CAS
Fisher & Griswold
Madagascar
100
-16.404
Park
13
45.306
Namoroka
343
344
2
2
Mahajanga
holotype
2275276585
2016-10-21
2016-10-31
2016-10-21
CAS, MOLC
C. Villemant
Madagascar
-16.40494
Namoroka National Park
1
45.31294
343
344
2
2
Mahajanga
holotype
2275277120
2016-10-23
2016-10-25
2016-10-23
MNHN
C. Villemant
Madagascar
Namoroka National Park
343
344
1
1
Mahajanga
holotype
2275276110
2011-01-02
2011-01-06
2011-01-02
MNHN
Madagascar
97
-15.157
Analagnambe
13
47.78417
Sofia District
343
344
1
1
Mahajanga
holotype
2275276474
MOLC
Madagascar
-17.75
Toamasina
1277
48.716667
343
344
1
1
Toamasina
holotype
2275276411
FSAE
Madagascar
-17.75
Chrome
1277
47.983334
343
344
1
1
Toamasina
holotype
2275276485
[262,889,1809,1835]
1992-02-01
FSAE
Madagascar
Morarano-Chrome
343
344
1
1
Toamasina
holotype
2275276750
[889,1282,1809,1835]
ISNB
Foulpointe
Madagascar
Toamasina
343
344
1
Toamasina
holotype
2275276290
FSAE
Ivondro
Madagascar
Toamasina
343
344
1
1
Toamasina
holotype
2275276327
[392,1388,1844,1872]
1957-06
MNHN
Pierre Andria Robinson
Madagascar
Andekaleka
Rogez
343
344
1
1
Toamasina
holotype
2275276296
2003-03-17
2003-03-21
2003-03-17
MNHN
Fisher & Griswold
Madagascar
600
-15.286333
Akirindro Mt.
13
49.54233
343
344
1
1
Toamasina
holotype
2275275954
2005-11-28
CAS
Brian L. Fisher
Madagascar
390
-17.921
Camp Rendrirendry
13
49.193165
Betampona Reserve
343
344
2
2
Toamasina
holotype
2275276294
2009-02-21
2009-02-23
2009-02-21
CAS
B. L. Fisher
Madagascar
780
344
345
-17.759167
Onibe River
21
48.85472
Zahamena National Park
343
344
1
1
Toamasina
holotype
2275275869
1994-11-13
1994-11-21
1994-11-13
CAS
B. L. Fisher
Madagascar
1280
-14.75
Anjanaharibe-sud Reserve
1287
49.466667
344
345
1
1
Toamasina
holotype
2275275974
DEUCD
Bekily
Madagascar
Toliara
344
345
1
1
Toliara
holotype
2275275959
[468,952,225,251]
MNHN, MRAC
Beroboka
Madagascar
Toliara
344
345
2
2
Toliara
holotype
2275276616
2008-08-03
2008-08-11
2008-08-03
NHMUK
Antsarongaza & M. Irwin & R. Harin'Hala
Madagascar
128
-19.9775
Beroboka village
12
44.665333
344
345
1
1
Toliara
holotype
2275275780
[531,896,297,323]
MOLC
Belalanda
Madagascar
Toliara
344
345
1
1
Toliara
holotype
2275276475
2002-02-21
2002-02-25
2002-02-21
FSAE
Fisher & Griswold
Madagascar
80
-24.652166
Mahafaly Plateau
12
43.991333
344
345
1
1
Toliara
holotype
2275275875
2009-07-03
2009-07-16
2009-07-03
MOLC
M. Irwin & R. Harin'Hala
Madagascar
130
-24.650333
Androimpano Forest
12
43.963165
344
345
1
1
Toliara
holotype
2275275915
[1083,1389,405,431]
CAS
Behara
Madagascar
Toliara
344
345
1
1
Toliara
holotype
2275276495
MRAC
Fort-Dauphin
Madagascar
Toliara
344
345
