Nearctic Anthomyzidae: a monograph of Anthomyza and allied genera (Diptera) Roháćek, Jindřich Barber, Kevin N. Acta Entomologica Musei Nationalis Pragae 2016 suppl. 2016-12-30 56 1 412 Roháćek & Barber, 2016 Roháćek & Barber 2016 [429,681,1307,1332] Insecta Anthomyzidae Anthomyza Animalia Diptera 299 296 Arthropoda species gilviventris sp. nov.  ( Figs 482–484, 486–507, 510–515, 517)   Type material.  HOLOTYPE: ♂, “ CAN: BC: Fernie, Annex Pk,  17.vii.2011, KNBar-ber, sweeps, wet ditch,  Carex utriculata 49°30.72’N 115°04.13’W” and “Holotypus ♂  Anthomyza gilviventris sp. n., J. Roháček& K. N. Barberdet. 2014” (red). The specimen is in good condition, with well-exposed gonostyli (see Fig. 483) ( CNCI, intact).   CANADA: ALBERTA: Banff N. P.,   15 miE Mt. Eisenhower Jct.,  27.vii.1967, 2 ♀♀, S. P. Whitneyleg. ( USNM) ; 7.0 mi W Banff,  11.vii.1966, 1 ♀, K. Valleyleg. (all USNM); ~   22.7 kmS Bellevue, Hwy 774, 49°22.62'N 114°22.58'W, sweeps, roadside ditch, mostly   Carexspp.,   Equisetum, grasses,  17.vii.2011, 3 ♂♂ 5 ♀♀; ~   18.3 kmNW Cadomin, Hwy#40, 53°10.02'N 117°29.72'W, sweeps, edge of roadside pond,   Carexspp.,  22.vii.2008, 16 ♂♂ 10 ♀♀, all K. N. Barberleg. (all CNCI);  Elkwater Lk.,  10.vi.1956, 3 ♂♂ 4 ♀♀, swept from sedge,  9.vii.1956, 1 ♀, O. Peckleg.; ~  0.9 kmWNW Exshaw, 51°03.78'N 115°10.99'W, sweeps, mostly emergent   Carexsp.,  25.vii.2008, 1 ♀, K. N. Barberleg. (all CNCI);  Calgary, Fish Creek Prov.Pk., pond near Shannon Terrace, 50°55.60'N 114°07.427'W, swept from sedgesand   Equisetum,  12.viii.2011, 2 ♂♂, J. E. Swannleg. ( BDUC);  Calgary, Fish Creek Prov. Pk., Shannon Terrace, sweep around pond by 2 ndbridge,  9.vii.2010, 1 ♀, J. E. Swannleg. ( BDUC);  Fish Creek P. Pk., 50°55.61'N 114°07.43'W, sweeps, mostly   Carex utriculataand   Equisetum ssuviatile,  12.vii.2011, 3 ♂♂ 3 ♀♀;  Fish Creek P. Pk., 50°55.74'N 114°07.32'W, sweeps, mostly   Carex utriculataand   Equisetumsp.,  12.vii.2011, 4 ♂♂ 4 ♀♀, all J. E. Swann& K.N. Barberleg. (all BDUC); ~  22.5 kmNW Highwood House, ~  4 kmW Mist Ck, 50°31.40'N 114°53.12'W, sweeps,   Carexsp. (small),  25.vii.2008, 1 ♂ 1 ♀, K. N. Barberleg. ( DEBU); ~  3.4 kmSSW Hinton, Hwy.#40, 53°21.27'N 117°37.32'W, sweeps,   Equisetum ssuviatile,  22.vii.2008, 3 ♂♂; ~  4.4 kmSSW Hinton,Hwy#40, 53°20.77'N 117°36.83'W, sweeps,   Equisetum ssuviatile,   Carexsp. [   C. aquatilisWahlenb.],  22.vii.2008,  1 ♀; ~  15 kmSSW Hinton, gas right-of-way nr. Wildhorse LakeP[rovincial] R[ecreation] A[rea], 53°16.65'N 117°44.57'W, sweeps, roadside   Equisetum palustre,  23.vii.2008, 4 ♂♂;  Kananaskis Country, Sibbald Area, Hwy.68,  3.1 kmW Powderface Trail, 51°03.10'N 114°54.72'W, sweeps, mostly   Carex utriculata,  15.vii.2011, 7 ♂♂ 6 ♀♀( 2 ♂♂genit. prep.), all K. N. Barberleg. (all CNCI); ~  4.4 kmNNE Kananaskis Village, Mt. Lorette Ponds, 50°58.11'N 115°06.59'W, sweeps, mostly   Equisetum ssuviatile,  25.vii.2008, 1 ♂, K. N. Barberleg. ( DEBU); ~  14.4 kmE Obed, Range Rd.213 @ RR crossing, 53°32.19'N 117°01.02'W, sweeps, mostly   Carex utriculata,  25.vii.2011, 4 ♂♂ 3 ♀♀, K. N. Barberleg. ( CNCI);  Peter Lougheed P. Pk., 50°41.82'N 115°06.92'W, sweeps, fen,   Carex utriculatawith cf.  Poa pratensis,  13.vii.2011, 98 ♂♂ 30 ♀♀( DEBU 01502929–3056, 2 ♂♂ 1 ♀genit. prep.),  14.vii.2011, 2 ♂♂ 2 ♀♀( DEBU 01503292–95),  15.vii.2011, 1 ♂ 1 ♀( DEBU 01503333, -34),  16.vii.2011, 2 ♂♂ 3 ♀♀( DEBU 01503390–94); Peter Lougheed P.Pk.,  Upper Lake Drive, 50°36.98'N 115°07.13'W, sweeps, edge of fen,   Carex utriculata,  14.vii.2011, 1 ♀( DEBU 01503314); same locality but 50°36.94'N 115°07.16'W, sweeps, edge of fen,   Carex utriculata,  16.vii.2011, 3 ♂♂ 2 ♀♀( DEBU 01503647–51), all K. N. Barber; ~  20 kmSSE Robb, Hwy#40, Lovett RiverP[rovincial] R[ecreation] A[rea], 53°03.89'N 116°49.02'W, sweeps, streamside, mostly grasses,  23.vii.2008, 1 ♂, K. N. Barberleg. ( DEBU);  Spray Valley P.Pk., 50°48.95'N 115°09.84'W, sweeps, fen,   Carex utriculata? and  Poasp.,  13.vii.2011, 39 ♂♂ 40 ♀♀( DEBU 01502773–851);  W. A. Switzer P. Pk.,  BeaverRanch Trail, 53°29.80'N 117°48.02'W, sweeps, edge of Jarvis Ck.,   Carex utriculata,  22.vii.2011, 13 ♂♂ 2 ♀♀( DEBU 01503707–21); same locality but 53°29.67'N 117°48.00'W, sweeps, sedge-nlled oxbow,   Carex utriculata?,  22.vii.2011, 1 ♂( DEBU 01503777);  W. A. Switzer P. Pk., off Hay River Rd. W, 53°33.44'N 117°48.43'W, sweeps, mostly  Carex utriculata,  22.vii.2011, 8 ♂♂ 2 ♀♀( DEBU 01503792–801),  24.vii.2011, 22 ♂♂ 7 ♀♀( DEBU 01503847–75, 1 ♀genit. prep.), all K. N. Barberleg.;  Waterton Lakes Nat. Pk.,  14–20.vii.1980, 1 ♀, H. J. Teskeyleg. ( CNCI).  BRITISH COLUMBIA: Brisco,  19.vi.1932, 2 ♀♀, O. B[ryant]leg. ( USNM, 1 ♀missing one wing);  Buckinghorse Provincial Campground, Alaska Highway, DC-175 ( 281.6 km),  27.vi.1978, 2 ♀♀, P. H. Arnaud Jr.leg. ( CASC);  Fernie, Annex Pk., 49°30.72'N 115°04.13'W, sweeps, wet ditch, mostly  Equisetum arvense&  E. laevigatum,  17.vii.2011, 1 ♂ 1 ♀, sweeps, wet ditch,   Carex utriculata,  17.vii.2011, 13 ♂♂ 14 ♀♀,  18.vii.2011, 13 ♂♂ 16 ♀♀,  19.vii.2011, 16 ♂♂ 8 ♀♀;  Fernie, Annex Pk., 49°30.66'N 115°04.16'W, sweeps, pond margin,   Carex utriculata,  18.vii.2011, 14 ♂♂ 4 ♀♀( 1 ♂genit. prep.), all K. N. Barberleg. (all CNCI);   4 kmNW Field, CD 1416, 31.v.1992, 1 ♂, A. Borkentleg.;  ~ 3.5 kmNNW Fort Steele, Hwy 95 rest area, 49°38.90'N 115°38.82'W, sweeps, graminoids, mostly   Carex pellita,  18.vii.2011, 1 ♀, K. N. Barberleg. (both CNCI);  Graham Is., Slatechuck Mt., roadside sweep,  12.vii.1988, 1 ♀, T.A. Wheelerleg. ( DEBU, genit. prep.);  Kaslo Cr.,  18.vi.[-], 1 ♀, R. P. Currieleg. ( USNM);  Kinbasket Lake, BC Hydrodrawdown study, Malaise trap,  13.vi.2008, ( WIL08-01), 1 ♀, ( WIL08-15), 1 ♂, Cooper Beauchesne &Assoc. Ltd.leg.( RBCM);  ~8.0 km SE Valemount, edge of Kinbasket Lake, 52°46.65'N 119°10.38'W, sweeps, mostly   Carex utriculata,  23.vii.2011, 18 ♂♂ 9 ♀♀, K. N. Barberleg.;   King SalmonL., 58°43'N 132°54'W, 1750',  Carex, grass,  Equisetumbeside lake,  17.vii.1960, 5 ♂♂ 4 ♀♀, W. W. Mossleg. (all CNCI);  Merritt,  1.5 kmNE Lundbom L.,  1.vii.1988, 1 ♀, G. E. Hutchingsleg. ( RBCM, ENT991-6166);  Moresby Camp, Q[ueen] C[harlotte] Islands,  29.vi.1957, 1 ♀, E. E. MacDougallleg ( CNCI).;  Mt. Robson Prov. Pk., Hwy #16, small road towards Mt. Robson, 53°03'N 119°15'W, forest noor, swamp, (Universität Bielefeld, Ca1519),  6.viii.2002, 5 ♂♂ 1 ♀, M. v. Tschirnhausleg. ( ZSMC, in ethanol);  Nanaimo Lakes, forested area,  11.vi.1988, 1 ♂ 1 ♀, G. E. Hutchingsleg. ( RBCM);  Parson, Crestbrook Rd., 51°03.68'N 116°39.06'W,sweeps,wet ditch,   Carex utriculatawith  Equisetum palustre& E.× litorale,  18.vii.2011, 2 ♂♂ 5 ♀♀, K. N. Barberleg. ( CNCI, 1 ♂ 1 ♀ genit. prep.);  Pete Lake, 57°56'N 131°56'W, 4000',  19.viii.1960, 1 ♂ 2 ♀♀, R. Pilfreyleg.; Revelstoke,  2.vii.1973, 1 ♀, H. J. Teskeyleg. (all CNCI);  Robson,  3.vi.1958, 1 ♀,  6.vi.1967, 1 ♀,  5.vii.1967, 1 ♀, H. R. Foxleeleg. ( UBCZ);  Qualicum Beach, Little Qualicum River Estuary Regional Conservation Area, ex.  Carex4, aspirator,  4.vi.1980, 1 ♀, Canadian Wildlife Serviceleg. ( RBCM, ENT991-17202);   SalmonArm, CD980,  8.vi.1988, 1 ♀, CD1210,  8.vi.1990, 1 ♀, A. Borkentleg.;  ~ 27 kmN Sparwood, Lower ElkValley Rd., 49°50.24'N 114°53.29'W, sweeps, edge of creek,   Carex utriculata?,  20.vii.2011, 1 ♂ 1 ♀, K. N. Barberleg.;  Summit Lake, mi.392 Alaska Hwy., 4500',  23–24.vi.1959, 1 ♀, E. E. MacDougallleg. (all CNCI);  Vancouver  50 kmE,  3.vii.1988, 1 ♂ 2 ♀♀, A. Freidbergleg. ( TAUI).   LABRADOR: Cartwright,  9.vii.1955, 1 ♂,  12.vii.1955, 3 ♀♀( 1 ♀genit. prep.), E. F. Cashmanleg,   17.vii.1955, 5 ♂♂ 6 ♀♀( 3 ♂♂ 3 ♀♀genit. prep.),  19.vii.1955, 1 ♂, E. E. Sternsleg. ( CNCI).   MANITOBA:  10 kmSE Churchill, Farnworth Lake (Landing L.), 58.7045°N 94.0525°W,  14 ma.s.l., tundra,  14.vii.2010, 1 ♀(Sample ID:10PROBE-14241, BOLD ID:JWDCI851-11), 1 ♂(Sample ID:10PROBE-14244, BOLD ID:JWDCI854-11), 1 ♀(Sample ID:10PROBE-14245, BOLD ID:JWDCI855-11),  23.vii.2010, 1 ♀(Sample ID:10PROBE-16182, BOLD ID:JWDCK227-11), 1 ♂(Sample ID:10PROBE-16183, BOLD ID:JWDCK228-11),  28.vii.2010, 1 ♂(Sample ID:10PROBE-15878, BOLD ID:JWDCJ1443-11), 1 ♂(Sample ID:10PROBE-15879, BOLD ID:JWDCJ1444-11),  31.vii.2010, 1 ♂(Sample ID:10PROBE-12448, BOLD ID:JWDCH242-10),  6.viii.2010, 1 ♀(Sample ID:10PROBE-13043, BOLD ID:JWDCH837-10), 1 ♀(Sample ID:10PROBE-13045, BOLD ID:JWDCH839-10);   26 kmSE Churchill, Twin Lakes, 58.6332°N 93.7871°W,  29 ma.s.l., fen,  1.viii.2010, 1 ♂(Sample ID:10PROBE-12672, BOLD ID:JWDCH466-10),  7.viii.2010, 1 ♀(Sample ID:10PROBE-13220, BOLD ID:JWDCH020-10);   12 kmESE Churchill, Launch Road, 58.7541°N 93.9974°W,  10 ma.s.l., fen,  14.vii.2010, 1 ♀(Sample ID:10PROBE-14512, BOLD ID:JWDCJ172-11),  19.vii.2010, 1 ♂(Sample ID:10PROBE-16713, BOLD ID:JWDCL663-11), 1 ♀(Sample ID:10PROBE-16714, BOLD ID:JWDCL664-11);   23 kmE Churchill, Ramsay Creek, 58.7306°N 93.7804°W,  10 ma.s.l., boreal forests,  23.vii.2010, 1 ♀(Sample ID:10PROBE-16481, BOLD ID:JWDCL241-11), all J. Wangleg. (all BIOUG, Barcode of Life, in ethanol);  ~ 14 kmSW Falcon Lake, jct. Hwy#1 & Rd. 86E, 49°38.24'N 95°29.89'W, sweeps, graminoids, mostly   Carexspp.,  29.vii.2011, 1 ♀, K. N. Barberleg. ( CNCI);   5 kmN Gardenton, Tallgrass Prairie Reserve, 49°10.71'N 96°40.76'W, sweep in tallgrass prairie,  17.vi.1999, 1 ♀, V. Creccoleg. ( LEMQ 0039607);  Int. Peace Gardens, Turtle Mtn. For. Res.,  17.vii.1958, 1 ♀;  5 miN Minnedosa,  8.vii.1958, 1 ♀, both R. L. Hurleyleg. (both CNCI).   