Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): description of 10 new species from the L. australis, L. monteithi, and L. villosus species groups Gunter, Nicole L. Weir, Thomas A. Zootaxa 2019 2019-11-08 4695 3 201 252 3T8V4 in Matthews 1974 [366,594,1798,1825] Insecta Scarabaeidae Lepanus Animalia Coleoptera 21 222 Arthropoda species monteithi    1. Clypeal edge with a blunt projection lateral to clypeal teeth ( Fig. 8A); dorsal part of eyes wide, separated by 7 eye widths; medial lobe of metaventrite narrowly margined beside mesocoxae ( Fig. 8D); male metatibiae without a notched inner edge; aedeagus as in Fig. 10D, with a small tuft of setae on apex of right paramere; northern Northern Territory...............................................................................   Lepanus topendGunter & Weir, new species  - Clypeal edge evenly rounded lateral to clypeal teeth ( Fig. 8B); dorsal parts of eyes narrower, separated by more than 12 eye widths; medial lobe of metaventrite with margin beside mesocoxae either very narrowly margined or virtually absent ( Figs. 8H–I); male metatibiae with or without a notch on inner edge; aedeagus with or without small tufts of setae on apices of parameres............................................................................................... 2   2. Lateral lobe of metaventrite with a pit containing a tuft of setae ( Fig. 8G), virtually impunctate anteriorly, inner border beside mesocoxae not defined; head and disc of pronotum very finely punctate; aedeagus as in Fig. 10A, without tufts of setae on parameres; northern Queensland, Iron Range area north to tip of Cape YorkPeninsula................................................................................................   Lepanus geoffiGunter & Weir, new species  - Lateral lobe of metaventrite without a setose pit, punctate anteriorly, inner border beside mesocoxae well defined; head and disc of pronotum with punctation fine or not; aedeagus with or without tufts of setae on apices of parameres................. 3   3. Outer edge of protibiae with only 2 distinct teeth, 3 rdor basal tooth reduced to little more than a denticle ( Fig. 8E); male metatibiae with a notch on inner edge (eg Fig. 8F); aedeagus with small tufts of setae on apices of both right and left parameres... ................................................................................................... 4  - Outer edge of protibiae with 3 distinct teeth; male metatibiae with or without a notch on inner edge; aedeagus with or without small tufts of setae on apices of parameres................................................................. 5   4. Head without a basal carina; head and disc of pronotum very finely punctate; elytral intervals virtually impunctate; lateral lobe of metaventrite with inner border beside mesocoxae anteriorly curving to meet lateral border ( Fig. 8H); pygidium not reticulate basally and virtually impunctate ( Fig. 11C); aedeagus as in Fig. 10C, with small tufts of setae on apices of both right and left parameres, left paramere pointed; northern Queensland, Wet Tropics......   Lepanus reidiGunter & Weir, new species[part]  - Head with short basal carina present only adjacent to eyes ( Fig. 8C); head and disc of pronotum with punctation not so fine; elytral intervals distinctly punctate; lateral lobe of metaventrite with inner border bedside mesocoxae anteriorly straight, lateral border virtually absent ( Fig. 8I); pygidium often finely reticulate basally and with distinct punctures ( Fig. 11B); aedeagus as in Fig. 10B, with small tufts of setae on apices of both right and left parameres, left paramere somewhat spatulate; northern Queensland, Iron Range area................................................   Lepanus monteithi Matthews, 1974   5. Basal margin of pronotum without a row of small punctures; head and disc of pronotum very finely punctate; elytral striae obsolete; lateral lobe of metaventrite with inner border beside mesocoxae anteriorly curving forward to meet lateral border ( Fig. 8H); pygidium not reticulate basally and virtually impunctate ( Fig. 11C); male metatibiae with a notch on the inner edge ( Fig. 8F); aedeagus as in Fig. 10C, with small tufts of setae on the apices of both right and left parameres; northern Queensland, Wet Tropics.....................................................   Lepanus reidiGunter & Weir, new species[part]  - Basal margin of pronotum with a distinct row of small punctures; head and disc of pronotum with punctation not so fine, easily discernable; elytral striae clearly visible; lateral lobe of metaventrite with inner border beside mesocoxae anteriorly straight, lateral border virtually absent (as in Fig. 8I); pygidium finely reticulate basally and with distinct punctures ( Fig. 11E); male metatibiae without a notch on inner edge; aedeagus as in Fig. 10E, without small tufts of setae on apices of parameres; northern Queensland, Iron Range area north to Torres Strait.....................   Lepanus yorkensisGunter & Weir, new species