1
1
Toliara
holotype
2275276676
2003-05-18
2003-05-26
2003-05-18
MRAC
M. Irwin & F. Parker & R. Harin'Hala
Madagascar
37
-25.590557
Marie Special Reserve
Cap
344
345
1
1
Toliara
holotype
2275276015
[232,776,512,539]
2003-02-25
2003-03-07
2003-02-25
CAS
M. Irwin & F. Parker & R. Harin'Hala
Madagascar
37
-25.590557
Marie Special Reserve
Cap
344
345
1
1
Toliara
holotype
2275275945
2008-09-10
2008-09-17
2008-09-10
CAS, MOLC
B. L. Fisher
Madagascar
30
-20.069168
Foret de Kirindy
21
44.660557
344
345
3
3
Toliara
holotype
2275276303
2002-09-23
2002-10-03
2002-09-23
CAS
R. Harin'Hala
Madagascar
30
-22.903667
Mikea Forest
12
43.4755
344
345
2
2
Toliara
holotype
2275276169
2003-07-06
2003-07-17
2003-07-06
CAS
R. Harin'Hala
Madagascar
30
-22.903667
Mikea Forest
12
43.4755
344
345
1
1
Toliara
holotype
2275276461
2003-10-30
2003-11-09
2003-10-30
CAS
M. Irwin & F. Parker & R. Harin'Hala
Madagascar
180
-24.936832
Tsimelahy
12
46.626667
Andohahela National Park
344
345
1
1
Toliara
holotype
2275276775
[195,867,728,755]
1951-02-20
2004-01-15
1951-02-20
CAS
M. Irwin & F. Parker & R. Harin'Hala
Madagascar
180
-24.936832
Tsimelahy
12
46.626667
Andohahela National Park
344
345
1
1
Toliara
holotype
2275275664
2003-03-23
2003-04-02
2003-03-23
CAS
R. Harin'Hala
Madagascar
165
-23.6865
Beza Mahafaly Reserve
12
44.591
344
345
1
1
Toliara
holotype
2275275916
[531,1237,801,827]
2002-02-01
2002-02-08
2002-02-01
MOLC
R. Harin'Hala
Madagascar
165
-23.6865
Beza Mahafaly Reserve
12
44.591
344
345
1
1
Toliara
holotype
2275276020
2002-02-08
2002-02-15
2002-02-08
CAS
R. Harin'Hala
Madagascar
165
-23.6865
Beza Mahafaly Reserve
12
44.591
344
345
1
1
Toliara
holotype
2275275623
2002-03-18
2002-03-22
2002-03-18
CAS
Fisher & Griswold
Madagascar
Tsimanampetsotsa National Park
40
-24.041666
Beheloka
12
43.751835
Mitoho Cave
344
345
1
1
Toliara
holotype
2275275805
1922-02-19
2003-03-24
1922-02-19
CAS
M. Irwin & F. Parker & Harin'Hala
Madagascar
85
-25.006666
Berenty Special Reserve
12
46.303333
344
345
1
1
Toliara
holotype
2275276392
[231,904,980,1007]
1922-02-15
2003-02-15
1922-02-15
CAS
M. Irwin & F. Parker & Harin'Hala
Madagascar
85
-25.006666
Berenty Special Reserve
12
46.303333
344
345
1
1
Toliara
holotype
2275276946
1922-02-16
2003-02-25
1922-02-16
CAS
M. Irwin & F. Parker & Harin'Hala
Madagascar
85
-25.006666
Berenty Special Reserve
12
46.303333
344
345
1
1
Toliara
holotype
2275276346
[325,1051,1017,1043]
2002-10-22
2002-12-27
2002-10-22
CAS
M. Irwin & F. Parker & Harin'Hala
Madagascar
85
-25.006666
Berenty Special Reserve
12
46.303333
344
345
1
1
Toliara
holotype
2275275570
[1065,1387,1017,1043]
CAS
Berenty