NEWFOUNDLAND:  Bayd’Espoir, low veg. nr. river,  11. vii.1985, 1 ♂, L. Hollettleg.; Coal Brook,  24.vi.1983, 1 ♀(genit. prep.), A. Borkentleg. (both CNCI);  Avalon Pen[insula], Portugal Cove, Indian Meal Line,  15.vii.1982, 1 ♂ 3 ♀♀,  15.vii.1982, b. l., 1 ♀, [no collector] (all genit. prep.); N branch St. George’s R.,  17.vi.1979, 1 ♀(genit. prep.), Larson & Swalesleg. (all NFRC);  St. John’s, Agric. Exp. Stn.,  26.vii.1967, 3 ♂♂ 4 ♀♀, J. F. McAlpineleg. ( CNCI, 1 ♀genit. prep.).   NORTHWEST TERRITORIES: Norman Wells,  23.vi.1969, 1 ♀, G. E. Shewellleg. ( CNCI).   NOVA SCOTIA: Paquette Lk.,  5 kmS South Harbour,  5.vii.1983, 1 ♀, A. Borkentleg. ( CNCI, genit. prep.).   ONTARIO: Algonquin P. Pk., nr. Sitting Duck Lake, 45.4522°N 78.4713°W, Malaise trap, hardwood forest (SD-MAT),  5–21.v.2008, 1 ♀, E. Proctorleg. ( DEBU);   10 kmNW Penetanguishene, Awenda P. Pk., Second Lake, sweep in kettle marsh,  3.vii.2001, 1 ♀, M. Pollettleg. ( LEMQ 0040381);  Bruce Peninsula N. P.,  CraneRiver below Lake Scugog, 45°07.0'N 81°32.1'W, sweeps, riverside vegetation,  3.vii.1999, 2 ♀♀( 1 ♀genit. prep.);  Bruce Peninsula N. P., Dorcas BayRd. at WillowCreek, 45°09.39'N 81°34.43'W, sweeps,   Carex aquatilis,  22.vi.2008, 1 ♂;  Bruce Peninsula N. P., Singing Sands, 45°11.50'N 81°34.61'W, sweeps, sedgesin fen,  22.vi.2008, 5 ♀♀(4 ♀♀ genit.prep), all K.N. Barberleg.(all DEBU);  ~ 13.9 kmW Chapleau, 47°49.20'N 83°35.42'W, hydro right-of-way, sweeps, mostly   Carex utriculata, grasses,  13.vi.2013, 1 ♀,  23.vi.2013, 1 ♀(genit. prep.);  ~ 40 kmNE Chapleau, 47°59.76'N 82°55.04'W, wet roadside sweeps, mostly   Carex utriculata,  23.vi.2013, 2 ♂♂ 1 ♀;  ~13.5 km S Cochrane, 48°56.65'N 81°00.18'W, hydro right-of-way, sweeps, mostly   Carex utriculata,  9.vii.2012, 3 ♂♂ 1 ♀,  22.vi.2013, 7 ♂♂ 3 ♀♀,  12.vii.2013, 5 ♂♂ 1 ♀( 1 ♂genit. prep.),  13.vii.2013, 1 ♂ 1 ♀,  18.viii.2013, 1 ♂ 2 ♀♀, all K. N. Barberleg. (all CNCI);  Grey Co., Dornoch, Anderson Lake,  1.vii.1992, 1 ♀, R. A. Canningsleg. ( RBCM);  Dryden, 49°47.27'N 92°48.62'W, sweeps, mixed graminoids/herbs,  17.vii.2008, 1 ♀;  Dubreuilville, along MagpieRiver, 48°21.12'N 84°34.04'W, sweeps,  Equisetum ssuviatile,  Carex,  10.vii.2010, 1 ♀, both K. N. Barberleg. (both CNCI); same locality but sweeping  Equisetum ssuviatile,  Carexspp. on muddy river bank,  10.vii.2010, 2 ♂♂ 2 ♀♀, J. Roháčekleg. ( SMOC, 2 ♂♂ 1 ♀ genit. prep.);  ~ 3.8 kmENE Dugwal, 48°35.33'N 80°57.90'W, sweeps, wet ditch,  Carex utriculata,  Equisetum ssuviatile,  Scirpus,  23.vi.2013, 1 ♂ 2 ♀♀;  Echo Bay, Echo BayMarsh, 46°29.66'N 84°04.12'W, sweeps, near lookout, mostly   Carex utriculata,  8.vi.2013, 1 ♂ 1 ♀,  15.vi.2013, 2 ♂♂ 2 ♀♀; same locality but 46°29.71'N 84°04.04'W, sweeps, nr. lookout, mostly  Carexincl.  C. utriculata,  24.vi.2012, 1 ♂ 1 ♀(pair in copula);  Elliot Lake, 46°22.23'N 82°36.49'W, sweeps, mixed graminoids incl.   Carex utriculata,  29.vi.2013, 1 ♂;  ~ 14.5 kmSSE Elliot Lake, ~ 3.7 kmNNE jct Hwys 17&108, 46°14.71'N 82°33.53'W, sweeps, mostly   Carex utriculata,  29.vi.2013, 1 ♂ 1 ♀, all K. N. Barberleg. (all CNCI);  ~ 59 kmNNW Elliot Lake, s. of Rocky Is. Lake, 46°50.16'N 83°03.05'W, sweeps, mostly   Carex aquatilisin fen,  3.vii.2010, 1 ♀, K. N. Barberleg. ( CNCI),   455 m, sweeping, mostly   Carex rostrata?[more likely  C. utriculata] in fen,  3.vii.2010, 5 ♂♂ 10 ♀♀, J. Roháčekleg. ( SMOC, 1 ♂ 2 ♀♀ genit. prep.);  Finland, S of Caliper Lake, sweep sedgeat beaverdam,  10.vii.1992, 1 ♂, T.A.Wheelerleg.( LEMQ 0039596,genit. prep.);  ~7.0 km E Foleyet, 48°14.34'N 82°20.75'W, hydro right-of-way, sweeps, mostly   Carex utriculata,  23.vi.2013, 2 ♂♂,  13.vii.2013, 2 ♀♀;  Goulais River, end of Island Rd., 46°43.57'N 84°24.45'W, sweeps, mud nats,   Equisetum ssuviatile,   Dulichium arundinaceum,  9.vii.2007, 1 ♀(genit. prep.); same locality but 46°43.33'N 84°24.72'W, sweeps, oxbow nats,   Equisetum ssuviatile, sedges, herbs,  16.vii.2007, 1 ♀(genit. prep.), all K. N. Barberleg. (all CNCI);  Goulais River, Sand Bay, 46°44.81'N 84°32.68'W, sweeping  Juncusand  Carexat margin of fen pools,  10.vii.2010, 5 ♀♀, J. Roháčekleg. ( SMOC, 1 ♀ genit. prep.);  Greenwater P. Pk., Sandbar Lk. Trail, 49°13.10'N 81°17.35'W, sweeps, lakeshore   Equisetumspp., graminoids,  Caltha,  21.vii.2009, 3 ♀♀, K.N.Barberleg.( DEBU 01502099–101);  ~2.0 km W Hallebourg, 49°40.36'N 83°32.11'W, sweeps, wet ditch, mostly   Carex utriculata,  22.vi.2013, 1 ♂;  ~ 43 kmW Hearst, hydro right-of-way opposite Forde Lk., 49°44.07'N 84°14.74'W, sweeps, mostly   Carex utriculata,  21.vi.2013, 2 ♀♀;  Icewater Creek WS [Watershed],  12.7 kmNNE Searchmont, mi.10.5 Whitman Dam Rd., alderthicket,  21.vi.1986, 1 ♂, all K. N. Barberleg. (all CNCI);  Lake Superior P. Pk., Hwy 17 near jct. Agawa Rock, 47°22.31'N 84°41.23'W, sweeps, mostly   Carex utriculata,  12.vii.2014, 2 ♂♂( DEBU 01503948, -49);  Lake Superior P.Pk., CrescentLk.campground, 47°16.66'N 84°33.00'W, sweeps, shoreline sedges,  29.v.2010, 2 ♂♂ 2 ♀♀( DEBU 01502431–34), all K. N. Barberleg.;  Manitoulin Island,  10 kmW Gore Bay, 45°53'N 82°34'W, sweep in grassland alvar,  20.vi.1996, 2 ♀♀, P. Bouchardleg. ( LEMQ 0039584, -85, 1 ♀ genit. prep.);  Manitoulin I., Misery BayProv. Nat. Res., 45°47'48"N 82°44'58"W, alvar,  27.v.2010, 1 ♀, S. A. Marshallleg. ( DEBU 00324518, genit. prep.); Hwy#17, ~ 8.5 kmNW Marathon, 48°47.69'N 86°26.11'W, sweeps, emergent   Equisetum ssuviatilewith  Carexsp.,  16.vi.2007, 3 ♂♂ 1 ♀, sweeps, emergent  Carexsp.,  16.vi.2007, 2 ♂♂ 1 ♀, sweeps, emergent  Equisetum ssuviatile,  16.vi.2007, 1 ♀(genit. prep.); same locality but 48°47.69'N 86°26.07'W, sweeps, roadside graminoids,  31.vii.2008, 1 ♀, all K. N. Barberleg. (all CNCI); Hwy#17, ~ 8.5 kmNW Marathon, 48°47.69'N 86°26.11'W,  28.iv.2012, ex.   Equisetum ssuviatile, wet stalks on [sic “near”] surface, [reared] bulk pails, misted daily, 22°C, 16L:8D, 60–70% RH, emerged:  12.v.2012, 1 ♀, K. N. Barberleg. ( CNCI);  Mer Bleue,  7.vi.1923, 2 ♂♂ 1 ♀, C. H. Curranleg. ( CNCI);  Moosonee, 51.24622°N 80.67281°W, Repl. 1 mesic, Malaise trap,  18–21.vi.2010, 1 ♂;  Moosonee, 51.27717°N 80.64778°W, Repl. 3 wet, Malaise trap,  19–22.vi.2010, 1 ♀, both NBP Field Partyleg. (both LEMQ);  Moosonee, 51°16.68'N 80°38.65'W, sweeps, mostly C[ arex]  utriculata,  C. aquatilis, wet sedgemeadow,  10.vii.2014, 6 ♂♂ 2 ♀♀(1 ♀ genit. prep.);  Moosonee, 51°16.69'N 80°38.86'W, general sweeps, sedgemeadow,  9.vii.2014, 1 ♀;  Moosonee, 51°16.36'N 80°39.11'W, sweeps, railside ditch, mostly   Equisetum ssuviatile,  Carexspp.,  10.vii.2014, 2 ♂♂ 1 ♀;  Moosonee, 51°16.55'N 80°39.01'W, sweeps, mostly   Carexspp., wet forest trail,  11.vii.2014, 5 ♂♂;   OtterRapids, 50°10.80'N 81°38.59'W, sweeps,  Carexspp.,  19.vii.2009, 3 ♂♂, sweeps,  Equisetum ssuviatile,  19.vii.2009, 1 ♀, sweeps, roadside  Equisetumspp.,  19.vii.2009, 2 ♂♂ 1 ♀;  ~ 7 kmSSE OtterRapids, 50°07.52'N 81°35.92'W, sweeps,   Scirpussp.,  20.vii.2009, 1 ♂, all K. N. Barberleg. (all CNCI);  Pancake BayP. Pk., 46°58.11'N 84°42.72'W, sweeps from boardwalk, mostly emergent sedges/  Equisetum,  24.vii.2004, 5 ♂♂ 4 ♀♀( DEBU 01500556–64),  2.viii.2004, 6 ♂♂ 5 ♀♀( DEBU 01500848–58, 2 ♀♀genit. prep.),  7.viii.2004, 2 ♂♂ 1 ♀( DEBU 01501074–76),  4.ix.2004, 1 ♀( DEBU 01501430),  27.vi.2005, 2 ♂♂ 7 ♀♀( DEBU 01501683–91, 3 ♀♀genit. prep.),  7.vii.2007, 3 ♀♀( DEBU 01501899–901), all K. N. Barberleg.;  Pancake BayProv. Park, 46°58.11'N 84°42.72'W, sweeping from boardwalk, mostly emergent sedges/  Equisetum,  9.vii.2010, 2 ♀♀, J. Roháčekleg. ( SMOC, 1 ♀ genit. prep.);  Pancake BayP. Pk., 46°58.12'N 84°42.75'W, sweeps, mostly graminoids/  Typhanear wetland boardwalk,  2.viii.2004, 1 ♂( DEBU 01500963);  Pancake BayP. Pk., 46°58.10'N 84°42.71'W, sweeps, mostly   Carexnr.wetland boardwalk,  24.vii.2004, 11 ♂♂ 7 ♀♀( DEBU 01500702–19), sweeps, mostly emergent  Carex/  Equisetumnr. boardwalk,  16.vi.2007, 2 ♀♀( DEBU 01501853, -54, both genit. prep.), all K. N. Barberleg.;  Petawawa,  7.vii.1961, 1 ♂, J. R. Vockerothleg. ( CNCI); Hwy 101 at Prairie BeeRiver(west side bridge), 47°51.81'N 83°54.33'W, sweeps, mostly   Carex utriculata,  14.vii.2013, 5 ♂♂(1 ♂ genit. prep.);  ~ 10.8 kmW Jct Hwys 556 & 129, km64.3 Ranger Lk. Rd., 