Review of the family Coccidae (Hemiptera: Coccomorpha) in Laos Choi, Jinyeong Soysouvanh, Pheophanh Lee, Seunghwan Hong, Ki-Jeong Zootaxa 2018 2018-08-17 4460 1 1 62 5SVQ 536010 Williams & Watson, 1990 Williams & WatsOn 1990 [238,728,548,574] Insecta Coccidae Milviscutulus GBIF Animalia Hemiptera 41 42 Arthropoda genus   Type species:  Lecanium mangiferae Green, 1889, by original designation.   Diagnosis.Dorsum with anal plates together kite-shaped, each plate with anterolateral margin over 1.7 times as long as posterolateral margin ( Figs 37D, 38E); dorsal setae clavate or flagellate ( Fig. 38F); tubular ducts absent; duct tubercles present or absent ( Fig. 38D); and preopercular pores present ( Fig. 38C). Marginal setae with pointed or fimbriate apices ( Fig. 38M). Stigmatic clefts small, each containing 3 stigmatic spines ( Fig. 38M). Venter with multilocular disc-pores each with 7–10 loculi ( Fig. 38K); ventral tubular ducts of 2 types (type I: each with a broad inner ductule; type II: each with a narrow inner ductule) ( Figs 37C, 38H); antenna 6 to 8 segmented, usually with 7 segments ( Fig. 38P); legs with tibio-tarsal articulatory scleroses ( Fig. 38J) ( Williams & Watson 1990; Hodgson 1994).   Remarks.  Milviscutuluscontains 4 described species, most of which are confined to Indonesia, Fiji,  PapuaNew Guinea, Solomon Islandsand  WesternSamoain the Orientaland Australian regions, except for one tropicopolitan species,  M. mangiferae. The genus is similar to  Protopulvinaria, but is easily differentiated from the latter by the distribution of ventral tubular ducts which are scarce or absent on the head and prothorax; in contrast,  Protopulvinariahas abundant ventral tubular ducts in these areas ( Williams & Watson 1990; Hodgson 1994).