Tetramorium moravicum Kratochvil , 1941: 86 Tetramorium moravicum Tetramorium Tetramorium forte Tetramorium rhenanum Schulz, 1996: 391 T. moravicum Schlick-Steiner et al. 2005: 1 Taxonomic revision of the Palaearctic Tetramorium chefketi species complex (Hymenoptera: Formicidae). Csösz S. Radchenko, A. Schulz, A. Zootaxa 2007 1405 1 38 urn:lsid:biosci.ohio-state.edu:osuc_concepts:144999 Kratochvil, 1941 Kratochvil 1941 Insecta Formicidae Tetramorium Animalia Tetramorium moravicum Kratochvil Hymenoptera 1 Arthropoda species moravicum   Tetramorium moravicum  Kratochvil, 1941: 86([[worker]], [[queen]]); also described as new in  Kratochvil1944: 71([[worker]], [[male]]); TYPE MATERIAL: PARATYPES“ Tetramorium moravicum” [-] “ 1941” [/] “ Mohelno” [now CZECH REPUBLIC] [/] “l. Dr Kratochvil” [-] Paratypus [/] “ Tetramorium” [/] “ moravicum Kratochv.” (1[[worker]], 2[[males]]/ ZMMU); junior synonym of Tetramorium forte: Radchenko 1992b: 51; revived from synonymy: Seifert 1996: 160, not Radchenko et al 1998: 110, Czechowski et al. 2002: 66, Radchenko et al. 2004: 77;  Tetramorium rhenanum Schulz, 1996: 391([[worker]], [[queen]], [[male]]) TYPE MATERIAL: HOLOTYPEand PARATYPES: Hessen, Rheinland, [ GERMANY] 3km N. Lorchhausen, 35km W. Wiesbaden, 100mH, 15.06.1994. leg. Schulz(3[[workers]], 1[[queen]] / MHNG, 12[[workers]], 3[[queens]], 2 [[males]] / PCAS); junior synonym of T. moravicum Schlick-Steiner et al. 2005: 1;  Redescription of worker (figs 46-49). Medium to large size, CS 833 [720, 953]. Whole body and appendages dark brown to black. Head nearly square, CL/CW 1.02 [0.97, 1.06], with very feebly convex sides, straight occipital margin and narrowly rounded occipital corners. Eyes small, EYE 0.170 [0.165, 0.183]. Frons moderately narrow, FR/CS 0.36 [0.34, 0.38], frontal lobes usually as wide as frons, rarely slightly wider, FL/FR 1.01 [1.0, 1.04]. Scape long, SL/CS 0.80 [0.77, 0.83], with well developed longitudinal dorsal carina basally, parallel costulae extending scape. Promesonotal dorsum slightly convex, metanotal groove deep. Propodeal teeth long. Petiolar node trapezoidal in profile, NOH/NOL 1.07 [0.90, 1.29], petiole relatively high, PEH/NOL 1.94 [1.72, 2.23]. General appearance coarsely rugoso-reticulate, ground surface microreticulate, dull. Head dorsum longitudinally rugose and microreticulate, occiput rugoso-reticulate. Alitrunk dorsum, mesopleuron and dorsum of petiolar node rugoso-reticulate, ground surface coarsely microreticulate, dorsum of postpetiole longitudinally rugulose and microreticulate. Polygonal striation continuous on 1st gastral tergite (see fig. 8.). Ventral surface of head with several short and few moderately long, straight, or few iCi-shapesetae arising posteriorly to buccal cavity (see fig. 5). Redescription of gyne (figs 43-45). Medium to large size, CS 1198 [1045, 1275]. Whole body and appendages