46°53.46'N 83°27.01'W, sweeps,  Carexsp.,  23.vi.2007, 1 ♀, sweeps,   Equisetum ssuviatileand  Carexsp.,  23.vi.2007, 2 ♀♀, sweeps,   Equisetum ssuviatileand  Dulichium arundinaceum,  23.vi.2007, 1 ♂, all K. N. Barberleg. (all CNCI);  Richmond Fen, ex. forest, 45°05'48.73"N 75°50'39.70"W,  11.vi.2010, 1 ♀(genit. prep.), J. B. Sinclar[sic B. J. Sinclair] leg.;  ~ 4.5 kmE Rosseau, on Aspdin Rd., 45°15.88'N 79°34.88'W, sweeps, mostly   Carexin sedgemeadow,  7.vii.2005, 1 ♀, K. N. Barberleg. (both CNCI);  S[ault] S[ainte] Marie, S. of Algoma U[niversity] College, 46°29.9'N 84°17.2'W, sweeps, graminoids mostly   Carex aquatilis,  11.vi.1997, 4 ♂♂ 5 ♀♀(1 ♂ genit. prep.), sweeps,  Carex aquatilis,  12.vii.1997, 1 ♂ 3 ♀♀,  3.viii.1997, 1 ♂,  22.vi.1998, 1 ♂,  9.vi.2001, 1 ♂, sweeps, mostly  Carex aquatilis,  5.vi.2001, 1 ♂,  12.vi.2001, 1 ♂,  15–16.vi.2001, 1 ♂, sweeps,  Carex lacustris,  12.vii.1997, 2 ♂♂ 9 ♀♀, sweeps,  Scirpussp.,  12.vii.1997, 1 ♂ 5 ♀♀, sweeps,  Calamagrostis canadensis,  26.vi.1998, 1 ♀;  S[ault] S[ainte] Marie, S. of Algoma University, 46°29.88'N 84°17.19'W, sweeps, mostly   Carexspp.,  Scirpus cyperinus,  5.vii.2008, 1 ♀, all K. N. Barberleg. (all CNCI);  S[ault] S[ainte] Marie, Finn Hill, 46°31.63'N 84°17.29'W, sweeps,  Scirpus microcarpus,  19.vii.2004, 12 ♂♂ 9 ♀♀(1 ♂ wing illustration),  4.viii.2004, 3 ♂♂ 2 ♀♀,  12.vi.2005, 1 ♂,  9.vii.2005, 1 ♂ 2 ♀♀(1 ♂ genit.prep.),  27.vi.2007, 2 ♂♂,  13.vii.2007, 4 ♂♂,  12.vi.2008, 15 ♂♂ 9 ♀♀,  6.vii.2008, 1 ♂ 2 ♀♀,  18.vi.2016, 2 ♂♂ 4 ♀♀( CNCI),  29.vi.2016, 126 ♂♂ 92 ♀♀( AMNH, BDUC, CMNH, CASC, CSCA, INHS, KNWR, LACM, LEMQ, MCZN, NMPC, OSAC, PMAE, RBCM, SEMC, SMOC, UBCZ, USNM 5 ♂♂ 4 ♀♀each; CNCI 36 ♂♂ 20 ♀♀), sweeps,  Carex stipata stipata,  25.vi.2009, 1 ♂( CNCI), K. N. Barberleg.; same locality but 46°31.63'N 84°17.33'W, sweeps, mostly   Carex stipata stipata,  4.vi.2013, 1 ♀, sweeps,  Carex stipata stipata,  18.vi.2016, 2 ♂♂ 3 ♀♀,  29.vi.2016, 4 ♂♂ 2 ♀♀,  2.vii.2016, 1 ♂; same locality but 46°31.48'N 84°17.36'W, sweeps, mostly  Scirpus microcarpus,  18.vi.2005, 1 ♀, all K. N. Barberleg. (all CNCI); same locality but sweeping, mostly   Scirpus microcarpus,  7.vii.2010, 2 ♂♂ 5 ♀♀, J. Roháčekleg. ( SMOC, 1 ♂ 1 ♀ genit. prep.);  S[ault] S[ainte] Marie, Landslide Rd., Coldwater Ck. noodplain, 46°33.8'N 84°16.6'W, sweeps, mostly   Carex aquatilis,  13.vi.2001, 1 ♂;  S[ault] S[ainte] Marie, River Road, 46°30.4'N 84°15.3'W, sweeps, mostly   Carexspp.,  11.vi.2001, 1 ♀;  S[ault] S[ainte] Marie, 2 ndLine Ext., 46°32.5'N 84°17.0'W, sweeps, mostly   Carexsp.,  13.vi.2001, 1 ♂;  ~ 10 kmW S[ault] S[ainte] Marie, Sunnyside Beach Rd., 46°30.51'N 84°32.56'W, sweeps,   Equisetum ssuviatile, sedges,  22.vi.2007, 2 ♂♂ 1 ♀;  Searchmont, N Hwy #552, 46°50.3'N 84°04.4'W, sweeps, roadside sedges/ grasses,  10.vii.1998, 1 ♂;  Smooth Rock Falls, 49°16.04'N 81°36.08'W, sweeps,   Carex utriculata,  22.vi.2013, 1 ♂ 3 ♀♀, all K. N. Barberleg. (all CNCI);  Sudbury,  22.vii.1922, 1 ♀, A. L.Melanderleg. ( USNM);  ~61.5 km N Thessalon, Hwy#129, 47°05.00'N 83°09.40'W, sweeps, emergent  Equisetum ssuviatile/ sedges,  23.vi.2007, 1 ♀(genit. prep.);  ~ 74 kmNNE Thessalon, 46°53.94'N 83°16.23'W, shore of Mississagi R., sweeps, graminoids, herbs,  Equisetumspp.[  E. ssuviatile&  E.palustre],  5.vii.2010, 1 ♀(genit. prep.),  17.vii.2010, 1 ♀(genit. prep.), all K. N. Barberleg. (all CNCI);  Toronto,  16.v.1896, 1 ♀, A. L. Melanderleg. ( USNM);   60 kmS White River,  30.vi.1992, 1 ♂, T. A. Wheelerleg. ( LEMQ 0039595).   QUEBEC: Beechgrove, 45°39'N 76°08'W,  27.vi.1984, 2 ♂♂ 1 ♀, J. R. Vockerothleg. ( CNCI);  Lac St-Francois Nat. Wildlife Area, NW of Aménag. Therrien, close to Ruisseau Th[errien], 45°00.39'N 74°30.99'W,  Carexmeadow, sweeping, T1a,  5.vi.1999, 4 ♂♂ 3 ♀♀( LEMQ 0039559, -63–65, -70, -71, -74), T1b,  5.vi.1999, 5 ♂♂ 2 ♀♀( LEMQ 0039561, -62, -66, -67, -69, -72, -75), T1c,  5.vi.1999, 1 ♂ 1 ♀( LEMQ 0039568, -73), T2c,  5.vi.1999, 1 ♂( LEMQ 0039560), all F. Beaulieuleg.;  La Trappe,  27.v.1934, 1 ♀, J. Ouelletleg. ( AMNH);  Mistassini,  3.vii.1956, 1 ♂ 1 ♀( ♀genit. prep.),  7.vii.1956, 1 ♀(genit. prep.), J. R. McGillisleg.;  Mistassini Post,  23.vi.1956, 1 ♀,  4.vii.1956, 1 ♂, J. R. Lonswayleg.;  Old Chelsea, Summit King Mt., 1150',  11.vii.1971, 1 ♀, J. R. Vockerothleg. (all CNCI);  near Schefferville, 54°49.507'N 66°51.393'W, sweep, grass/ sedgemeadow,  8.vii.2008, 1 ♂ 1 ♀, A. Rogicleg. ( LEMQ, 1 ♀genit. prep.).   SASKATCHEWAN: Kenosee,  15.vi.1958, 1 ♀, A. R. Brooksleg.;   Saskatoon, S. Sask[atchewan] R. Valley,off Central Ave., 52°10.4'N 106°36.2'W, sweeps, riverside graminoids,  17.vii.1999, 1 ♀, K. N. Barberleg.;  Uranium City,  21.vi.1962, 1 ♂(genit. prep.), J. G.Chillcottleg. (all CNCI).   YUKON: Alaska Highwayat Yukon River crossing, 60°34'N 134°40'W, sweep grass/ sedgesalong river margin,  2.vii.1997, 1 ♂ 1 ♀, T.A.Wheelerleg.( LEMQ 0039608, -609);  Alaska Highwayat Yukon River crossing, sweep sedgesand grassesalong river,  2.vii.1997, 1 ♀, S. Boucherleg. ( LEMQ 0039610);  Dawson, 1300',  29.vii.1962, 1 ♂ 2 ♀♀, R. E. Leechleg. ( CNCI).   UNITED STATES OF AMERICA: ALASKA: Anchorage,  18.vii.1951, 1 ♂ 1 ♀, R. S. Bigelowleg. ( CNCI);  KenaiNWR [ National Wildlife Reserve], ~ 0.5 miE of Berg Lk., LTEMP site 3271, wetland, sweep net sample,  20.vi.2004, 1 ♂, S. Grimesleg. ( KNWR, ID:3884);   King Salmon, Naknek River,  4.viii.1952, 1 ♂, W. R. Masonleg. ( CNCI);  Knik Lake, NW of Wasilla, sweeping vegetation edge of lake,  18.vii.1978, 4 ♂♂ 4 ♀♀(1 ♂ 1 ♀ genit. prep.);  Moon Lake, Alaska Highway, DC-1331,  8.vii.1978, 1 ♂, all P. H. Arnaud Jr.leg. (all CASC);  Unalakleet,  13.vii.1961, 1 ♂, R. Madgeleg.( CNCI).   COLORADO: Rio Grande Co.,  BeaverCreek, 10000', Malaise trap,  21.vi.1972, 1 ♂ 1 ♀(1 ♀ genit.prep.), W. W. Wirthleg.;  Boulder Co., Nederland,  24.vi.1961, 1 ♀, G. C. Steyskalleg. (all USNM);  Electra Lake, F.4367E, ~ 37°33'N 107°48'W, ~8400',  28.vi.–1.vii.1919, 1 ♂ 1 ♀, [no collector] ( AMNH);  Teller Co.,  3.5 miS Florissant, Sanborn Ranch, along tributary of PlumCreek,  14.vii.2004, 1 ♂ 6 ♀♀, B.A. Footeleg. ( CMNH, 3 ♀♀ genit. prep.);   2 miS Gould,  13.viii.1968, 1 ♀, Omanleg. ( OSAC, genit. prep.);   3 miN Nederland, 8500', marshy stream margin,  2.vii.1961, 5 ♂♂ 2 ♀♀, J. G. Chillcottleg. ( CNCI, 1 ♂ 1 ♀ genit. prep.);  Platte Canyon, nr. Idlewild,  10.vi.1927, 1 ♂, J. M. Aldrichleg. ( USNM).   IDAHO: Boundary[Co.], Perkins Lake, 48°45.6'N 116°05.5'W,  760 m,  3.vi.2006, 1 ♂ 2 ♀♀, W. N. Mathisleg. ( USNM, 2 ♀♀ genit. prep.);  Idaho Co.,  Pollock,  1.v.1959, 1 ♂,  2.v.1959, 1 ♀, [no collector] ( WFBM, each with empty puparial case in gelatin capsule).   MICHIGAN: Clinton Co., Bath,  6.vi.1940, 1 ♀, C. W. Sabroskyleg. ( USNM).   MONTANA:  2 miE Babb,  19.vii.1967, 2 ♀♀, B. A. Footeleg.;   15 miS Big Fork,  26.vi.1967, 4 ♀♀(3 ♀♀ genit. prep.), D. Allenleg. (all USNM);  Granite Co.,  14 miS Clinton, 4000',  27.vi.1971, 1 ♀, J. R. Powersleg. ( CASC);  Gallatin Co., Gallatin R.,  2 miN Gallatin Gtwy[Gateway], 4100',  23.vi.2001, 1 ♂, J. B. Runyonleg. ( MTEC);  33.0 mi W Kalispell,  30.vi.1966, 1 ♂ 1 ♀;   33 miW Kalispell,  3.viii.1966, 1 ♀;  1.0 mi S Swan Lake,  1.vii.1966, 1 ♀(genit. prep.),  2.viii.1966, 2 ♂♂ 1 ♀( 1 ♀genit. prep.),  20 miS Swan Lake,  7.vii.1967, 2 ♂♂ 1 ♀,  14.vii.1967, 1 ♀(genit. prep.), all B. A. Footeleg. (all USNM);  33.0 mi N West Glacier,  5.vii.1966, 3 ♀♀, K. Valleyleg. ( USNM, 1 ♀ with puparial case in gelatin capsule).   NEW YORK: Whiteface Mt.,  19.vii.1962, 4600–4872', 1 ♀, J. R. Vockerothleg. ( CNCI).   WASHINGTON: Pacinc Co., Ft. Canby St. Park,  29.vi.1988, 1 ♀, W. N. & D. Mathisleg.;  Pend Oreille Co., Ione, ca.  10 kmW, Rt. 20, 48°40.4'N 117°28.2'W,  975 m,  5.vi.2006, 4 ♀♀, W. N. Mathisleg.;  Mt. Constitution,  17.vii.1909, 1 ♂, [no collector], 1 ♂, J. M. Aldrichleg.,  2 ♂♂ 2 ♀♀, A. L. Melanderleg.,   22.vii.1909, 2 ♂♂, A. L. Melanderleg.,  1 ♂, C. F. Bakerleg. (all USNM).   WISCONSIN: WashburnCo., T39N R12W B 32, 4.vii.1953, 1 ♀, R. H. Jonesleg. ( USNM, genit. prep.).   WYOMING: Yellowstone Pk., Twin Lakes,  10.vii.1923, 2 ♂♂ 2 ♀♀, A. L. Melanderleg. ( USNM).  Other material examined (not included in typeseries).   CANADA: ALBERTA:W.A. SwitzerP. Pk.,  BeaverRanch Trail, sweeps, 53°29.80'N 117°48.02'W, edge of Jarvis Ck.,  Carex utriculata,  22.vii.2011, 1 ♀, K. N. Barberleg. ( DEBU 01503722, thorax crushed).   ONTARIO: Bruce Peninsula N. P.,  CraneRiverbelow Lake Scugog, 45°07.0'N 81°32.1'W, sweeps, riverside vegetation,  3.vii.1999, 1 ♀, K. N. Barberleg. ( DEBU, headless, genit. prep.); ~   59 km NNWElliot Lake, s. of Rocky Is. Lake, 46°50.16'N 83°03.05'W,  455 m, sweeping, mostly  Carex rostrata? [more likely  C. utriculata] in fen,  3.vii.2010, 2 ♂♂ 1 ♀, J. Roháčekleg. ( SMOC, 2 ♂♂used for molecular work, 1 ♀headless);  Manitoulin Island,  10 kmW Gore Bay, 45°53'N 82°34'W, sweep in grassland alvar,  20.vi.1996, 1 ♂, P. Bouchardleg. (  LEMQ0039583, headless);  Pancake BayP. Pk., 46°58.11'N 84°42.72'W, sweeps from boardwalk, mostly emergent sedges/  Equisetum,  7.viii.2004, 1 ♀, K. N. Barberleg. (  DEBU01501077, abdomen lost);  Sault Ste. Marie, Finn Hill, 46°31.48'N 84°17.36'W, sweeping, mostly  Scirpus microcarpus,  7.vii.2010, 1 ♂, J. Roháčekleg. ( SMOC, used for molecular work).   