Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae) Goulding, Tricia C. Khalil, Munawar Tan, Shau Hwai Dayrat, Benoit ZooKeys 2018 763 1 111 7G727 http://zoobank.org/91D9C26C-2483-4330-BBA2-2196C5B91CA2 Goulding Gastropoda Onchidiidae Wallaconchis CoL Animalia Wallaconchis achleitneri Systellommatophora 0 1 Mollusca species achleitneri  Type locality. Australia, Queensland, Bowen, Doughty Creek, 20°01.26'S, 148°14.35'E, st 117, Avicenniaand Rhizophoratrees by a creek with sandy mud.  Type material. Holotype, 7/5 mm [3534], designated here (st 117, MTQ).  Additional material examined. Australia, Queensland, Bowen, Doughty Creek, 20°01.26'S, 148°14.35'E, 64 specimens 16/10 mm [#3] to 8/5 mm [3535]; 14/8 mm [#1], 14/8 mm [#2], 14/8 mm [#4] and 9/6 mm [2645], st 117, Avicenniaand Rhizophoratrees by a creek with sandy mud (st 117, MTQ).  Distribution. Australia: Queensland (type locality).  Habitat (Fig. 50 A-B, Table 3). Animals were found abundantly in a small mangrove patch near a small river, on mud with large grain sand. Wallaconchis achleitneriwas found at only one station in Queensland despite five weeks of sampling, suggesting it may require something particular in this habitat, possibly the sediment.  Etymology.  Wallaconchis achleitneriis dedicated to Stefan Achleitner, a team member in our expedition to Queensland, and whose German translation helped to clarify parts of old and challenging species descriptions.  Diagnosis (Table 5). Externally, Wallaconchis achleitnericannot be distinguished from brown or grey specimens of other Wallaconchisspecies, except by its small size. Internally, the combination of a smooth, long, and narrow penis within a large penial vestibule, a spherical spermatheca, a free (not attached to the body wall by fibers), and a slightly enlarged oviduct distinguishes W. achleitnerifrom other Wallaconchisspecies. Finally, even though its known species distribution may change in the future, W. achleitneriis not presently considered to be sympatric with any other Wallaconchisspecies.  Color and morphology of live animals (Fig. 50 C-E). The dorsal notum is brownish grey. The ocular tentacles are orange-brown. The hyponotum and foot are yellow-grey.  External morphology. The number of dorsal papillae with eyes is three to four papillae with three to four eyes per papilla. There is a retractable papilla with three to four eyes in the center of the dorsal notum, but which is not raised above the other papillae.  Digestive system (Fig. 51, Table 4). Examples of radular formulae are presented in Table 4. The length of the rachidian teeth is approximately 15 µm, significantly smaller than that of the lateral teeth. The length of the hook of the lateral teeth gradually increases (from 30 to 35 µm) from the inner to the outer teeth (excluding the innermost and outermost lateral teeth which are significantly smaller). The intestinal loops are of type I.  Figure 51. Radula, Wallaconchis achleitneri, Australia, Queensland (MTQ st117). A Rachidian and innermost lateral teeth, scale bar 6 μm[#1] B Rachidian and innermost lateral teeth, holotype, scale bar 20 μm[3534] C Transition between inner lateral teeth and outer lateral teeth, holotype, scale bar 30 μm[3534] D Outermost lateral teeth, holotype, scale bar 10 μm[3534] E Lateral teeth with basal lateral spine decreasing in size from right to left, scale bar 10 μm[#1] F Left lateral teeth, scale bar 100 μm[#2].  Reproductive system (Fig. 52A). In the posterior (hermaphroditic) part of the reproductive system, the oviduct is larger in circumference than the deferent duct. The spherical spermatheca joins the distal region of the oviduct through a short duct.  Figure 52. Reproductive system, Wallaconchis achleitneri, Australia, Queensland (MTQ st117). A Hermaphroditic (female), posterior parts, scale bar 1 mm [#4] B Anterior, male copulatory parts, scale bar 1 mm [#1]. Abbreviations: dd deferent duct fgm female gland mass hd hermaphroditic duct hg hermaphroditic gland ov oviduct ps penial sheath rm retractor muscle rs receptaculum seminis sp spermatheca v vestibule.  Copulatory apparatus (Figs 52B, 53). The penis (approximately 0.6-0.9 mm long) is narrow and smooth with no hooks (Fig. 53). The penial vestibule is cylindrical and the penial sheath is narrow. The distal end of the penis lies within the vestibule (Fig. 52B). The distal end of the deferent duct is approximately the same width as the penial sheath. The deferent duct is convoluted. The retractor muscle is slightly longer than the penial sheath and inserts at the posterior end of the body cavity, near the rectum.  Figure 53. Penis, Wallaconchis achleitneri, Australia, Queensland (MTQ st117). A Scale bar 100 μm[2645] B Holotype, scale bar 100 μm[3534].