dark brown to black. Head distinctly wider than long, CL/CW 0.88[0.84, 0.92], with straight, subparallel sides, straight occipital margin and widely rounded occipital corners. Frons narrow, FR/CS 0.36 [0.34, 0.38], frontal lobes as wide as frons, FL/FR 1.0 [1.0, 1.02]. Scape moderately short, SL/CS 0.73 [0.71, 0.75], with well developed longitudinal dorsal carina basally, parallel costulae extending scape. Head as wide as scutum, MW/CS 1.07 [1.05, 1.13]. Propodeal teeth long. Dorsal crest of petiolar node convex, with well visible protuberance medially in frontal view; in profile petiolar node dorsum blunt. Petiole and postpetiole relatively narrow, WAIST 0.92 [0.86, 1.0]. General appearance coarsely rugose, ground surface microreticulate, dull. Head dorsum, occiput and sides rugoso-reticulate, ground surface microreticulate. Frons longitudinally rugose and microreticulate. Scutum longitudinally rugose, anteriorly smooth, scutellum more or less smooth, sides finely rugulose. Sides of alitrunk, ruguloso-reticulate and microreticulate, ventral part of katepisternum usually smooth and shiny. Dorsum of petiolar node and postpetiole coarsely reticulate; median protuberence of petiolar node smooth. Polygonal striation disrupted on 1st gastral tergite, sometimes continuous basally. Ventral surface of head with several short and few moderately long, straight, or few C-shape setae arising posterior to buccal cavity. Redescription of male. Whole body and appendages black. Head with feebly convex sides, straight occipital margin and rounded occipital corners. Head narrower than scutum. Propodeal teeth very short, propodeum nearly rounded in profile. Dorsal crest of petiolar node blunt not emarginate in frontal wiew. Head, alitrunk and waist finely sculptured, partly shiny, the rest of ground surface microreticulate. Head finely reticulate ground surface microreticulate. Scutum finely rugulose, laterally and anteriorly smooth and shiny. Scutellum longitudinally rugulose, usually shiny medially. Sides of alitrunk finely rugose and microreticulate. Dorsum of petiolar node feebly reticulate and microreticulate, postpetiole generally smooth and shiny. Polygonal striation disrupted on 1st gastral tergite.  Material examined (64 nest series including 374 workers, 28 gynes and 43 males).  AUSTRIA- Setzberg, 1,5km N. Spitz, Wachau, 48°23'N 15°25'E 300mH,  05.05.2001. nr.251. & nr.253. leg. Schulz(4[[workers]] / PCAS);  BULGARIA- Blagoierbad,  15.06.1972. leg. Poldi(2[[workers]] / MSNM);  Malko Trnovo,  25.05.1958, leg. Pisarski(7[[workers]] / HNHM);  FRANCE- Breil-sur-Roya 6km N, Dep. Alpes Maritimes, vic. Maurioun, 450- 700mH,  17.07.1994. leg. Schulz(3[[workers]] / MHNG, 20[[workers]], 4[[males]] / PCAS);  Camargue, Albaron,  15.06.1974. leg. Poldi(2[[workers]], 2[[queens]] / MSNM),   20.06.1974. leg. Poldi(2W, 2[[males]] / MSNM);  GEORGIA- Nr.3672. Kavkaz, Gumista, Suchumi,  21.11.1963, leg Pisarski(6[[workers]], 3 [[males]] / HNHM, 6[[workers]], 4 [[males]] / MIZ);  GERMANY- Hessen, Mittleres Rheintal, oberhalb Lorchhausen, 250m,  18.IV.2003, leg. A. Schulz, R. Guesten& M. Sanetra(3[[workers]] / HNHM, 6[[workers]] / PCAS);  Kaiserstuhl, Badberg ¾km w Steinbr., Bd.