QUEBEC: Mistassini,  3.vii.1956, 1 ♀, J. R. McGillisleg. ( CNCI, headless).   UNITED STATES OF AMERICA: ALASKA: Fairbanks,  2.vii.1921, 1 ♀, J. M. Aldrichleg. ( USNM, headless).   COLORADO: Summit Co., Frisco,  3.viii.2001, 1 ♀, I. S. Winklerleg. ( BYUC, headless).   WASHINGTON: Mt. Constitution,  17.vii.1909, 1 ♀,  22.vii.1909, 1 ♀, A. L. Melanderleg. ( USNM, both headless).   Description. Male.Total body length 2.22–2.94 mm; externally (including colouration) extremely similar to both relatives, the Palaearctic  A. tschirnhausiand the above described  A. shewelli(though usually more robust than the latter species), therefore mainly differences against these species are stressed in the description below. Head ( Fig. 483) slightly higher than long, with face somewhat more receding than that of  A. shewelliand hence more angular in pronle (as in  A. tschirnhausi). Occiput and frons closely resembling that of  A. shewelliin form, colour and microtomentum, but dark V-shaped area on frons often paler and anteromedially (above frontal lunule) usually interrupted by small pale (ochreous to yellow) area or short stripe. Frontal lunule, face, parafacialia, gena and postgena hardly different from those of  A. shewelli. Mouthparts coloured as in  A. shewellibut clypeus usually paler brown to ochreous. Cephalic chaetotaxy largely as described above for  A. shewellibut both long ors distinctly shorter than oc, anterior ors setula in front of the middle ors often situated closer to the latter, postocular setulae (up to 10) and peristomal setulae (up to 6, sometimes some of them can be duplicated) usually more numerous, and palpus with preapical seta and ventral setulae longer. Eye broadly subovoid, with longest diameter strongly oblique and 1.3–1.5 times as long as the shortest. Smallest genal height about 0.14–0.16 times as long as shortest eye diameter. Antenna entirely yellow as in both relatives, but arista darker than in  A. shewellihaving basal segments pale brown and terminal seta blackish brown. Thorax ( Fig. 483) generally as in both relatives but more densely microtomentose (grey, often with some bluish tinge) and duller than that of  A. shewelli, hence more resembling that of  A. tschirnhausi. Small pale ochreous to ochreous yellow spot on propleuron above fore coxa usually more distinct than that in  A. shewelli. Thoracic chaetotaxy not particularly different from that of relatives although ac microsetae (particularly in larger specimens) usually more numerous, in up to 6 rows in front of suture. Holotypesomewhat aberrant in having 3 strong stpl setae (posterior stpl duplicated) otherwise there are only 2 stpl as in both allied species. Legs with the same colouration as those of typical specimens of  A. shewelliincluding the variably darkened (usually rather pale) last tarsal segments. Pedal chaetotaxy as in  A. shewelli, having ctenidial spine of f 1similarly variable in length, ranging from only as long as to distinctly longer than maximum width of t 1; t 2with usual (for the  A. tschirnhausigroup) short posterior ventroapical seta and rather variable anterior ventroapical setula being small and nne to thicker and longer, although always markedly shorter than the former seta. Wing ( Fig. 486) largely as described above for  A. shewellibut generally broader, R 2+3less upcurved apically, and R 4+5and M usually straighter and more parallel distally. Wing measurements: length 2.36–3.20 mm, width 0.74–0.99 mm, Cs 3: Cs 4= 1.00–1.44, rm\dm-cu: dm-cu = 2.26–3.78. Haltere pale to whitish yellow as that of pale specimens of  A. shewelli. Abdomen with preabdominal terga brown but paler than thorax, and, particularly, T1 (sometimes partly also T2) paler than T3–T5, all similarly shaped to those of  A. shewelli. Colouration of preabdominal sterna variable, ranging from pale ochreous yellow to light brown. S1 bare, with very narrow, brownish-darkened posterior margin, transversely trapezoidal, and wider and shorter than S2 (as in  A. shewelli). S2–S5 setose as in  A. shewelli, becoming wider, larger and sometimes also darker posteriorly. S2 about as long as wide and with more or less darkened anterior third. S3–S5 wider than long, transversely trapezoidal (broader posteriorly), all generally wider than those of  A. shewelli; S5 widest, most transverse and more or less brownish-darkened laterally. T6 very similar to that of relatives but somewhat shorter, being less than half length of T5. S6 and S7 always paler (up to ochreous yellow) than S8 and both with usual brown anterior marginal ledge. S6 with 2–4, S7 with 3–4 short setae. S8 long, tapered posteriorly and setose as that of  A. shewelli, similarly brown to T5, thus not as dark as epandrium. Genitalia externally more resembling those of  A. tschirnhausithan those of  A. shewelli. Epandrium ( Figs 487, 489) blackish brown, high and distinctly tapered dorsally, about as long as  A. tschirnhausibut slightly narrower and more densely setose than that of the latter species, with several (2–4 pairs) only slightly longer and thicker dorsolateral setae; anal nssure small, narrow, rounded (often asymmetrically) triangular, and narrower than that of  A. tschirnhausi. Cercus small, with nne setae, as in relatives. Medandrium ( Fig. 487) very high, dorsally tapered and with narrow end dorsally emarginate as in relatives, resembling that of  A. shewelli(dorsally somewhat narrower than in  A. tschirnhausi). Gonostylus ( Figs 487, 489, 493, 510–512) most similar to that of  A. tschirnhausi, yellow but with apex more or less darkened (brownish, particularly in large specimens), slightly shorter than epandrial height, slender, elongate, tapered and pointed apically, with anterior side concave ( Fig. 510) to almost straight ( Figs 493) and posterior side always strongly convex and with micropubescence usually covering larger portion of its outer side. Hypandrium ( Fig. 490) of moderate size, distally slightly larger than that of  A. tschirnhausiand with anterior internal lobes much larger, darker, more heavily sclerotized and often projecting above dorsal hypandrial margin. Transandrium narrow ( Fig. 491), medially more robust than in  A. tschirnhausiand with a pair of small, low, dark dorsal lobes (as in  A. shewelli); caudal process distinctly developed, narrow, nat, ventrally slightly narrowed, medially somewhat less sclerotized and laterally coalesced to secondary sclerotizations of basal membrane. Pregonite ( Fig. 490) fused with hypandrium, low, posteroventrally with a small, inconspicuous lobe carrying 2 (subequal or 1 slightly longer) setae; anterior nat part of pregonite with 3 setae (all on inner side, posterior seta longest, as in both relatives). Postgonite ( Fig. 490) very slender and elongate, darker proximally and paler distally where bent, apically more acute than in  A. shewellibut not as pointed as in  A. tschirnhausi, with 1 longer seta at middle of anterior margin, outer side with several sensillae. Basal membrane ventrally somewhat bilobed, with nat spines being pigmented ventrolaterally, and less numerous and hyaline medially ( Figs 490, 491); lateral parts adjacent to caudal process of transandrium secondarily sclerotized and amber yellow pigmented as in  A. shewelli. Aedeagal part of folding apparatus ( Fig. 492) with numerous, dense, nat spine-like excrescences, often darker than in relatives; connecting sclerite inconspicuous, very slender, weakly sclerotized, bordering the former. Phallapodeme slender with robust fulcrum, ventrolaterally projecting apex and shortly forked base as in  A. shewelli. Aedeagus slender ( Fig. 492), with small elongate phallophore similar to that of  A. shewelli, indistinctly separated from large distiphallus. Saccus relatively slender, shorter than nlum, sclerotized in proximal half where it originates by lateral rounded sclerite with 1 internal spine; distally membranous and normally with 5 (rarely up to 8) thick spines on apex, thus armature resembling that of  A. shewelli(not that of  A. tschirnhausiwith only 3 terminal spines). Filum formed by single, long, twisted and largely dark sclerite terminating in curved, pointed and nnely bicuspid apex ( Fig. 488) provided with a number (more than 10) subterminal spines; most of nlum (particularly its membranous proximal part) with numerous spinulae (yet more than in  A. shewelli). Ejacapodeme very small, with a small, pointed projection ( Fig. 492).  