Madagascar
Toliara
344
345
1
1
Toliara
holotype
2275276776
1983-05-05
1983-05-15
1983-05-05
NHMUK
J. S. Noyes & M. C. Day
Madagascar
Berenty
344
345
1
1
Toliara
holotype
2275275930
2001-11-09
2001-11-19
2001-11-09
NHMUK
M. E. Irwin & F. D. Parker & R. Harin'Hala
Madagascar
840
-22.8865
Zombitse National Park
12
44.692165
344
345
2
2
Toliara
holotype
2275275633
2009-02-09
2009-02-10
2009-02-09
CAS
B. L. Fisher
Madagascar
1270
-23.463612
Ampanihy
21
46.463055
Kalambatritra Special Reserve
344
345
1
1
Toliara
holotype
2275276642
2003-10-27
2003-11-02
2003-10-27
CAS
Frontier Wilderness Project
Madagascar
20
-23.046383
Ranobe
1
43.610283
344
345
1
1
Toliara
holotype
2275275444
2003-11-08
2003-11-15
2003-11-08
CAS
Frontier Wilderness Project
Madagascar
20
-23.03945
Ranobe
1
43.610283
344
345
1
1
Toliara
holotype
2275276845
2003-08-05
2003-08-10
2003-08-05
CAS
Frontier Wilderness Project
Madagascar
65
-23.222517
Fiherenana
1
43.880882
344
345
1
1
Toliara
holotype
2275276137
2002-10-31
2002-11-06
2002-10-31
CAS
Frontier Wilderness Project
Madagascar
100
-23.172832
Fiherenana
12
43.9565
344
345
1
1
Toliara
holotype
2275276679
[605,1284,1377,1403]
2002-10-22
2002-10-31
2002-10-22
CAS
Frontier Wilderness Project
Madagascar
100
-23.172832
Fiherenana
12
43.9565
344
345
4
4
Toliara
holotype
2275276382
2002-05-15
2002-05-29
2002-05-15
CAS, MOLC
Frontier Wilderness Project
Madagascar
70
-23.523167
Manderano
12
44.086334
344
345
2
2
Toliara
holotype
2275276459
2010-11-24
2010-12-12
2010-11-24
CAS
B. L. Fisher
Madagascar
500
-21.2185
Makay Mts.
12
45.321
344
345
2
2
Toliara
holotype
2275276386
2010-12-26
2011-01-03
2010-12-26
CAS
M. Irwin & R. Harin'Hala
Madagascar
600
-22.666666
National Park
1259
44.833332
344
345
1
1
Toliara
holotype
2275276151
1990-03-08
CAS, SANC
C. D. Eardley
South Africa
-22.183332
Altyddroog Farm
1261
29.883333
344
345
1
1
Limpopo
holotype
2275275965
1998-01-24
1998-02-21
1998-01-24
S. van Noort & B. Fisher
South Africa
-34.456833
Walker Bay Nature Reserve
11
19.3565
344
345
1
Western Cape
holotype
2275276437
2009-09-10
2009-11-04
2009-09-10
SAMC
Rooiberg Sandstone Fynbos & S. van Noort
South Africa
940
-33.72772
Gamkaberg Nature Reserve
1
21.96
344
345
1
1
Western Cape
holotype
2275276479
[356,737,1701,1727]
SAMC
Uganda
Uganda
UGANDA
344
345
1
1
holotype
2275275209
2008-08-04
2008-08-26
2008-08-04
OLL
S. van Noort
Uganda
National Park
1523
0.5639333
Makerere
1
30.361668
Kanyawara
344
345
1
1
Kibale
holotype
2275276061
2002-08-21
2002-10-28
2002-08-21
SAMC
A. van Harten
Yemen
Manakhah
344
345
2
2
holotype
2275277056
[474,619,1809,1835]
MOLC
Yemen
344
345
1
1
holotype