-Wtmbg., 320m,  13.IV.2003, leg. A. Schulz, R Guesten& M. Sanetra(3[[workers]] / HNHM, 6[[workers]] / PCAS);  Kaiserstuhl, Schelingen, Bd.-Wtmbg., ( Suedteil), 300-320m,  12.IV2003, leg. A. Schulz & R. Guesten(3[[workers]] / HNHM, 6[[workers]] / PCAS),  (Nordteil), 320-340m,  13.IV2003, leg. A. Schulz, R. Guesten& M. Sanetra(3[[workers]] / HNHM, 6[[workers]], 3[[queens]], 6[[males]] / PCAS);  Mittleres Rheintal, obh. Bacharach-Steeg 200m Rheinland-Pfalz,  18.IV2003, leg. A. Schulz, R. Guesten& M. Sanetra(3[[workers]] / HNHM, 6[[workers]] / PCAS);  Nahetal, 1km e Burg Layen, 120m, Rheinland-Pfalz,  14.IV2003, leg. G. Heller, A. Schulz & R. Guesten(3[[workers]] / HNHM, 6[[workers]] / PCAS);  Nahetal, 2km SE Schlossboeckelheim, 150m, Rheinland-Pfalz,  14.IV2003, leg. G. Heller, A. Schulz & R. Guesten(3[[workers]] / HNHM, 10[[workers]], 8[[queens]], 8[[males]] / PCAS);  Nahetal, 2km w Norheim, 120m, Rheinland-Pfalz,  14.IV2003, leg. G. Heller, A. Schulz & R. Guesten(3[[workers]] / HNHM, 6[[workers]] / PCAS);  GREECE- Ossa Mt.,  28.05.1989, nr.GR001. leg. R. Sciaky(6[[workers]] / HNHM);  Parnon, 4km WSW. Kastanitsa, 37°17'N 22° 40'E, Peleponnes, Prov. Arkadia, 1200-1400mH,  22.04.2000. nr.03-19. leg. A. Schulz(4[[workers]] / PCAS);  Mani, 20 km SW Githeo, 36°40' N 22°24'E, Peleponnes, Prov. Lakonia, 50 mH,  23.04.2000. nr.04-31. leg. A. Schulz(1[[queen]],5[[workers]] / PCAS);  Peloponesos Aetos,  27.06.1986. leg. Sabbadini(4[[workers]] / MSNM);  Pilion, vic. Chania, Prov. Magnissia, 1000- 1400mH,  14.05.1996. nr.11-213., nr.11-214. & nr.11-215. leg. A. Schulz, K. Vock(12[[workers]] / PCAS);  Smolikos Mt, N.slope,  28.05.1989, nr.GR003. leg. R Sciaky(2[[workers]] / HNHM);  HUNGARY-  Matrafuered, Szent-Ivany,  02.05.1937. leg.  Moczar(8[[workers]] / HNHM);   Soshartyan,  24.04.1999. nr.04. leg.  Balint(1[[worker]] / HNHM);  ITALY- Casola, 670m Appennines, Parmense, 09.1991. leg. Grasso(4[[workers]] / MSNM);  Em.Rom (Re) Cinquecerri,  24.04.1978. leg. Ferri(4[[workers]] / MSNM);  Forti di Genova,  15.04.1993. leg. Poldi(3[[workers]] / MSNM);  Mts. Subasio, (PG)  05.06.1993. leg. Platania(2[[workers]], 1[[queen]] / MSNM);  Novafeltria, Emilia,  21.09.1951, leg. E.C.M.Yarrow(11[[workers]] / NHM);  Piemonte, Cuneo prov., Valdieri env. 800mH,  16.06.1988, leg. Rigato(8[[workers]], 1[[queen]], 1[[male]] / NHM);  Sulmona, AQ,  05.02.1962. leg. Poldi(2[[workers]] / MSNM);  Umbria, Spoleto Protte [?],  01.04.1989. leg. Poldi(5[[workers]] / MSNM);  Valdieri (CN) 800-850m,  03.07.1989. leg. Rigato(2[[workers]], 1[[queen]] / MSNM);  ROMANIA-  Kolozsvar(Cluj Napoca), Szenafuevek, Transylvania,  07.08.2003. no.AA087. leg.  