Figs 481–484. Primary types and females of the Nearctic species of the  Anthomyza tschirnhausigroup. 481 –  A. shewelli sp. nov., holotype male, laterally, body length 2.30 mm; 482 –  A. gilviventris sp. nov., dark form, paratype female, laterally, body length 3.42 mm (USA: Colorado); 483 –  A.gilviventris sp. nov., holotype male, laterally, body length 2.89 mm; 484 – same species, topotypic paratype female, laterally, body length 3.57 mm. Photo by J. Roháček.  Figs 485–486. Wings of the Nearctic species of the  Anthomyza tschirnhausigroup. 485 –  A. shewelli sp. nov., paratype male, wing length 2.7 mm (Canada: Ontario); 486 –  A. gilviventris sp. nov., paratype male, wing length 3.0 mm (Canada: Ontario). Photo by K. N. Barber.  Female.Similar to male unless mentioned otherwise. Total body length 2.50–3.89 mm. 1st antennal nagellomere more or less darkened anterodorsally (more extensively on inner side) as in females of  A. shewelli. Peristomal setulae usually yet more numerous (up to 9) than in male; also palpus often with more setulae (up to 15). f 1with ctenidial spine variable as in male. Wing measurements: length 2.56–3.73 mm, width 0.77–1.21 mm, Cs 3: Cs 4= 0.97–1.42, rm\dm-cu: dm-cu = 2.15–3.15. Abdomen in typical specimens with preabdominal terga (T1–T5) very pale ( Fig. 484), dirty yellow to orange ochreous, at most somewhat brownishdarkened on lateral sides and/or posterior corners. More rarely (see below) abdomen is markedly darkened, with T1–T5 (and also postabdominal T6, T7+S7) largely brown to dark brown ( Fig. 482) or brown-spotted laterally and medially and hence resembling that of  A. shewelli. T1–T5 more transverse than in male; T1 distinctly narrower than T2; T2–T5 subequal in length but T2 slightly narrower than T3–T5, all relatively shortly setose (as in  A. shewelli). Preabdominal sterna light ochreous to pale yellow, only S2 with somewhat darkened anterior third to two-thirds and S1 (bare) with narrowly darkened posterior margin; S2–S5 becoming distinctly larger and wider posteriorly, only S2 as long as wide, S3–S5 slightly to distinctly wider than long, all slightly trapezoidal to suboblong; S5 widest and largest of them, as wide as postabdominal S6 but usually paler than the latter. Postabdomen ( Figs 495, 496, 502, 503, 515), interestingly, more similar to that of  A. shewelli, thus longer, more slender and elongate than that of  A. tschirnhausi. T6 trapezoidal, wider and more transverse than in  A. shewelli, setose as in  A. shewelli; typical specimens with T6 completely yellow, dark specimens with T6 brown-darkened to almost completely dark brown ( Fig. 502). S6 narrower than in  A. tschirnhausibut wider than in  A. shewelli, with setae relatively dense (as in the former species); yellow ( Fig. 496) to brown-darkened ( Fig. 503), rarely with darkened lateral margins. Conical tergosternum T7+S7 ( Fig. 495, 496, 502, 503, 515) long and very slender as in  A. shewelli, somewhat emarginate anterodorsally; in typical specimens largely dark yellow to orange ochreous ( Fig. 495) or narrowly brownishdarkened at posterior margin ( Fig. 484), often with a pair of submedial brown stripes ( Fig. 484) and usually with narrow unpigmented posteromedial area (of variable size); ventral part of T7+S7 ( Fig. 496) mostly yellow to orange, prolonged, forming distinctly darker (orange brown to contrastingly brown) anterolateral pouch-like lobes, and with variably paler to unpigmented areas in anterior third that are usually not (below 7th spiracle) depressed and/ or separated from darker dorsal part by a lateral ridge (compare Fig. 517 to Fig. 516); most of ventral side of T7+S7 covered by whitish micropubescence that is denser and stronger anterolaterally than medially ( Fig. 496); other parts of tergosternum without micropubescence and glossy; setae largely as in  A. shewelli. In dark variant T7+S7 is largely brown ( Fig. 502, 503, 515), paler only anterodorsally or completely brown, and with anterolateral lobes dark. On the other extreme, in paler specimens (e.g. some southwestern Alberta populations) with orange yellow T7+S7, the anteroventral lobes are pale or only slightly darkened, and the apical marginal band is very narrow, diffuse, or absent, often with a submedial pair of ventral stripes (as in Fig. 484). T8 ( Figs 495, 499, 502, 506, 513, 515) elongate, very narrow and dark brown as in  A. shewelli, without micropubescence, setulae longer than in the latter species. S8 ( Figs 496, 499, 500, 503, 506, 507, 513) slightly longer and often narrower than that of  A. shewelli, half length or more of T8, longitudinally divided into two dark brown sclerites and lacking micropubescence as in  A. tschirnhausi, with longer setulae and with 2 pairs of setae in posterior half. Genital chamber ( Figs 497, 499, 500, 504, 506, 507, 513) with one pair of posterior elongate nat sclerites, variable in length, usually as long as or longer than T8 ( Figs 499, 506), but sometimes shorter ( Fig. 513); annular sclerite asymmetrical and situated as in  A. shewelli. Ventral receptacle ( Figs 497, 504) membranous, tubular as in  A. shewelli, with wider proximal part and attenuated distal part terminated by spirally vermicular apex. Accessory gland also closely resembling that of  A. shewelli, including duct. Spermathecae (1+1) relatively small, the larger 0.6–0.8 times as long as medial length of S10, more broadly subovoid and with more densely transversely striated surface ( Figs 494, 498, 501, 505, 514) than in  A. shewelli, less elongate and with surface less densely striated than in  A. tschirnhausi(cf.  ROHÁĆEK2009a: Figs 66, 67) and its smooth basal part with numerous nne setiform ( Figs 494, 498) to somewhat thickened ( Figs 501, 505) spinulae. T10 ( Figs 495, 499, 502, 506) small, elongately subpentagonal (as in  A. shewelli), brown, with a pair of long medial setae and 1–2 pairs of short setulae, otherwise without micropubescence. S10 ( Figs 496, 499, 503, 506) distinctly larger than T10, also brown, relatively narrow and elongate, longer than in both relatives, and largely bare, with a single pair of medial setae, a few setulae at posterior margin and reduced micropubescence restricted to posterior fourth. Cercus ( Figs 495, 499, 502, 506) slightly longer than in  A. shewelli, but similarly shortly setose including distinctly exclinate apical setae, and with limited micropubescence that is restricted to its ventral side. Variability.Some variability of the male gonostylus is shown in Figs 493, 510–512. Compared to  A. shewelli, the variability in shape of the gonostylus is less extreme, but there is more variation in the extent of the micropubescence. On the other hand, the dorsal colouration of the female abdomen is more variable than in  A. shewelli, ranging from almost completely orange yellow (including T7+S7) to largely blackish brown, but it is to be stressed that the extreme colour variants do not occur in all populations. Only the dark form is known from Alaskan populations, but females with brown-darkened preabdominal terga have been recognized in some northeastern (Labrador, Newfoundland) and northwestern (British Columbia, Alaska) areas, as well as in Montana (Swan Lake, Big Fork, both ~ 1000–2000 m) and Colorado, including high-montane areas in the latter ( Nederland, 8500 ft[~ 2600 m], Beaver Creek, 10000 ft[~ 3000 m]). The frequency of occurrence of the dark females may be increased in colder climates. On the other extreme, those smaller females with an entirely orange yellow preabdomen and reduced or no darkening on the anterolateral lobes and posterior margin of T7+S7, seem to occur infrequently among most populations, but are predominant in several populations in southwestern Alberta (Peter Lougheed P. Pk. [~ 1600 m]) and Churchill, Manitoba ( 10–14 m).   Discussion.  