A revision of the genus Rudolfina Roháček (Sphaeroceridae: Limosininae) Paiero, Steven Mark Marshall, Stephen A. European Journal of Taxonomy 2020 2020-01-16 593 1 48 Paiero & Marshall, 2020 Paiero & Marshall 2020 [602,888,1573,1600] Insecta Sphaeroceridae Rudolfina Animalia Diptera 29 30 Arthropoda species megepandria sp. nov.   Figs 4, 13, 21B    Etymology The specific name, which has been a manuscript name since Marshall (1982), refers to the unusually large epandrium.    Material examined   Holotype   MEXICO • ♂; Jalisco, 18 mi.W of Atenquique;  9300 fta.s.l.; A. Newtonleg.; fir forest, dung; FMNHdebu01086084.   Fig. 13.  Rudolfina megepandria sp. nov., male terminalia (debu01086085). A. Epandrium, surstylus and cercus, lateral view. B. Same as preceding, caudal view. C. Sternites 4–7, ventral view. D. Phallus and postgonite, dorsal view. E. Phallus, postgonite and phallapodeme, dorsolateral view. F. Same as preceding, lateral view. Scale bars: A–B = 0.20 mm; C = 0.05 mm; D–F = 0.05 mm.   Paratypes   MEXICO–  Jalisco• 1 ♂, 1 ♀; same collection data as for holotype; DEBU. –   Oaxaca• 1 ♀; Yuvila Rd;  9400 fta.s.l.;  9–19 Aug. 1973; A. Newtonleg.; mesic oak, carrion; FMNH.    Description BODY. Length: 1.9–2.2 mm. Eye height 2.5 × genal height. Head with 3 interfrontal setae and row of 4–5 small setulae on inner margin of orbital plate. Gena with 2 strong setae and 7–8 smaller setulae. Acrostichal setulae in 4–6 rows. Costagial seta extending to subcostal break. Second costal sector 0.75– 0.8× third costal sector. Length of M 1between crossveins dm-cu and r-m– 3.0× dm-cu; CuA 1stub vein ~10.0 × M 1stub vein (M 1stub vein extremely reduced). Male mid femur with 14–15 strong setae in ventrobasal cluster. Male mid tibia with ventral comb of 4–5 robust, short setae on apical ½. Mid tibia without midventral seta (both sexes). MALE ABDOMEN ( Fig. 13). Sternite 5 posterior margin with 2 small nipple-like teeth on each side of small emargination; emargination ~1/15 width and ~1/9 length of sternite 5. Transverse part of sternite 6 arcuate. Epandrium swollen, broader than preceding abdominal segments; subanal plate broad, complete. Surstylus with posterior lobe elongate, length ~4.0× basal width, with a hirsute medial lateral lobe; apex truncate. Cercus elongate-conical, projecting posteriorly. Postgonite elongate, narrowly rounded distally and narrow basally. Dorsal sclerite of distiphallus ( Fig. 13D–F) without dorsal swellings and not extending beyond apex of acrophallus. FEMALE ABDOMEN ( Fig. 4). Tergite 7 with posterior margin entire. Medial part of tergite 8 elongate, length ~2.0–3.0 × width, with posterior margin emarginate. Epiproct broadly rounded anteriorly. Cercus ~0.75× length of epiproct; fused anterolaterally to epiproct. Sternite 7 with posterior margin broadly rounded. Spermathecae cup-shaped; paired spermathecae with stems ~1.5× width of spermatheca.    Remarks   Rudolfina megepandria sp. nov.is diagnosed by its swollen male epandrium and distinctively shaped sternite 5. It is most closely related to  R. bucki sp. nov., which has a less pronounced epandrium and differs in details of the surstylus and cercus. 2564277645 FMNH A. Newton Mexico 2835 Atenquique 29 30 1 1 Jalisco holotype 2564277647 [189,1069,302,329] DEBU A. Newton Mexico 2835 Atenquique 31 32 2 1 1 Jalisco paratype 2564277671 1973-08-09 1973-08-19 1973-08-09 FMNH A. Newton Mexico 2865 Yuvila Rd 31 32 1 1 Oaxaca paratype