Marko(10[[workers]] / HNHM);   Kolozsvar, Szenafuevek, Transylvania,  17.09.2003. leg.  Marko(5[[workers]] / HNHM);  SERBIA-MONTENEGRO- Brezoviza, Mts. Sar, Kosovo, 900-1200mH,  23.05.1971, nr.16. leg Papp & Horvatovich(6[[workers]] / HNHM);  Nis. Jugo nr.384, 18.1984. leg. Poldi(1[[worker]] / MSNM);  Pec Pecka, Banja, Kosovo, 600mH,  18.05.1971. nr.14. leg. Papp & Horvatovich(2[[workers]] / HNHM);  Titograd,  10.06.1972. leg. Poldi(2[[workers]], 1[[queen]], 1[[male]] / MSNM);  TURKEY- Aydintepe, vil. Guemueshane1200mH,  01.07.1975. nr.10. leg. Osella(3[[workers]], 3[[queens]], 1[[male]] / MSNM);  Bitlis, 10km SW Kuecueksu15km S Tatvan 1700mH, Rasenflaechean Bachlauf  16.06.1993. nr.1084. leg. Schulz(6[[workers]] / PCAS);  Erzurum, 5km SW Aydogdu 20km SW Goele, 1400mH,  26.06.1993. nr.1149. leg. Schulz(6[[workers]],1[[male]]/ PCAS);  Erzurum, Sac Gecidi 40km SE Horasan, 2300mH, Hochsteppe und Bachlauf, Wiese  22. 06. 1993. nr. 1114. leg. Schulz(6[[workers]] / PCAS);  Kars, Dagpinar 25km SE Kars 1800mH, Steppe mit Vulkangesteinen  23.06.1993. nr.1121. leg. Schulz(6[[workers]] / PCAS);  Kars, nahe Posof, 1700mH, an der Georgischen Grenze  25.06.1993. nr.1144. nr.1145. nr.1146. leg. Schulz(30[[workers]], 6[[queens]], 5[[males]] / PCAS);  Kars, zwischen Cildir und Camlicatak, ca 20-40km E Ardahan, 1800-2000mH,  24.06.1993. nr.1125. leg. Schulz(3[[workers]] / PCAS);  Rize, 20km NW Ovitdagi Gecidi 30km SE Rize 1000-1400mH, Laubwald 50% auf Sand  30.06.1993. nr.1199. leg. Schulz(6[[workers]] / PCAS);  Van, 20 km NNW Catak 2300-2700mH, Steppenvegetation  18.06.1993. nr.13-1098. leg. Schulz(5[[workers]] / PCAS);  Van, 5km SE Dedeli 30km SE Patnos 1700mH, Hochsteppe  20.06.1993. nr.1103. & nr.1105. leg. Schulz(12[[workers]] / PCAS);  Van, 8km E Budakli 40 km W Gevas, 2200mH, feuchte Wiese und Steppe  17.06.1993. nr.1086., nr.1088. & nr.1090. leg. Schulz(29[[workers]], 1[[queen]] / PCAS);  Van, Migaros,  15.08.1987. leg. Pavesi(2[[workers]] / MSNM);  UKRAINE- Manhup-Kale, Krim, SW Bachcysaraj, 300-500mH,  12.08.1995. nr.04-T524. leg. M. Sanetra(6[[workers]] / PCAS);  Simferopol, S-Krim, vic. Zalis’s’a. 300-500mH,  13.08.1995. nr.05-T527. & nr.05-T528. leg. leg. Sanetra(3[[workers]] / MHNG, 11[[workers]]/ PCAS);  Morphometrics: (108 workers and 12 gynes measured).  Diagnosis. Workers of T. moravicumare distinguishable from most species of the chefketispecies complex (except for T. rhodiumand T. syriacum) by their usually finely costulate scapes bearing a well visible and long dorsal carina basally. The most distinctive features, to separate T. moravicumfrom T. rhodiumworkers, the shape of the petiole in profile (NOH/NOL, PEH/NOL, Table 1.), and the relative length of scape (SL/CS, Table 1.); the frontal width (FR/CS, Table 1.) gives appropriate discrimination of the workers of T. moravicumand T. syriacum. Gynes of T. moravicumdiffer from known gynes of other species of the chefketispecies complex by the relatively wide scutum, MW/CS 1.07 [1.05, 1.13] and by the dorsal carina of the scape. For further combination of morphometric characters see Table 1-2.  