Anthomyza gilviventris sp. nov., appears to be very similar and closely allied to both the Nearctic  A. shewelliand the Palaearctic  A. tschirnhausi. Apart from  A. shewellihaving a clearly dissimilar gonostylus, the differentiation of males of  A. gilviventrisand  A. tschirnhausiis more problematic because they both have a similarly formed gonostylus with convex posterior and concave to almost straight anterior margins (cf. Figs 510–512and ROHÁĆEK 2009a: Fig. 57). However, the Palaearctic  A. tschirnhausidiffers from the Nearctic  A. gilviventrisin having a broader epandrium and anal nssure, lower anterior internal lobes of the hypandrium, a less-developed caudal process of the transandrium, and, most importantly, a saccus with only 3 terminal spines. Identincation of females of  A. gilviventrisis more difncult because of a wide colour variability of the abdomen. Typical specimens can be distinguished from those of the above relatives by a strikingly orange yellow to orange ochreous abdomen that is usually accompanied by a pair of contrasting anteroventrolateral dark spots (see Fig. 484) on the pouch-like lobes of T7+S7. Because these spots can sometimes be reduced or missing (especially in some populations in Alberta), these specimens can closely resemble the (albeit very rare) palest females of  A. shewelli. On the other hand, the darkest females of  A. gilviventrishave an abdomen with T1–T6 brown to blackish-brown, darkened as in typical females of  A. shewelli, but the T7+S7 of  A. gilviventrisis almost completely (also anteriorly) dark. In these extreme cases they can be best distinguished from  A. shewellifemales by the relative size and shape of the spermathecae (in  A. gilviventristhe larger spermatheca is only 0.6–0.8 times as long as S10 and more broadly ovoid) and T7+S7 is usually lacking the lateral ridge and the depressed unpigmented micropubescent area below the 7th spiracle. Also, the microtomentum of the mesonotum is characteristically denser, paler and often with a bluish tinge. It is to be stressed that the  A. gilviventrisfemales with dark abdomens are also very similarly coloured to those of  A. tschirnhausi, but the latter species differs by its less posteriorly tapered T7+S7, a wider T8 and S8 (this also shorter) and more elongate spermathecae with striae covering a larger part of their surface (cf.  ROHÁĆEK2009a: Figs 63–69).   Etymology.The new species is named for the largely yellow abdomen of most females (an adjective compounded from gilvus (= Latin yellow) and venter (= Latin abdomen)).   Biology.As discussed for  A. shewelli, the habitat of  A. gilviventrisis also primarily characterized by moist areas including various wetland areas including “fen”, “edge of fen”, and “margin of fen pools”. There are fewer references to wooded areas such as “forest noor”, “forested area”, “boreal forest”, and “wet forest trail”. The most commonly referenced plants are the sedges  Carexand  Scirpus, but other plants are commonly (  Equisetum) or occasionally (  Typha,  Caltha) found intermixed. Grassesare also present in these habitats and are sometimes even predominating. Particularly large collections at one site ( Ontario: Sault Ste. Marie – Finn Hill; see  A. shewelli– Biology) were made from  Scirpus microcarpus, which was a perennially reliable habitat for  A. gilviventris, yielding only a few  A. shewelli. This portion of the sedgemeadow receives and retains signincant runoff from the bordering wooded ridge to the west leading to soil saturation over an extended period of the growing season. A western site ( Alberta: Spray Valley P. Pk.) was predominated by a  Carexthat was not identined to species, but matched other identined specimens of  C. utriculatafrom similar sites sampled in southwestern Alberta(e.g. Peter Lougheed P. Pk.). The nies from this southwestern Albertan locality include those  A. gilviventristhat are smaller in body size and do not have the darkened anteroventral lobes of T7+S 7 inthe female. All the specimens from Manitoba: Churchill, a northern site, were similarly pale and were taken in open areas of tundra and fen, but also boreal forest, although the latter might include wet openings. A single female emerged from a pail of overwintered stalks of  Equisetum ssuviatilecollected in the spring of 2012 ( Ontario: Marathon; see  A. vockerothi– Biology). This was a rare event and the local population of  A. gilviventrisis more likely to be associated with the intermixed  Carex aquatilis, as is also likely for the single reared male of  A. gibbiger. The night period for  A. gilviventrisruns from 1 May ( Idaho: Pollock) to 4 September ( Ontario: Pancake BayP. Pk.).   Distribution.  Anthomyza gilviventrishas a much more extensive distribution compared to that of  A. shewelli. Though it is known from fewer eastern jurisdictions from Newfoundland and Labrador to New York, it ranges west to British Columbia and Washington, and then south to Colorado and north to Alaska, Yukon, and Northwest Territories. Records are available for Canada: Alberta, British Columbia, Labrador, Manitoba, Newfoundland, Northwest Territories, Nova Scotia, Ontario, Quebec, Saskatchewan, Yukon; United States of America: Alaska, Colorado, Idaho, Michigan, Montana, New York, Washington, Wisconsin, Wyoming(see Table 2, Fig. 598). 2011-07-17 CNCI CAN J. Rohacek & K. N. Barber 49.512 Annex Pk 11 -115.06883 Fernie 299 292 1 1 BC holotype 2985226433 [113,1054,1459,1479] 1967-07-27 USNM S. P. Whitney Canada Grey Co. K. Valley 14 48.673332 Banff N. P. 110 -117.47 Fish Creek Prov. 299 292 2 2 ALBERTA paratype 2985226213 1966-07-11 USNM K. Valley Banff 299 296 1 1 paratype 2985226254 Carex, Equisetum 2011-07-17 49.377 Bellevue 11 -114.376335 299 296 8 5 3 paratype 2985225923 Carex 2008-07-22 CNCI K. N. Barber 53.167 Cadomin 10 -117.49533 299 296 26 10 16 paratype 3989399301 [342,1089,1565,1586] 1956-06-10 1956-07-09 1956-06-10 O. Peck Elkwater Lk. Elkwater Lk. 299 296 8 5 3 paratype 3989399302 Carex 2008-07-25 CNCI K. N. Barber 51.063 Exshaw 11 -115.18317 Exshaw 300 297 1 1 paratype 2985226738 Equisetum 2011-08-12 BDUC J. E. Swann 50.926666 Fish Creek Prov. Pk., pond near Shannon Terrace 9 -114.12379 Calgary 300 297 2 2 paratype 2985226554 2010-07-09 BDUC J. E. Swann Fish Creek Prov. Pk., Shannon Terrace Calgary 300 297 1 1 paratype 2985225493 Carex utriculata, Equisetum ssuviatile 2011-07-12 50.926834 Fish Creek P. Pk. 11 -114.12383 300 297 6 3 3 paratype 2985226996 Carex utriculata, Equisetum 2011-07-12 sweeps BDUC J. E. Swann & K. N. Barber 50.929 Fish Creek P. Pk 11 -114.122 300 297 8 4 4 paratype 2985225957 Carex 2008-07-25 sweeps DEBU K. N. Barber 50.523335 Highwood House 11 -114.88533 Highwood House 300 297 2 1 1 paratype 2985226962 Equisetum ssuviatile 2008-07-22 sweeps 53.3545 Hinton 10 -117.622 Hinton 300 297 3 3 paratype 2985225774 Equisetum ssuviatile, Carex 2008-07-22 sweeps C. aquatilis Wahlenb 53.34617 Hinton, Hwy # 40 10 -117.61383 300 297 1 paratype 2985226181 Equisetum palustre 2008-07-23 sweeps 53.2775 Wildhorse Lake 10 -117.742836 300 297 5 1 4 paratype 2985225759 Carex utriculata 2011-07-15 sweeps CNCI K. N. Barber 51.051666 Sibbald Area 11 -114.912 Kananaskis Country 300 297 15 6 9 paratype 2985226963 Equisetum ssuviatile 2008-07-25 sweeps DEBU K. N. Barber 50.9685 Mt. Lorette Ponds 11 -115.10983 Kananaskis Village 300 297 1 1 paratype 2985225642 Carex utriculata 2011-07-25 sweeps CNCI K. N. Barber 53.5365 Range Rd. 10 -117.017 Obed 300 297 7 3 4 paratype 2985225948 Carex utriculata 2011-07-13 2011-07-16 2011-07-13 sweeps DEBU 50.697 Peter Lougheed P. Pk. 11 -115.11533 300 297 DEBU 01502929-3056, DEBU 01503292-95, DEBU 01503333, DEBU 01503390-94 142 37 105 paratype 2985226832 Carex utriculata 2011-07-14 sweeps DEBU 50.616333 Upper Lake Drive 11 -115.118835 300 297 DEBU 01503314 1 1 paratype 2985225449 Carex utriculata 2011-07-16 sweeps DEBU K. N. Barber 50.615665 Upper Lake Drive 11 -115.11933 300 297 DEBU 01503647-51 5 2 3 paratype 2985225909 2008-07-23 DEBU K. N. Barber 53.064835 Lovett River 10 -116.817 300 297 1 1 paratype 2985226208 Carex utriculata? and Poa sp. 2011-07-13 DEBU 50.815834 Spray Valley P. Pk. 11 -115.164 300 297 DEBU 01502773-851 79 40 39 paratype 2985226893 Carex utriculata 2011-07-22 DEBU 53.496666 W. A. Switzer P. Pk. 10 -117.80033 Beaver Ranch Trail 300 297 DEBU 01503707-21 15 2 13 paratype 2985226421 Carex utriculata? 