Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations Williams, Kevin A. Cambra, Roberto A. Bartholomay, Pedro R. Luz, David R. Quintero, Diomedes Pitts, James P. Zootaxa 2019 2019-03-05 4564 1 101 136 Williams Williams & Cambra & Bartholomay & Luz & Quintero & Pitts 2019 [151,528,297,323] Insecta Mutillidae Lomachaeta GBIF Animalia Hymenoptera 11 112 Arthropoda species eironeia sp. nov.  ( Figs 59, 69, 70)   Diagnosis.MALE. This species can be immediately recognized by the unique genitalia, wherein the cuspis and lateral paramere margins have numerous elongate (~5 × paramere width) setae. The following characters are also useful for diagnosis: the body is entirely blackish; the head and mesosoma have coarse separated punctures; the mandible is unarmed ventrally; the forewing has its veins encompassing the basal 0.7 × of the wing; the T1 shape is sub-disciform; the T2 disc is densely coarsely punctate; the T2 fringe is composed of simple setae.   Description.MALE. Body length 4 mm. Coloration( Fig. 40). Entire body black, except mandible, tegula, and legs largely brown. Tibial spurs white. Wings hyaline, veins brown. Body setae whitish, except frons, vertex, and mesoscutum with interspersed erect silvery and brownish brachyplumose setae. Head.Rounded posteriorly, with coarse punctures, coarser on face. Mandible tridentate apically, unarmed ventrally. Gena not carinate. Ocelli small; ocellocular distance 6 × length of lateral ocellus, interocellar distance 2 × lateral ocellar diameter. F1 1.0 × pedicel length; F2 1.6 × pedicel length. Mesosoma.Pronotum and mesoscutum with coarse separated punctures. Tegula smooth, margins setigerously punctate. Mesopleuron with coarse contiguous punctures. Metapleuron glabrous. Scutellum slightly convex, with coarse confluent punctures. Axilla sessile with mesoscutum, with smooth curved sharp posterior tooth. Propodeum areolate dorsally, lateral face mostly smooth with faint areolations posteriorly. Forewing with veins encompassing basal 0.7 × wing. Metasoma.T1 shape sub-disciform, with defined anterior and dorsal faces. T2 with coarse dense punctures, intervals smooth; T2–4 fringe setae slightly thicker than disc setae, but not forming bristles. S2 with coarse separated punctures. T3–6 punctate. T7 smooth. Hypopygium punctate, sharply bidentate posteriorly. Genitalia( Figs 69, 70). Paramere dorsoventrally flattened, obliquely downcurving to apex, sub-truncate apically, with long (~5 × paramere width) ventral setae around inner and lateral margins, setae longer and denser apically. Cuspis with posteroventral setae ~5 × cuspis length. Penis valve unidentate apically. FEMALE. Unknown.   Material examined.   Holotype, ♂, MEXICO,  Sonora, km 23, Hwy16, 5.5 kmENE [Los] Hornos,  23– 26.VII.2008, 27 o43.71’N 109 o50.64’W,  40m, M.E. Irwin& R.L. Wescott, malaise in dense riparian forest (EMUS, Figs 59, 69, 70).   Distribution.Known only from the typelocality in Sonora, Mexico   Etymology.From the Greek,  eironeia, irony, in reference to the fact that this specimen was recognized as a new species by KAW only a few months after Williams & Pitts 2009was published.   Remarks. The genitalia of this species are truly bizarre ( Fig. 59), even for a genus with such variable parameres ( Williams & Pitts 2009). 2012757423 2008-07-23 2008-07-26 2008-07-23 M. E. Irwin & R. L. Wescott Mexico 40 27.7285 Hornos 12 -109.844 Hwy 11 112 1 1 Sonora holotype