Distribution. Widespread in the western Palaearctic from S France to Caucasus. The westernmost known locality of this species is Camargue, France. 923906574 1941 ZMMU Dr Kratochvil CZECH REPUBLIC Mohelno 1 paratype 923906575 1994-06-15 MHNG, PCAS Schulz GERMANY Rheinland, 3 km N. Lorchhausen, 35 km W. Wiesbaden, 100 mH 1 Hessen holotype 923906576 2001-05-05 PCAS Schulz AUSTRIA 48.383335 Setzberg, 1,5 km N. Spitz, Wachau, 300 mH 15.416667 1 nr. 251, nr. 253 Lower Austria 923906577 1972-06-15 MSNM Poldi BULGARIA Blagoierbad 1 923906582 1958-05-25 HNHM Pisarski Bulgaria Malko Trnovo 1 923906578 1994-07-17 MHNG, PCAS Schulz FRANCE Breil-sur-Roya, vic. Maurioun, 450-700 mH 6 km N 1 Dep. Alpes Maritimes 923906579 1974-06-15 MSNM Poldi France Albaron 1 Camargue 923906581 1974-06-20 MSNM Poldi France Albaron 1 Camargue 923906585 1963-11-21 HNHM, MIZ Pisarski GEORGIA Kavkaz, Gumista, Suchumi 1 Nr. 3672 923906583 2003-04-18 HNHM, PCAS A. Schulz, R. Guesten, M. Sanetra GERMANY Mittleres Rheintal, oberhalb Lorchhausen, 250 m 1 Hessen 923906580 2003-04-13 HNHM, PCAS A. Schulz, R Guesten, M. Sanetra Germany Kaiserstuhl, Badberg 0.75 km w Steinbr., 320 m 1 Baden-Württemberg 923906584 2003-04-12 HNHM, PCAS A. Schulz, R. Guesten Germany Kaiserstuhl, Schelingen (Südteil), 300-320 m 1 Baden-Württemberg 923906589 2003-04-13 HNHM, PCAS A. Schulz, R. Guesten, M. Sanetra Germany Kaiserstuhl, Schelingen (Nordteil), 320 - 340 m 1 Baden-Württemberg 923906586 2003-04-18 HNHM, PCAS A. Schulz, R. Guesten, M. Sanetra Germany Mittleres Rheintal, obh. Bacharach-Steeg 200 m 1 Rheinland-Pfalz 923906587 2003-04-14 HNHM, PCAS G. Heller, A. Schulz, R. Guesten Germany Nahetal, 1 km e Burg Layen, 120 m 1 Rheinland-Pfalz 923906591 2003-04-14 HNHM, PCAS G. Heller, A. Schulz, R. Guesten Germany Nahetal, 2 km SE Schlossboeckelheim, 150 m 1 Rheinland-Pfalz 923906590 2003-04-14 HNHM, PCAS G. Heller, A. Schulz, R. Guesten Germany Nahetal, 2 km w Norheim, 120 m 1 Rheinland-Pfalz 923906588 1989-05-28 HNHM R. Sciaky GREECE Ossa Mt. 1 nr. GR 001 923906594 2000-04-22 PCAS A. Schulz Greece 37.283333 Parnon, 4 km WSW. Kastanitsa, Peleponnes, 1200-1400 mH 22.666666 1 nr. 03 - 19 Prov. Arkadia 923906592 2000-04-23 PCAS A. Schulz Greece 36.666668 Mani, 20 km SW Githeo, Peloponnes, 50 mH 22.4 1 nr. 04 - 31 Prov. Lakonia 923906595 1986-06-27 MSNM Sabbadini Greece Peloponesos Aetos 1 923906596 1996-05-14 PCAS A. Schulz, K. Vock Greece Pilion, vic. Chania, 1000-1400 mH 1 nr. 11 - 213, nr. 11 - 214, nr. 11 - 215 Prov. Magnissia 923906593 1989-05-28 HNHM R Sciaky Greece Smolikos