2011-07-22 DEBU 53.4945 W. A. Switzer P. Pk. 10 -117.8 Beaver Ranch Trail 300 297 DEBU 01503777 1 1 paratype 2985226612 2011-07-22 2011-07-24 2011-07-22 DEBU K. N. Barber Canada Grey Co. 14 48.673332 W. A. Switzer P. Pk., off Hay River Rd. W 110 -117.47 Fish Creek Prov. 300 297 DEBU 01503792-801, DEBU 01503847-75 40 10 30 Alaska paratype 2985226541 [258,887,993,1012] 1980-07-14 1980-07-20 1980-07-14 CNCI H. J. Teskey Waterton Lakes Nat. Pk. 300 297 1 1 paratype 2985226192 1932-06-19 USNM O. B [ryant] Brisco 300 297 3 3 paratype 2985225875 1978-06-27 CASC P. H. Arnaud Jr. Alaska Highway Buckinghorse Provincial Campground 300 297 2 2 paratype 2985225438 Carex utriculata 2011-07-17 2011-07-19 2011-07-17 49.512 Annex Pk. 11 -115.06883 Fernie 300 297 82 39 43 paratype 2985225626 Carex utriculata 2011-07-18 CNCI K. N. Barber 49.511 Annex Pk. 11 -115.069336 Fernie 300 297 19 4 15 paratype 2985226676 [494,995,1151,1172] 1992-05-31 A. Borkent 300 297 1 1 paratype 2985226337 Carex pellita 2011-07-18 CNCI K. N. Barber 49.648335 Hwy 95 rest area 11 -115.647 300 297 1 1 paratype 2985226043 1988-07-12 DEBU T. A. Wheeler Slatechuck Mt. Graham Is. 300 297 1 1 paratype 2985226036 [344,789,1233,1252] 18. vi. USNM R. P. Currie Kaslo Cr. 300 297 1 1 paratype 2985226657 2008-06-13 Malaise trap RBCM Cooper Beauchesne & Assoc. Ltd. BC Hydro Kinbasket Lake 300 297 WIL08-01, WIL08-15 2 1 1 paratype 2985226698 Carex utriculata 2011-07-23 K. N. Barber 52.7775 edge of Kinbasket Lake 10 -119.173 300 297 27 9 18 paratype 2985226579 1960-07-17 CNCI W. W. Moss 58.716667 King Salmon L. 1042 -132.9 300 297 9 4 5 paratype 2985226101 1988-07-01 RBCM G. E. Hutchings Merritt 300 297 ENT991-6166 1 1 paratype 2985225895 1957-06-29 CNCI E. E. MacDougall Moresby Camp 300 297 1 1 Queen Charlotte Islands paratype 2985225600 2002-08-06 ZSMC M. v. Tschirnhaus 53.05 Mt. Robson Prov. Pk. 1079 -119.25 300 297 6 1 5 paratype 2985226615 [233,956,1445,1466] 1988-06-11 RBCM G. E. Hutchings Nanaimo Lakes 300 297 2 1 1 paratype 2985226620 Carex utriculata with Equisetum palustre & E. × litorale 2011-07-18 CNCI K. N. Barber 51.061333 Crestbrook Rd. 11 -116.651 Parson 300 297 7 5 2 paratype 3989552301 1960-08-19 R. Pilfrey 0 57.933334 Pete Lake 1047 -131.93333 Pete Lake 300 297 3 2 1 paratype 3989552302 [354,833,1526,1546] 300 297 2985226312 1958-06-03 1967-07-05 1958-06-03 UBCZ Foxlee Robson 300 297 3 3 paratype 2985225627 1980-06-04 RBCM Canadian Wildlife Service 301 294 Qualicum Beach 300 297 ENT991-17202 1 1 paratype 2985226469 1988-06-08 1990-06-08 1988-06-08 A. Borkent Salmon Arm 301 294 2 2 paratype 2985226259 Carex utriculata? 2011-07-20 K. N. Barber 49.837334 Lower Elk Valley Rd. 11 -114.88817 301 294 2 1 1 paratype 2985226488 1959-06-23 1959-06-24 1959-06-23 CNCI E. E. MacDougall 0 Summit Lake 301 294 1 1 paratype 2985226720 1988-07-03 TAUI A. Freidberg Vancouver 301 294 3 2 1 paratype 3989552303 1955-07-09 1955-07-12 1955-07-09 CNCI E. F. Cashman Cartwright 301 294 4 3 1 LABRADOR paratype 3989552305 1955-07-17 1955-07-19 1955-07-17 CNCI E. E. Sterns Cartwright 301 294 11 6 5 LABRADOR paratype 2985226405 tundra 2010-07-14 2010-08-06 2010-07-14 ID 14 58.7045 Farnworth Lake (Landing L.) 6 -94.0525 301 294 ID:10PROBE-14241, ID:JWDCI851-11, ID:10PROBE-14244, ID:JWDCI854-11, ID:10PROBE-14245, ID:JWDCI855-11, ID:10PROBE-16182, ID:JWDCK227-11, ID:10PROBE-16183, ID:JWDCK228-11, ID:10PROBE-15878, ID:JWDCJ1443-11, ID:10PROBE-15879, ID:JWDCJ1444-11, ID:10PROBE-12448, ID:JWDCH242-10, ID:10PROBE-13043, ID:JWDCH837-10, ID:10PROBE-13045, ID:JWDCH839-10 10 5 5 MANITOBA paratype 2985226374 2010-08-01 2010-08-07 2010-08-01 ID 29 58.6332 Twin Lakes 6 -93.7871 301 294 ID:10PROBE-12672, ID:JWDCH466-10, ID:10PROBE-13220, ID:JWDCH020-10 2 1 1 paratype 2985226636 2010-07-14 2010-07-19 2010-07-14 ID 10 58.7541 Launch Road 6 -93.9974 301 294 ID:10PROBE-14512, ID:JWDCJ172-11, ID:10PROBE-16713, ID:JWDCL663-11, ID:10PROBE-16714, ID:JWDCL664-11 3 2 1 paratype 2985226857 boreal forests 2010-07-23 ID J. Wang 10 58.7306 Ramsay Creek 6 -93.7804 301 294 ID:10PROBE-16481, ID:JWDCL241-11 1 1 paratype 2985225378 Carex spp. 2011-07-29 CNCI K. N. Barber 49.637333 11 -95.49817 301 294 1 1 paratype 3989552306 1999-06-17 LEMQ V. Crecco 49.1785 Tallgrass Prairie Reserve 11 -96.67934 301 294 LEMQ 0039607 1 1 paratype 3989552304 1958-07-08 1958-07-17 1958-07-08 CNCI R. L. Hurley Turtle Mtn. For. Res. Int. Peace Gardens 301 294 2 2 paratype 2985225310 1983-06-24 1985-07-11 1983-06-24 CNCI L. Hollett & A. Borkent Bay d'Espoir 301 294 2 1 1 NEWFOUNDLAND paratype 2985225541 1979-06-17 1982-07-15 1979-06-17 NFRC Larson & Swales Portugal Cove 301 294 6 5 1 Avalon Peninsula paratype 2985226203 1967-07-26 CNCI J. F. McAlpine Agric. Exp. Stn. St. John's 301 294 8 5 3 paratype 2985225877 1969-06-23 CNCI G. E. Shewell Norman Wells 301 294 1 1 NORTHWEST TERRITORIES paratype 2985225761 1983-07-05 CNCI A. Borkent Paquette Lk. 301 294 1 1 NOVA SCOTIA paratype 2985225979 2008-05-05 2008-05-21 2008-05-05 Malaise trap DEBU E. Proctor 45.4522 Algonquin P. Pk. 6 -78.4713 301 294 1 1 ONTARIO paratype 2985226677 2001-07-03 LEMQ M. Pollett Awenda P. Pk. 301 294 LEMQ 0040381 1 1 paratype 2985225951 1999-07-03 45.116665 Bruce Peninsula N. P. 113 -81.535 301 294 3 3 paratype 2985225424 Carex aquatilis 2008-06-22 45.1565 Bruce Peninsula N. P. 11 -81.57383 301 294 1 1 paratype 2985226950 2008-06-22 DEBU K. N. Barber 45.191666 Singing Sands 11 -81.576836 Bruce Peninsula N. P. 301 294 5 5 paratype 2985226656 Carex utriculata 2013-06-13 2013-06-23 2013-06-13 47.82 11 -83.59033 301 294 2 2 paratype 2985226142 Carex utriculata 2013-06-23 47.996 11 -82.917336 301 294 3 1 2 paratype 2985226113 Carex utriculata 2012-07-09 2013-08-18 2012-07-09 CNCI K. N. Barber 48.944168 11 -81.003 301 294 26 8 18 paratype 2985225961 1992-07-01 RBCM A. Cannings Grey Co. Anderson Lake Dornoch 301 294 1 1 paratype 2985227090 2008-07-17 49.787834 Dryden 11 -92.81033 Dryden 301 294 1 1 paratype 2985225891 2010-07-10 CNCI K. N. Barber 48.352 along Magpie River 11 -84.56734 Dubreuilville 301 294 1 1 paratype 2985226601 2010-07-10 SMOC J. Rohacek 48.352 along Magpie River 11 -84.56734 Dubreuilville 301 294 4 2 2 paratype 2985225620 2013-06-23 48.588833 11 -80.965 301 294 3 2 1 paratype 2985226422 Carex utriculata 2013-06-08 2013-06-15 2013-06-08 46.494335 Echo Bay Marsh 11 -84.068665 Echo Bay 301 294 6 3 3 paratype 2985225685 Carex utriculata 2012-06-24 2013-06-15 2013-06-08 46.495167 Echo Bay Marsh 11 -84.06734 Echo Bay 301 294 2 1 1 paratype 2985226523 Carex utriculata 2013-06-29 46.3705 Elliot Lake 11 -82.60817 301 294 1 1 paratype 2985226074 Carex utriculata 2013-06-29 CNCI K. N. Barber 46.245167 11 -82.55883 301 294 2 1 1 paratype 2985226369 Carex aquatilis 2010-07-03 CNCI K. N. Barber 302 299 46.836 Rocky Is. Lake 11 -83.050835 301 294 1 1 paratype 2985225826 Carex rostrata? 2010-07-03 SMOC J. Rohacek 455 302 299 15 10 5 paratype 2985225658 1992-07-10 LEMQ T. A. Wheeler Finland Finland 302 299 LEMQ 0039596 1 1 paratype 2985227044 Carex utriculata 2013-06-23 2013-07-13 2013-06-23 48.239 11 -82.34583 302 299 4 2 2 paratype 2985226003 Equisetum ssuviatile, Dulichium arundinaceum 2007-07-09 46.726166 Goulais River 11 -84.4075 302 299 1 1 paratype 2985225562 Equisetum ssuviatile 2007-07-16 CNCI K. N. Barber 46.722168 Goulais River 11 -84.412 302 299 1 1 paratype 2985227071 2010-07-10 SMOC J. Rohacek 46.746834 Goulais River 11 -84.54467 302 299 5 5 paratype 2985227082 Equisetum spp. 2009-07-21 DEBU K. N. Barber 49.218334 Greenwater P. Pk. 11 -81.28917 302 299 DEBU 01502099-101 3 3 paratype 2985225655 Carex utriculata 2013-06-22 49.67267 11 -83.535164 302 299 1 1 paratype 3990052313 Carex utriculata 2013-06-21 CNCI K. N. Barber 49.7345 hydro right-of-way opposite Forde Lk. 11 -84.24567 302 299 2 2 paratype 3990052317 Carex utriculata 1986-06-21 CNCI K. N. Barber 49.7345 Icewater Creek WS [Watershed] 11 -84.24567 302 299 1 2 paratype 2985226090 Carex utriculata 2014-07-12 DEBU 47.371834 Lake Superior P. Pk. 11 -84.687164 302 299 DEBU 01503948 2 2 paratype 2985226746 2010-05-29 DEBU K. N. Barber 47.277668 Lake Superior P. Pk. 11 -84.55 302 299 DEBU 01502431-34 4 2 2 paratype 2985226800 1996-06-20 LEMQ P. Bouchard 45.883335 Manitoulin Island 1127 -82.566666 302 299 LEMQ 0039584 2 2 paratype 2985226956 2010-05-27 DEBU S. A. Marshall 45.796665 Misery Bay Prov. Nat. Res. 18 -82.74944 302 299 DEBU 00324518 1 1 Manitoulin I. paratype 2985225373 Equisetum ssuviatile with Carex sp. 2007-06-16 48.794834 11 -86.435165 302 299 8 3 5 paratype 2985227074 Equisetum ssuviatile with Carex sp. 2008-07-31 CNCI K. N. Barber 48.794834 11 -86.4345 302 299 1 1 paratype 2985226100 Equisetum ssuviatile 2012-04-28 2012-05-12 2012-04-28 CNCI K. N. Barber 48.794834 11 -86.435165 302 299 1 1 paratype 2985225602 [406,918,885,906] 1923-06-07 CNCI C. H. Curran Mer Bleue Mer Bleue 302 299 3 1 2 paratype 2985226561 2010-06-18 2010-06-21 2010-06-18 Malaise trap 51.24622 Moosonee 1 -80.67281 Moosonee 302 299 1 1 paratype 2985225858 2010-06-19 2010-06-22 2010-06-19 Malaise trap LEMQ NBP Field Party 51.27717 Moosonee 1 -80.64778 Moosonee 302 299 1 1 paratype 2985226428 2014-07-10 51.278 Moosonee 11 -80.644165 Moosonee 302 299 8 2 6 paratype 2985226321 [151,915,993,1012] 2014-07-09 51.278168 Moosonee 11 -80.64767 Moosonee 302 299 1 1 paratype 2985226870 Equisetum ssuviatile, Carex spp. 2014-07-10 51.272667 Moosonee 11 -80.65183 Moosonee 302 299 3 1 2 paratype 2985227055 Carex spp. 2014-07-11 51.275833 Moosonee 11 -80.65017 Moosonee 302 299 5 5 paratype 2985226328 2009-07-19 50.18 Otter Rapids 11 -81.643166 Otter Rapids 302 299 7 2 5 paratype 2985226188 Scirpus sp. 2009-07-20 CNCI K. N. Barber 50.12533 11 -81.59866 302 299 1 1 paratype 2985226279 2004-07-24 2007-07-07 2004-07-24 DEBU K. N. Barber 46.9685 Pancake Bay P. Pk. 11 -84.712 302 299 DEBU 01500556-64, DEBU 01500848-58, DEBU 01501074-76, DEBU 01501430, DEBU 01501683-91, DEBU 01501899-901 41 26 15 paratype 2985226239 2010-07-09 SMOC J. Rohacek 46.9685 Pancake Bay Prov. Park 11 -84.712 302 299 2 2 paratype 2985226492 2004-08-02 DEBU 46.968666 Pancake Bay P. Pk. 11 -84.7125 302 299 DEBU 01500963 1 1 paratype 3990052323 Carex nr. wetland boardwalk 2004-07-24 2007-06-16 2004-07-24 DEBU K. N. Barber 46.968334 Pancake Bay P. Pk. 11 -84.71183 302 299 DEBU 01500702-19, DEBU 01501853 20 9 11 paratype 3990052308 1961-07-07 CNCI J. R. Vockeroth Petawawa Petawawa 302 299 1 1 paratype 2985226819 Carex utriculata 2013-07-14 47.8635 Prairie Bee River 11 -83.9055 302 299 5 5 paratype 2985225845 Equisetum ssuviatile and Carex sp., Equisetum ssuviatile and Dulichium arundinaceum 2007-06-23 CNCI K. N. Barber 46.891 km 64.3 Ranger Lk. Rd. 11 -83.450165 302 299 4 3 1 paratype 2985226995 2010-06-11 J. B. Sinclar & B. J. Sinclair 45.096867 Richmond Fen 1 -75.84436 Richmond Fen 302 299 1 1 paratype 2985226277 Carex in sedge meadow 2005-07-07 CNCI K. N. Barber 45.264668 on Aspdin Rd. 11 -79.58134 302 299 1 1 paratype 2985226520 Carex aquatilis 1997-06-11 2001-06-16 1997-06-11 303 296 46.498333 S. of Algoma University College 112 -84.28667 Sault Sainte Marie 302 299 37 23 14 paratype 2985225376 Carex spp. 2008-07-05 CNCI K. N. Barber 46.498 S. of Algoma University 11 -84.2865 Sault Sainte Marie 303 296 1 1 paratype 2985225767 2004-07-19 2016-06-29 2004-07-19 CNCI, AMNH, BDUC, CMNH, CASC, CSCA, INHS, KNWR, LACM, LEMQ, MCZN, NMPC, OSAC, PMAE, RBCM, SEMC, SMOC, UBCZ, USNM K. N. Barber 46.52717 Finn Hill 11 -84.28817 Sault Sainte Marie 303 296 353 144 209 paratype 2985226760 Carex stipata stipata 2013-06-04 CNCI, AMNH, BDUC, CMNH, CASC, CSCA, INHS, KNWR, LACM, LEMQ, MCZN, NMPC, OSAC, PMAE, RBCM, SEMC, SMOC, UBCZ, USNM K. N. Barber 46.52717 Finn Hill 11 -84.28883 Sault Sainte Marie 303 296 13 6 7 paratype 2985225459 Carex stipata stipata 2005-06-18 CNCI K. N. Barber 46.524666 Finn Hill 11 -84.28934 Sault Sainte Marie 303 296 1 1 7 paratype 2985226468 Scirpus microcarpus 2010-07-07 SMOC J. Rohacek 46.524666 Finn Hill 11 -84.28934 Sault Sainte Marie 303 296 7 5 2 paratype 2985226437 Carex aquatilis 2001-06-13 46.56333 Landslide Rd. 112 -84.276665 Sault Sainte Marie 303 296 1 1 paratype 2985225900 Carex spp. 2001-06-11 46.506668 River Road 112 -84.255 Sault Sainte Marie 303 296 1 1 paratype 2985226336 Carex sp. 2001-06-13 46.541668 2 nd Line Ext. 112 -84.28333 Sault Sainte Marie 303 296 1 1 paratype 2985225748 Equisetum ssuviatile 2007-06-22 46.5085 Sunnyside Beach Rd. 11 -84.54266 303 296 3 1 2 paratype 2985225625 [113,967,725,746] 1998-07-10 46.838333 Searchmont 112 -84.07333 Searchmont 303 296 1 1 paratype 2985225725 Carex utriculata 2013-06-22 CNCI K. N. Barber 49.267334 Smooth Rock Falls 11 -81.60133 303 296 4 3 1 paratype 2985225972 [113,587,779,799] 1922-07-22 USNM A. L. Melander Sudbury Sudbury 303 296 1 1 paratype 2985226245 2007-06-23 47.083332 11 -83.15667 303 296 1 1 paratype 2985225309 2010-07-05 2010-07-17 2010-07-05 CNCI K. N. Barber 46.899 shore of Mississagi R. 11 -83.2705 303 296 2 2 paratype 2985225694 1896-05-16 USNM A. L. Melander Toronto Toronto 303 296 1 1 paratype 2985225597 [329,1008,885,906] 1992-06-30 LEMQ T. A. Wheeler 303 296 LEMQ 0039595 1 1 paratype 2985225421 1984-06-27 CNCI J. R. Vockeroth 45.65 Beechgrove 1129 -76.13333 Beechgrove 303 296 3 1 2 QUEBEC paratype 3990052310 1999-06-05 LEMQ F. Beaulieu 45.0065 NW of Amenag. Therrien 11 -74.5165 Lac St-Francois Nat. Wildlife Area 303 296 LEMQ 0039559, LEMQ 0039561, LEMQ 0039568, LEMQ 0039560 17 6 11 paratype 3990052316 [286,734,1019,1039] 1934-05-27 AMNH J. Ouellet La Trappe La Trappe 303 296 1 1 paratype 2985225756 1956-07-03 1956-07-07 1956-07-03 J. R. McGillis Mistassini Mistassini 303 296 4 3 1 paratype 3990052315 1956-06-23 1956-07-04 1956-06-23 J. R. Lonsway Mistassini Post Mistassini Post 303 296 2 1 1 paratype 3990052303 [238,987,1073,1092] 1971-07-11 CNCI J. R. Vockeroth 0 Summit King Mt. Old Chelsea 303 296 1 1 paratype 2985225509 2008-07-08 LEMQ A. Rogic 54.825115 1 -66.85655 303 296 3 2 1 paratype 3990052325 [226,744,1125,1146] 1958-06-15 A. R. Brooks Kenosee 303 296 1 1 SASKATCHEWAN paratype 3990052318 1999-07-17 K. N. Barber 52.173332 S. Sask [atchewan] R. Valley, 108 -106.60333 Saskatoon 303 296 1 1 paratype 2985226418 1962-06-21 CNCI J. G. Chillcott Uranium City Uranium City 303 296 1 1 paratype 2985225873 1997-07-02 LEMQ T. A. Wheeler 60.566666 Yukon River crossing 1031 -134.66667 Alaska Highway 303 296 LEMQ 0039608 2 1 1 YUKON paratype 2985226197 1997-07-02 LEMQ S. Boucher Yukon River crossing Alaska Highway 303 296 LEMQ 0039610 1 1 paratype 2985226229 [318,882,1258,1279] 1962-07-29 CNCI R. E. Leech 0 Dawson Dawson 303 296 3 2 1 paratype 2985225660 1951-07-18 CNCI R. S. Bigelow United States of America Anchorage Anchorage 303 296 2 1 1 ALASKA paratype 2985226007 2004-06-20 KNWR S. Grimes National Wildlife Reserve Kenai 303 296 ID:3884 1 1 paratype 2985225450 [287,921,1338,1359] 1952-08-04 CNCI W. R. Mason Naknek River King Salmon 303 296 1 1 paratype 3990052307 1978-07-18 Knik Lake Knik Lake 303 296 8 4 4 paratype 3990052324 1978-07-08 CASC P. H. Arnaud Jr. Alaska Highway Moon Lake 303 296 1 1 paratype 2985225917 1961-07-13 CNCI R. Madge Unalakleet Unalakleet 303 296 1 1 paratype 2985226350 1972-06-21 Malaise trap W. W. Wirth 0 Beaver Creek Rio Grande Co. 303 296 2 1 1 COLORADO paratype 2985225931 [264,890,1446,1466] 1961-06-24 USNM G. C. Steyskal Nederland Boulder Co. 303 296 1 1 paratype 2985225955 1919-06-28 1919-07-01 1919-06-28 AMNH 37.55 Electra Lake 1180 -107.8 Electra Lake 303 296 2 1 1 paratype 2985225637 2004-07-14 CMNH B. A. Foote Sanborn Ranch Teller Co. 303 296 7 6 1 paratype 2985226111 [113,669,1526,1546] 1968-08-13 OSAC Oman 303 296 1 1 paratype 2985226847 1961-07-02 CNCI J. G. Chillcott 0 303 296 7 2 5 paratype 2985226133 1927-06-10 USNM J. M. Aldrich 304 301 Platte Canyon Platte Canyon 303 296 1 1 paratype 2985225970 2006-06-03 USNM W. N. Mathis 760 48.76 Perkins Lake 110 -116.09167 Boundary 304 301 3 2 1 IDAHO paratype 2985225607 1959-05-01 1959-05-02 1959-05-01 WFBM Pollock Idaho Co. 304 301 2 1 1 paratype 2985225357 1940-06-06 USNM C. W. Sabrosky Bath Clinton Co. 304 301 1 1 MICHIGAN paratype 3990052311 [420,960,299,318] 1967-07-19 B. A. Foote MONTANA 304 301 2 2 MONTANA paratype 3990052319 1967-06-26 USNM D. Allen 304 301 4 4 paratype 3990052304 1971-06-27 CASC J. R. Powers 0 Granite Co. Granite Co. 304 301 1 1 paratype 3990052305 2001-06-23 MTEC J. B. Runyon 0 Gallatin R. Gallatin Co. 304 301 1 1 paratype 3990052302 [390,752,377,398] 1966-06-30 USNM 33.0 mi W Kalispell 304 301 2 1 1 paratype 3990052321 [757,1066,379,398] 1966-08-03 USNM 33 mi W Kalispell 304 301 1 1 paratype 3990052312 1966-07-01 1967-07-14 1966-07-01 USNM B. A. Foote 1.0 mi S Swan Lake 304 301 9 5 4 paratype 3990052309 1966-07-05 USNM K. Valley 33.0 mi N West Glacier 304 301 3 3 paratype 2985225328 1962-07-19 CNCI J. R. Vockeroth 0 Whiteface Mt. Whiteface Mt. 304 301 1 1 NEW YORK paratype 2985227025 1988-06-29 W. N. & D. Mathis Ft. Canby St. Park Pacinc Co. 304 301 1 1 WASHINGTON paratype 2985225652 2006-06-05 W. N. Mathis 975 48.673332 Ione 110 -117.47 Pend Oreille Co. 304 301 4 4 paratype 3990052314 [318,959,537,558] 1909-07-17 1909-07-22 1909-07-17 USNM J. M. Aldrich Mt. Constitution 304 301 1 1 paratype 3990052322 1909-07-17 1909-07-22 1909-07-17 USNM A. L. Melander Mt. Constitution 304 301 4 2 2 paratype 3990052320 [282,626,564,585] 1909-07-22 USNM A. L. Melander Mt. Constitution 304 301 2 2 paratype 3990052301 [632,929,564,585] 1909-07-22 USNM C. F. Baker Mt. Constitution 304 301 1 1 paratype 2985226701 1953-07-04 USNM R. H. Jones Washburn Washburn 304 301 1 1 WISCONSIN paratype 2985226715 1923-07-10 USNM A. L. Melander Twin Lakes Yellowstone Pk. 304 301 4 2 2 WYOMING paratype 3990052306 2011-07-22 DEBU Jarvis Ck. & K. N. Barber Canada Switzer 53.496666 Beaver Ranch Trail 10 -117.80033 Pk. 304 301 1 1 Alberta 2985226851 1999-07-03 DEBU Bruce Peninsula N. P. & K. N. Barber Canada 45.116665 Lake Scugog 113 -81.535 Crane River 304 301 1 1 Ontario 2985225876 2010-07-03 NNW, SMOC J. Rohacek Canada 455 46.836 Rocky Is. Lake 11 -83.050835 304 301 2 2 Ontario 2985226106 1996-06-20 LEMQ P. Bouchard Canada 45.883335 Manitoulin Island 1127 -82.566666 304 301 1 Ontario 2985225760 2004-08-07 DEBU K. N. Barber Canada 46.9685 Pk. 11 -84.712 Pancake Bay 304 301 1 1 Ontario 2985227107 2010-07-07 SMOC J. Rohacek Canada 46.524666 Finn Hill 11 -84.28934 Sault Ste. Marie 304 301 1 Ontario 2985226375 [493,1124,911,931] 1956-07-03 R, CNCI McGillis Canada Mistassini 304 301 1 1 Quebec 2985226218 [151,1124,938,958] 1921-07-02 USNM J. M. Aldrich United States of America Fairbanks 304 301 1 1 Alaska 2985226764 [151,894,964,985] 2001-08-03 BYUC I. S. Winkler United States of America Summit Co. Frisco 304 301 1 1 Colorado 2985225907 1909-07-17 1909-07-22 1909-07-17 USNM A. L. Melander United States of America Mt. Constitution 304 301 2 2 Washington