Mt, N. slope 1 nr. GR 003 923906598 1937-05-02 HNHM Moczar HUNGARY Matrafuered, Szent-Ivany 1 923906597 1999-04-24 HNHM Balint Hungary Soshartyan 1 nr. 04 923906600 1991-09 MSNM Grasso ITALY Casola, 670 m Appennines 1 Parmense 923906601 1978-04-24 MSNM Ferri Italy Cinquecerri 1 Emilia-Romagna 923906599 1993-04-15 MSNM Poldi Italy Forti di Genova 1 923906621 1993-06-05 MSNM Platania Italy Mts. Subasio 1 Perugia 923906606 1951-09-21 NHM E. C. M. Yarrow Italy Novafeltria 1 Emilia 923906605 1988-06-16 NHM Rigato Italy Valdieri env. 800 mH 1 Piemonte, Cuneo prov. 923906617 1962-02-05 MSNM Poldi Italy Sulmona 1 L'Aquila 923906609 1989-04-01 MSNM Poldi Italy Spoleto Protte [?] 1 Umbria 923906614 1989-07-03 MSNM Rigato Italy Valdieri (CN) 800 - 850 m 1 Cuneo 923906604 2003-08-07 HNHM Marko ROMANIA Kolozsvar (Cluj Napoca), Szenafuevek 1 no. AA 087 Transylvania 923906623 2003-09-17 HNHM Marko Romania Kolozsvar, Szenafuevek 1 Transylvania 923906612 1971-05-23 HNHM Papp, Horvatovich SERBIA-MONTENEGRO Brezoviza, Mts. Sar, 900-1200 mH 1 nr. 16 Kosovo 923906622 1984 MSNM Poldi SERBIA-MONTENEGRO Nis 1 Jugo nr. 384 923906613 1971-05-18 HNHM Papp, Horvatovich SERBIA-MONTENEGRO Pec Pecka, Banja, 600 mH 1 nr. 14 Kosovo 923906611 1972-06-10 MSNM Poldi SERBIA-MONTENEGRO Titograd 1 923906602 1975-07-01 MSNM Osella TURKEY Aydintepe, vil. Guemueshane 1200 mH 1 nr. 10 923906619 1993-06-16 PCAS Schulz Turkey 10 km SW Kuecueksu 15 km S Tatvan 1700 mH, Rasenflaeche an Bachlauf 1 nr. 1084 Bitlis 923906620 1993-06-26 PCAS Schulz Turkey 5 km SW Aydogdu 20 km SW Goele, 1400 mH 1 nr. 1149 Erzurum 923906616 1993-06-22 PCAS Schulz Turkey Sac Gecidi 40 km SE Horasan, 2300 mH, Hochsteppe und Bachlauf, Wiese 1 nr. 1114 Erzurum 923906615 1993-06-23 PCAS Schulz Turkey Dagpinar 25 km SE Kars 1800 mH, Steppe mit Vulkangesteinen 1 nr. 1121 Kars 923906607 1993-06-25 PCAS Schulz Turkey nahe Posof, 1700 mH, an der Georgischen Grenze 1 nr. 1144, nr. 1145, nr. 1146 Kars 923906603 1993-06-24 PCAS Schulz Turkey zwischen Cildir und Camlicatak, ca 20 - 40 km E Ardahan, 1800 - 2000 mH 1 nr. 1125 Kars 923906608 1993-06-30 PCAS Schulz Turkey 20 km NW Ovitdagi Gecidi 30 km SE Rize 1000 - 1400 mH, Laubwald 50 % auf Sand 1 nr. 1199 Rize 923906618 1993-06-18 PCAS Schulz Turkey 20 km NNW Catak 2300 - 2700 mH, Steppenvegetation 1 nr. 13 - 1098 Van 923906610 1993-06-20 PCAS Schulz Turkey 5 km SE Dedeli 30 km SE Patnos 1700 mH, Hochsteppe 1 nr. 1103, nr. 1105 Van 923906625 1993-06-17 PCAS Schulz Turkey 8 km E Budakli 40 km W Gevas, 2200 mH, feuchte Wiese und Steppe 1 nr. 1086, nr. 1088, nr. 1090 Van 923906629 1987-08-15 MSNM Pavesi Turkey Migaros 1 Van 923906624 1995-08-12 PCAS M. Sanetra UKRAINE Manhup-Kale, SW Bachcysaraj, 300 - 500 mH 1 nr. 04 - T 524 Crimea 923906649 1995-08-13 MHNG, PCAS Sanetra UKRAINE Simferopol, S-Krim, vic. Zalis’s’a. 300 - 500 mH 1 nr. 05 - T 527, nr. 05 - T 528 Crimea